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Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
Michelinoceras is the oldest known genus of the Michelinocerida, more commonly known as the Orthocerida, characterized by long, slender, nearly cylindrical orthocones with a circular cross section, long camerae, very long body chambers, and a central or near central tubular siphuncle free of organic deposits. Septal necks are straight; connecting rings cylindrical and thin. Cameral deposits are well developed. A radula has been found in one species, with seven teeth per row. It had ten arms, two of which formed longer tentacles.
Bathmoceras is a primitive cephalopod genus from the Middle and Upper Ordovician. It is a member of the order Cyrtocerinida and is the only genus in the family Bathmoceratidae.
Clarkoceras is a genus of breviconic ellesmerocerid cephalopods, one of only two genera known to have crossed from the Late Cambrian, Trempealeauan, into the Early Ordovician, Gasconadian. ; the other being Ectenolites.
The Gasconadian Stage is the first stage of the Ordovician geologic period in North America and of the Lower Ordovician Canadian Epoch, coming immediately after the Late Cambrian Trempealeauan and preceding the middle Canadian Demingian Stage. The Gasconadian is equivalent to the European Tremadocian and roughly to the Skullrockian of the Ibexian series.
The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic).
The Genus Hemichoanella is a small, extinct, orthoconic nautiloid cephalopod from the Lower Ordovician of Western Australia assigned to the orthoceratoid family, Baltoceratidae. Hemichoanella and the Baltoceratidae are part of the subclass of once diverse and numerous shelled cephalopods known as the Nautiloidea.
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.
Rioceras is an extinct genus of orthocerid cephalopods of the family Baltoceratidae that lived in what would be North America during the Early Ordovician from 480—472 mya, existing for approximately 8 million years.
Rhabdiferoceras is an extinct genus of orthocerids belonging to the Baltoceratidae that lived in what would be North America during the Cassinian Stage at the end of the Early Ordovician, existing for approximately two million years from about 474 -472 mya.
Metabaltoceras, first described by Rousseau Flower in 1964, is a fossil cephalopod genus in the family Baltoceratidae, with a small, slender, fairly fusiform shell, and a large empty siphuncle in contact with the ventral surface. The siphuncle is achoanitic, having virtually no septal necks, segments formed all but entirely by connecting rings. Shells are generally straight, beginning with a subcircular cross section in the adapical portion but becoming faintly depressed adorally. Sutures are straight and transverse except ventrally, where they produce deep, prominent lobes.
Macroloxoceras is a large pseuorthocerid from the upper Devonian of Central Colorado and Southern New Mexico with features resembling those found in actinocerids. Pseudorthocerids and actinocerids are extinct nautiloid cephalopods, generally with long straight shells and expanded siphuncle segments filled with organic deposits.
Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.
Tajaroceras is an extinct slender cephalopod from the uppermost Lower Ordovician of western North America, belonging to the Orthocerid family Troedssonellidae.
Troedssonellidae is a family of orthoceroid cephalopods from the Ordovician, derived from rod-bearing Baltoceratidae, that have a continuous lining within the siphuncle that resembles very thin and slender endocones. Shells are generally slender and orthoconic. The siphuncle is central or subcentral, composed of straight or slightly expanded segments. Septal necks generally short and connecting rings are thin. Thin cameral deposits are known, which along with the position of the siphuncle and thin connecting rings distinguishes them from the endocerids in which they have been included.
Bajkaloceras is a straight-shelled orthoceroid, and possibly a member of the Intejocerida, from the Angara River basin in central Russia, named by Balashov in 1962. Its age, as given in the Treatise on Invertebrate Paleontology is Arenigian.
Ellesmeroceras is the type genus for the Ellesmeroceratidae, a family of primitive nautiloid cephalopods, that is characterized by its small, generally compressed, gradually expanded, orthoconic shell, found in Lower Ordovician marine sediments. The septa are close spaced and the siphuncle is ventral, about 0.2 the diameter of the shell. Septal necks are typically orthochoanitic but may slant inwardly (loxochoanitic) or reach halfway to the previous septum (hemichoanitic). Connecting rings are thick. As common for the Ellesmerocerida, Ellesmeroceras has diaphragms within the siphuncle tube.
Cyclostomiceratidae is an extinct family of Early Ordovician, (Cassinian), ellesmerocerid cephalopods characterized by short, essentially straight shells, a fairly rapidly expanding phragmocone and a ventral siphuncle in which septal necks are almost non-existent and connecting rings are thick and layered. As typical of the Ellesmerocerida, chambers are short, septa close spaced.
References
- Flower, R.H. 1964, The Nautiloid Order Ellesmeroceratida (Cephalopoda)
Memoir 12; New Mexico Bureau of Mines and Mineral Resources, Socorro, NM
- Furnish W.M and Glenister, Brian F 1964, Ellesmerocerida in the Treatise on Invertebrate Paleontology pp K129-K157, section on Baltoceratidae, K154-156.