| Birkeniidae | |
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| Fossil and model of Birkenia elegans | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | † Anaspida |
| Order: | † Birkeniiformes |
| Family: | † Birkeniidae Traquair, 1899 [1] |
| Genera | |
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| Synonyms | |
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Birkeniidae is an extinct family of jawless fish belonging to the order Birkeniiformes. They were defined in 1899 by Ramsay H. Traquair, and were a successful family among the many jawless fish in the Silurian seas.
Birkeniidae had a large body which was ovular in shape, and a strong, mobile tail. Their eyes were positioned on either side of the head. Their mouth was large for the jawless fish, but contained no teeth. Birkeniidae are distinguished by the crown shaped projection on their back, and the other smaller projections which run down its tail. They also possessed fins which faced downwards, an uncommon trait among the jawless fish. They had typical scaly skin.
Cephalaspidomorphs are a group of jawless fishes named for Cephalaspis of the osteostracans. Most biologists regard this taxon as extinct, but the name is sometimes used in the classification of lampreys, because lampreys were once thought to be related to cephalaspids. If lampreys are included, they would extend the known range of the group from the Silurian and Devonian periods to the present day. They are the closest relatives of jawed fishes, who emerged from within them and they would survive if the jawed fish are included.
Acanthodii or acanthodians is an extinct class of gnathostomes, typically considered a paraphyletic group. They are currently considered to represent a grade of various fish lineages leading up to the extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans and chondrichthyans. In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians. Paraphyletic groupings are problematic, as one can not talk precisely about their phylogenic relationships, their characteristic traits and literal extinction.
Placodermi is a class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devonian period. Their head and thorax were covered by articulated armoured plates and the rest of the body was scaled or naked, depending on the species. Placoderms were among the first jawed fish; their jaws likely evolved from the first of their gill arches. Placoderms are thought to be paraphyletic, consisting of several distinct outgroups or sister taxa to all living jawed vertebrates, which originated among their ranks. This is illustrated by a 419-million-year-old fossil, Entelognathus, from China, which is the only known placoderm with a type of bony jaw like that found in modern bony fishes. This includes a dentary bone, which is found in humans and other tetrapods. The jaws in other placoderms were simplified and consisted of a single bone. Placoderms were also the first fish to develop pelvic fins, the precursor to hindlimbs in tetrapods, as well as true teeth. Paraphyletic groupings are problematic, as one can not talk precisely about their phylogenic relationships, characteristic traits, and complete extinction. 380-million-year-old fossils of three other genera, Incisoscutum, Materpiscis and Austroptyctodus, represent the oldest known examples of live birth. In contrast, one 2016 analysis concluded that placodermi are likely monophyletic, though these analyses have been further dismissed with more transitional taxa between placoderms and modern gnathosthomes solidying their paraphyletic status.
Ostracoderms are the armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic and thus does not correspond to one evolutionary lineage. However, the term is still used as an informal way of loosely grouping together the armored jawless fishes.
Heterostraci is an extinct subclass of pteraspidomorph jawless vertebrate that lived primarily in marine and estuary environments. The first identifiable heterostracans appear in the fossil record during the Early Silurian, and all, save for the Psammosteids, became extinct by the start of the late Devonian. This last group of heterostracans died out in the extinction event at the end of the Devonian.
Thelodonti is a class of extinct jawless fishes with distinctive scales instead of large plates of armor.
Anaspida is an extinct group of primitive jawless vertebrates that lived primarily during the Silurian period, and became extinct soon after the start of the Devonian. They were classically regarded as the ancestors of lampreys. Anaspids were small marine agnathans that lacked a heavy bony shield and paired fins, but have a striking highly hypocercal tail. They first appeared in the early Silurian, and flourished until the early Devonian, when they disappear from the fossil record.
The class Osteostraci is an extinct taxon of bony-armored jawless fish, termed "ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian.
Jamoytius kerwoodi was a species of primitive, eel-like jawless fish that lived in the Llandovery epoch of the Early Silurian period.
Slimonia is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Slimonia have been discovered in deposits of Silurian age in South America and Europe. Classified as part of the family Slimonidae alongside the related Salteropterus, the genus contains three valid species, S. acuminata from Lesmahagow, Scotland, S. boliviana from Cochabamba, Bolivia and S. dubia from the Pentland Hills of Scotland and one dubious species, S. stylops, from Herefordshire, England. The generic name is derived from and honors Robert Slimon, a fossil collector and surgeon from Lesmahagow.
Ramsay Heatley Traquair FRSE FRS was a Scottish naturalist and palaeontologist who became a leading expert on fossil fish.
Lasanius is a genus of basal anaspida from the Late Silurian. Its fossils are around 27 mm to 40 mm in length.
Furcacauda is a genus of thelodontid agnathan from the Lower Devonian of Canada, and is the type genus of the order Furcacaudiformes. Furcacaudiform thelodontids were deep water jawless vertebrates with symmetrical fork and lobed-finned tails and scales smaller than typical loganellid and nikoliviid thelodonti scales. Furcacaudiform thelodonts are noted as having a laterally compressed body, large anterior eyes, slightly posterior, lateral, and vertical to a small mouth, and a condensed curved row of branchial openings (gills) directly posterior to the eyes. Many but not all had laterally paired fins. Wilson and Caldwell also note the presence of a caudal peduncle and a long caudal fin made of two large lobes, one dorsal and one ventral separated by 8 to 14 smaller intermediate lobes, giving the appearance of a striated half-moon shaped tail resembling the tail of a heterostracan. A large square cavity within the gut connecting a small intestine to an anal opening lead many to believe that it is this genus that exhibits the first vertebrate stomach. According to Wilson and Caldwell their discovery, based on sediment infillings of fossils of the Furcacauda heintze, gives credence to the evolutionary development of stomach before jaws.
Drepanolepis is an extinct genus of thelodont which lived in Canada during the Early Devonian period. They are considered a "traditional" thelodont and are classified by their forked tails.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
Pteraspidiformes is an extinct order of heterostracan agnathan vertebrates known from extensive fossil remains primarily from Early Devonian strata of Europe and North America, and from Upper Silurian Canada.
Loganellia is a genus of jawless fish which lived between 430-370 million years ago, during the Silurian and Devonian periods of the Paleozoic. Loganellia belonged to the Thelodonti class and like other Thelodonts possessed scales instead of plate armor.
Birkeniiformes is an extinct order of jawless fish belonging to the class Anaspida.
Saarolepis is an extinct genus of jawless fish belonging to the family Birkeniidae.
Anaspidomorphi (anaspidomorphs) is an extinct superclass of jawless fish.
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