Birkenia Temporal range: | |
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Fossil and model of B. elegans | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | † Anaspida |
Order: | † Birkeniiformes |
Family: | † Birkeniidae |
Genus: | † Birkenia Traquair 1898 |
Type species | |
†Birkenia elegans Traquair, 1898 | |
Species | |
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Birkenia is a genus of extinct anaspid fish from Middle Silurian strata of Northern Europe, and Middle Silurian to possibly Earliest Devonian strata of Arctic Canada. [1] Birkeniid anaspids are covered by a series of small plates on the head and rod-shaped scales in a cheveron-like pattern on the trunk. [2]
Intact fossil specimens of B. elegans suggest the living animal reached a length of up to 10.0 cm (3.9 in), and was an active swimmer. In addition to whole specimens and scale microfossils of B. elegans, which are found in Great Britain and Scandinavia, [3] scales of a second species, B. robusta, [1] are found in Late Silurian strata of Scandinavia and Estonia. The scales of B. robusta differ from those of B. elegans in that, as the specific epithet suggests, the scales of the former are more robustly proportioned than those of the latter. Fossil specimens also show a rib that is modified by being narrow at the ends and gradually broadens and rises towards the middle of the scale, where a major elevation is developed that is expressed as a raised and slightly angled ridge. [2]
Chondrichthyes is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or bony fish, which have skeletons primarily composed of bone tissue. Chondrichthyes are aquatic vertebrates with paired fins, paired nares, placoid scales, conus arteriosus in the heart, and a lack of opercula and swim bladders. Within the infraphylum Gnathostomata, cartilaginous fishes are distinct from all other jawed vertebrates.
The Devonian is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.2 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.9 Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Old Red Sandstone, abbreviated ORS, is an assemblage of rocks in the North Atlantic region largely of Devonian age. It extends in the east across Great Britain, Ireland and Norway, and in the west along the eastern seaboard of North America. It also extends northwards into Greenland and Svalbard. These areas were a part of the paleocontinent of Euramerica (Laurussia). In Britain it is a lithostratigraphic unit to which stratigraphers accord supergroup status and which is of considerable importance to early paleontology. The presence of Old in the name is to distinguish the sequence from the younger New Red Sandstone which also occurs widely throughout Britain.
Acanthodii or acanthodians is an extinct class of gnathostomes. They are currently considered to represent a paraphyletic grade of various fish lineages basal to extant Chondrichthyes, which includes living sharks, rays, and chimaeras. Acanthodians possess a mosaic of features shared with both osteichthyans and chondrichthyans. In general body shape, they were similar to modern sharks, but their epidermis was covered with tiny rhomboid platelets like the scales of holosteians.
Placoderms are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates, and the rest of the body was scaled or naked depending on the species.
Pterygotus is a genus of giant predatory eurypterid, a group of extinct aquatic arthropods. Fossils of Pterygotus have been discovered in deposits ranging in age from Middle Silurian to Late Devonian, and have been referred to several different species. Fossils have been recovered from four continents; Australia, Europe, North America and South America, which indicates that Pterygotus might have had a nearly cosmopolitan (worldwide) distribution. The type species, P. anglicus, was described by Swiss naturalist Louis Agassiz in 1839, who gave it the name Pterygotus, meaning "winged one". Agassiz mistakenly believed the remains were of a giant fish; he would only realize the mistake five years later in 1844.
Anaspida is an extinct group of jawless fish that existed from the early Silurian period to the late Devonian period. They were classically regarded as the ancestors of lampreys, but it is denied in recent phylogenetic analysis, although some analysis show these group would be at least related. Anaspids were small marine fish that lacked a heavy bony shield and paired fins, but were distinctively hypocercal.
Jamoytius kerwoodi is an extinct species of primitive, eel-like jawless fish known from the Patrick Burn Formation in Scotland, dating to the Llandovery epoch of the Early Silurian period.
Rhyncholepis was an extinct genus of anaspid from the Late Silurian. Fossils of species R. butriangula and R. parvula have been found in Ringerike, Norway, and Oesel, Estonia. The genus has two species, described in 1911 and 2002.
Andreolepis is an extinct genus of prehistoric fish, which lived around 420 million years ago. It was described by Walter Gross in 1968 based on scales found in the Hemse Formation in Gotland, Sweden. It is placed in the monogeneric family Andreolepididae and is generally regarded as a primitive member of the class Actinopterygii based on its ganoid scale structure; however some new research regards it as a stem group of osteichthyans.
Entelognathus primordialis is an early placoderm from the late Silurian of Qujing, Yunnan, 419 million years ago.
The Cowie Formation is geological formation located on the Highland Boundary Fault between the fishing village of Cowie and Ruthery Head, in Aberdeenshire, Scotland. Age of this formation is controversial, that was originally estimated at the Middle Silurian, Wenlock to Ludlow, but zircon geochronology shows the Early Devonian, Lochkovian instead. In study published in 2023, according to spore microfossils and additional zircon data, the Middle Silurian, late Wenlock age is suggested again. However, in 2024 it is considered as unsustainable because this conclusion is based on adjacent structurally separated block with different stratigraphy and sedimentology to the block with fossil productive Fish Bed. This formation preserves fossils, including a millipedes such as Pneumodesmus and Cowiedesmus that were discovered by Mike Newman in 2001, and some agnathan like Cowielepis.
The Sundvollen Formation is a geologic formation cropping out along the northern and eastern shores of Steinsfjorden, Oslo Region, Norway. It preserves fossils dating back to the Gorstian to Ludfordian stages of the Late Silurian period.
Cyathaspidiformes is an extinct order of heterostracan vertebrates known from extensive fossil remains primarily from Silurian to Early Devonian strata of Europe, and North America, and from Early Devonian marine strata of Siberia.
Cowielepis is an extinct genus of jawless fish in the class Anaspida. It is from the Cowie Harbour fish bed of Stonehaven, Scotland, which age is considered as the Silurian, or Early Devonian (Lochkovian).
Septentrionia is an extinct genus of jawless fish belonging to the family Septentrioniidae. It is the type genus of its family.
Liivilepis is an extinct genus of jawless fish belonging to the family Septentrioniidae.
Tiiu Märss is an Estonian geologist and palaeoichthyologist.
Valentina Karatajūtė-Talimaa was a Lithuanian geologist and paleontologist. Best known for her contributions to the world of vertebrate fossils and vertebrate paleontology, She discovered over 100 new species during her career.