Dryopteris villarii

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Dryopteris villarii
Dryopteris villarii (cropped).JPG
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Division: Polypodiophyta
Class: Polypodiopsida
Order: Polypodiales
Suborder: Polypodiineae
Family: Dryopteridaceae
Genus: Dryopteris
Species:
D. villarii
Binomial name
Dryopteris villarii
(Bellardi) Woyn. ex Schinz & Thell.
Synonyms [1]
  • Nephrodium villarii(Bellardi) Beck.
  • Polypodium villarii(Bellardi)

Dryopteris villarii, commonly known as the rigid buckler fern, [2] is a perennial leptosporangiate fern native to Central and South East Europe as well as the Western Caucasus. It was first described in 1915. [1]

Contents

Description

Dryopteris villarii typically reaches heights around 30 to 60 cm. [3] The rigid, leathery fronds are of a matt glaucous to dark green colour, with scaly reddish petioles and rachis. [4] [5] Fronds may present tufted, erect or spreading. [5] The short bipinnate leaflets are slightly dentate and narrowly oblong-lanceolate, attenuating closer to the base, with yellowish aromatic glands on either side. [3] [4] [5] Rhizomes are far creeping and adventitious, horizontal to ascending. [5] Sori between 0.7 mm to 1 mm, usually with glandular indusium and arranged in two rows along the midrib, usually four to six per row. They do not cover the apex. [3] [4] [5]

Distribution

Dryopteris villarii appears within dolomite and limestone pavements, scree, cliffsides, rocky outcrops, scrub and non-alpine and non-saline grasslands. [5] [6] It is found between 500 m to 2000 m. [4] It is tolerant of any soil type although prefers alkaline. [6]

Related Research Articles

<span class="mw-page-title-main">Frond</span> Collection of leaflets on a plant

A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the large leaves of cycads, as well as palms (Arecaceae) and various other flowering plants, such as mimosa or sumac. "Frond" is commonly used to identify a large, compound leaf, but if the term is used botanically to refer to the leaves of ferns and algae it may be applied to smaller and undivided leaves.

<i>Sceptridium</i> Genus of ferns

Sceptridium is a genus of seedless vascular plants in the family Ophioglossaceae, closely allied to the genus Botrychium. It is also closely related to the genus Botrypus. Sceptridium species are commonly called the grape-ferns.

<i>Dryopteris</i> Genus of plants

Dryopteris, commonly called the wood ferns, male ferns, or buckler ferns, is a fern genus in the family Dryopteridaceae, subfamily Dryopteridoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). There are about 300-400 species in the genus. The species are distributed in Asia, the Americas, Europe, Africa, and the Pacific islands, with the highest diversity in eastern Asia. It is placed in the family Dryopteridaceae, subfamily Dryopteridoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Many of the species have stout, slowly creeping rootstocks that form a crown, with a vase-like ring of fronds. The sori are round, with a peltate indusium. The stipes have prominent scales.

<span class="mw-page-title-main">Dryopteridaceae</span> Family of ferns

The Dryopteridaceae are a family of leptosporangiate ferns in the order Polypodiales. They are known colloquially as the wood ferns. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Polypodiineae. Alternatively, it may be treated as the subfamily Dryopteridoideae of a very broadly defined family Polypodiaceae sensu lato.

<i>Dryopteris expansa</i> Species of fern

Dryopteris expansa, the alpine buckler fern, northern buckler-fern or spreading wood fern, is a species of fern native to cool temperate and subarctic regions of the Northern Hemisphere, south at high altitudes in mountains to Spain and Greece in southern Europe, to Japan in eastern Asia, and to central California in North America. The species was first described from Germany. It prefers cool, moist mixed or evergreen forests and rock crevices on alpine slopes, often growing on rotting logs and tree stumps and rocky slopes. It is characteristically riparian in nature, and is especially associated with stream banks.

<i>Gymnocarpium dryopteris</i> Species of fern

Gymnocarpium dryopteris, the western oakfern, common oak fern, oak fern, or northern oak fern, is a deciduous fern of the family Cystopteridaceae. It is widespread across much of North America and Eurasia. It has been found in Canada, the United States, Greenland, China, Japan, Korea, Russia, and most of Europe. It is a seedless, vascular plant that reproduces via spores and have a life cycle with alternating, free-living sporophyte and gametophyte phases.

<i>Minuartia</i> Genus of flowering plants in the carnation family Caryophyllaceae

Minuartia is a genus of flowering plants commonly known as sandworts in the family Caryophyllaceae.

<i>Dryopteris affinis</i> Species of fern in the family Dryopteridaceae

Dryopteris affinis, the scaly male fern or golden-scaled male fern, is a fern native to western and southern Europe and southwestern Asia. It is most abundant on moist soils in woodlands in areas with high humidity, such as the British Isles and western France. In the Mediterranean region and the Caucasus it is confined to high altitudes.

<i>Dryopteris marginalis</i> Species of fern

Dryopteris marginalis, vernacularly known as the marginal shield fern or marginal wood fern, is a perennial species of fern found in damp shady areas throughout eastern North America, from Texas to Minnesota and Newfoundland. It favors moderately acid to circumneutral soils in cooler areas but is fairly drought-resistant once established. In the warmer parts of its range, it is most likely to be found on north-facing non-calcareous rock faces. It is common in many altitudes throughout its range, from high ledges to rocky slopes and stream banks. Marginal wood fern's name derives from the fact that the sori are located on the margins, or edges of the leaflets.

<i>Dennstaedtia</i> Genus of ferns

Dennstaedtia is a mostly tropical and subtropical genus of ferns described as a genus in 1801. hayscented fern, or cup ferns, are common names for some species in this genus. It includes 58 species native to the tropical Americas, Madagascar, southern and eastern Asia, Australia, New Guinea, and the Pacific Islands.

<i>Dryopteris ludoviciana</i> Species of fern

Dryopteris ludoviciana, the southern woodfern, is fern native to southern United States from Florida west to Texas and as far north as Kentucky and North Carolina.

<i>Phegopteris hexagonoptera</i> Species of fern

Phegopteris hexagonoptera, commonly called the broad beech fern, is a common forest fern in the eastern United States and adjacent Ontario. It grows from a creeping rootstock, sending up individual fronds that more or less clump. Its native habitat includes moist, undisturbed, hardwood forests.

<i>Dryopteris intermedia</i> Species of wood fern

Dryopteris intermedia, the intermediate wood fern or evergreen wood fern, is a perennial, evergreen wood fern native to eastern North America. It is a diploid species, and is the parent of several species of hybrid origin, including Dryopteris carthusiana. Other common names for this species include intermediate shield fern, fancy wood fern, fancy fern, glandular wood fern, American shield fern and common wood fern.

<i>Sceptridium dissectum</i> Species of fern

Sceptridium dissectum is a common fern in the family Ophioglossaceae, occurring in eastern North America. Like other plants in this group, it normally only sends up one frond per year. It has long been the subject of confusion because the frond presents in one of two forms, either the normal form that resembles other plants in the genus, or the skeletonized form.

<span class="mw-page-title-main">Fern sports</span>

Fern sports are plants that show marked change from the normal type or parent stock as a result of mutation. The term Morphotype is also used for any of a group of different types of individuals of the same species in a population. Fern fronds in sports are typically altered in several ways, such as the frond apex divided and pinnae similarly duplicated.

<i>Chirosia betuleti</i> Species of fly

Chirosia betuleti is a species of fly, which causes knotting gall in ferns. The gall develops in the terminal shoots of ferns, such as broad buckler fern, male fern, lady fern, and ostrich fern.

<i>Dryopteris macropholis</i> Species of fern

Dryopteris macropholis is a species of fern. It is distributed on the Marquesas Islands.

<i>Dryopteris dilatata</i> Species of fern

Dryopteris dilatata, the broad buckler-fern, is a robust species of deciduous or semievergreen fern in the family Dryopteridaceae, native to Europe, particularly western and central Europe. In southern Europe, it is mostly found in mountainous regions. It is also found between the Black Sea and the Caspian Sea. It grows to 90 cm (35 in) tall by 120 cm (47 in) wide, with dark green tripinnate fronds, the ribs covered in brown scales.

<i>Dryopteris aemula</i> Species of fern

Dryopteris aemula, the hay-scented buckler-fern or hay-scented fern, is a species of perennial leptosporangiate fern.

<i>Dryopteris fragrans</i> Species of fern

Dryopteris fragrans, commonly known as the fragrant woodfern, a circumboreal fern, is the smallest of the Dryopteris species. It can resemble Woodsia ilvensis in the wild, with which it shares the same habitat of rocky areas, shady cliffs, screes, and limestone talus. It typically will not reach more than 25 cm in height, and accumulates dead fronds around its base.

References

  1. 1 2 "Dryopteris villarii (Bellardi) Woyn. ex Schinz & Thell". Plants of the World Online . Royal Botanic Gardens, Kew. Retrieved 9 April 2024.
  2. "Dryopteris villarii (Rigid Buckler Fern)". www.plants.ces.ncsu.edu. n.d. Retrieved 9 April 2024.
  3. 1 2 3 "Dryopteris villarii (Bellardi) Woyn. ex Schinz & Thell". www.tela-botanica.org. n.d. Retrieved 9 April 2024.
  4. 1 2 3 4 "Dryopteris villarii (Bellardi) Woyn. ex Schinz & Thell. - Villars fern". www.actaplantarum.org. 2009. Retrieved 9 April 2024.
  5. 1 2 3 4 5 6 Gilbert, O.L. (1954). "Dryopteris Villarii (Bellardi) Woynar (Aspidium Rigidum Sw.; Lastrea Rigida (Sw.) C. Presl)". Journal of Ecology. 58 (1 1970): 301–13. doi:10.2307/2258184 . Retrieved 9 April 2024.
  6. 1 2 "Dryopteris villarii". floraveg.eu. n.d. Retrieved 9 April 2024.