Dudleya cymosa subsp. paniculata

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Dudleya cymosa subsp. paniculata
Canyon Live-forever imported from iNaturalist photo 15273 on 21 December 2021.jpg
Status TNC T4.svg
Apparently Secure  (NatureServe) [1]
Scientific classification Red Pencil Icon.png
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Order: Saxifragales
Family: Crassulaceae
Genus: Dudleya
Species:
Subspecies:
D. c. subsp. paniculata
Trinomial name
Dudleya cymosa subsp. paniculata
(Jeps.) K.M.Nakai
Synonyms [2]
  • Cotyledon caespitosa var. paniculataJeps.
  • Dudleya humilisBritton & Rose
  • Dudleya paniculataBritton & Rose

Dudleya cymosa subsp. paniculata, known by the common name Diablo Range dudleya, [3] is a species of perennial succulent plant in the family Crassulaceae native to the Inner South Coast Range of California. It is characterized by pale yellowish flowers, oblong to oblanceolate leaves and a growth habit not limited to a single substrate. It is closely related to Dudleya abramsii subsp. setchellii. [4]

Contents

Description

This plant is not usually caespitose, rosettes are solitary to few. The rosettes are 4 to 11 cm wide. The stems are 1 to 2 cm wide. The leaves are 3 to 10 cm, 5 to 20 mm wide, shaped oblong, oblanceolate to lance-oblong, green to more or less white, and the foliage may or may not be glaucous, leaf tip acute. The peduncle is 5 to 25 cm long, and the inflorescence contains 2 to 3 branches that rebranch once or twice. The terminal branches are 1 to 5 cm long, with 4 to 10 flowers, on pedicels 3 to 12 mm long. The petals are 1.5 to 2.5 mm wide, colored a pale yellow to white-yellow, or pale-yellow pink in the San Joaquin Valley. [5]

Flowering is from May to June. [5]

Taxonomy

Taxonomic history

This plant was first described as Cotyledon caespitosa var. paniculata by Jepson in 1901. [6] In Nathaniel Lord Britton and Joseph Nelson Rose's reorganization of North American Crassulaceae in 1903, they described this plant as Dudleya paniculata and Dudleya humilis. [7] In the 1950s, Reid Moran regarded Cotyledon caespitosa var. paniculata as a synonym of Dudleya cymosa subsp. setchellii, although he noted that they could be separated on the basis of leaf shape. Botanist Kei M. Nakai later again separated paniculata as Dudleya cymosa subsp. paniculata, and reduced Dudleya humilis as a synonym for subsp. paniculata plants that have become edaphic dwarfs. [2]

Cladogram of Dudleya cymosa subsp. paniculata
Virens clade

D. abramsii subsp. setchellii

D. cymosa subsp. paniculata

D. cymosa subsp. cymosa

Characteristics

Dudleya cymosa subsp. paniculata can be differentiated from its close relative, [4] Dudleya abramsii subsp. setchellii, in a number of ways. D. c. ssp. paniculata plants have oblong to oblanceolate rosette leaves rather than oblong-triangular leaves, an inflorescence of 2 to 3 branches that rebranch once or twice rather than an inflorescence of 2 to 3 branches that do not rebranch, and pedicels that are 6 to 12 mm long against pedicels that are 4 to 7 mm long. D. a. ssp. setchellii is also only restricted to serpentine rock outcrops, while this species occurs on a variety of substrates. [2]

Distribution and habitat

This species occurs throughout the Inner South Coast Ranges, the eastern San Francisco Bay Area, and mountainous portions of the San Joaquin Valley. [5] It is particularly conspicuous on Mount Diablo and Mount Hamilton. It is found on rocky outcrops and in canyons. [2]

See also

Related Research Articles

<i>Dudleya</i> North American succulent genus

Dudleya, commonly known as liveforevers is a genus of succulent plants in the stonecrop family, Crassulaceae, consisting of about 68 taxa in southwestern North America and Guadalupe Island. The species come in multiple forms, some large and evergreen, others cryptic and deciduous. The flowers of Dudleya have parts numbered in 5, and when fruiting are filled with tiny, ovoid or crescent-shaped seeds.

<i>Dudleya abramsii <span style="font-style:normal;">subsp.</span> setchellii</i> Species of succulent

Dudleya abramsiisubsp. setchellii, known by the common name Dudleya setchellii, the Santa Clara Valley dudleya, or Santa Clara Valley liveforever, is a member of the Dudleya genus of succulent perennials, members of the family Crassulaceae. The Santa Clara Valley dudleya, endemic to the Santa Clara Valley region in the southern San Francisco Bay Area, was listed on 3 February 1995, as an endangered species. It is considered to be a subspecies of Dudleya abramsii, but its taxonomic status is still unclear. Its closest relative is Dudleya cymosa subsp. paniculata, which is a morphologically similar sister taxon.

<i>Dudleya cymosa</i> Species of succulent

Dudleya cymosa is a species complex of evergreen and deciduous succulent plants in the family Crassulaceae known by the common name: canyon liveforever. It is a loosely defined polyphyletic species with a diverse number of subspecies, varying highly in morphology, distribution, and habitat.

<i>Dudleya blochmaniae</i> Species of deciduous succulent plant from North America

Dudleya blochmaniae is a summer-deciduous succulent plant known by the common names Blochman's liveforever or Blochman's dudleya. This species of Dudleya survives part of the year with no aboveground presence, surviving as underground corm-like roots in deciduous months. It is characterized by white, star-shaped and spreading flowers that emerge after sufficient rainfall. It is found along the Pacific coast of the California Floristic Province, from the vicinity of San Luis Obispo in California to Punta Colonet in Baja California.

<i>Dudleya caespitosa</i> Species of coastal succulent plant from North America

Dudleya caespitosa is a succulent plant known by several common names, including sea lettuce, sand lettuce, and coast dudleya. It is endemic to California, where it grows along the coastline in the southern half of the state. Taxonomically, this species is a highly variable complex of polymorphic and polyploid plants, closely related to numerous neighboring species such as Dudleya farinosa, Dudleya greenei and Dudleya palmeri. It is delimited from neighboring species on an arbitrary basis of distribution and chromosome number, and may not be immediately separable from the other species it approaches.

<i>Dudleya abramsii <span style="font-style:normal;">subsp.</span> calcicola</i> Species of succulent

Dudleya abramsiisubsp.calcicola is a succulent plant known by the common name limestone liveforever, or limestone dudleya. It is endemic to California, where it is a rare resident of limestone outcroppings and rocky slopes in the southern Sierra Nevada and nearby mountains and foothills. It was formerly regarded as Dudleya calcicola.

<i>Dudleya candelabrum</i> Species of succulent

Dudleya candelabrum is a species of succulent plant known by the common names candleholder liveforever or candleholder dudleya. Endemic to California, this species grows wild only on the northern Channel Islands, where it is found in open rocky places and north-facing slopes. It is characterized by thin, spade-shaped green leaves and an inflorescence covered in long, reflexed bracts, with pale yellow flowers. It has been threatened by poachers shipping plants to South Korea.

<i>Dudleya palmeri</i> Species of succulent

Dudleya palmeri is a species of succulent plant in the family Crassulaceae known by the common name Palmer's liveforever. This Dudleya is endemic to California where it grows along the coast. It is characterized by orange to red over yellow or pink flowers. It is a polyploid species that closely resembles Dudleya lanceolata but has a coastal habit, and hybridizes with Dudleya caespitosa and Dudleya cymosa.

<i>Dudleya greenei</i> Species of succulent

Dudleya greenei is a perennial species of succulent plant known by the common names Greene's liveforever, or Greene's dudleya. It is endemic to the Channel Islands of California, where it grows along the cliffs of four of the eight islands. It is a highly variable plant, presenting with multiple forms and varying levels of ploidy. Taxonomically, this species is an insular segregate of Dudleya caespitosa, and was placed as a stopgap taxon by Reid Moran in his 1951 thesis on the genus. It is characterized by white or green leaf rosettes, loomed over by inflorescences bearing pale yellow to white flowers. It is a member of the subgenus Dudleya, as it cannot be propagated from leaf cuttings, does not grow from a corm, and has tight petals.

<i>Dudleya abramsii</i> Species of succulent

Dudleya abramsii is a species complex of succulent plants native to California and parts of Baja California. There are numerous subspecies, some critically endangered, with varying habits and lifestyles, but most often characterized by a smaller size, yellow flowers, and an affinity for rocky habitats. The subspecies may be polyphyletic.

<i>Dudleya verityi</i> Species of succulent

Dudleya verityi is a rare species of succulent plant known by the common name Verity's liveforever. It is endemic to Ventura County, California, where it is known from only three occurrences in the vicinity of Conejo Mountain between Camarillo and Thousand Oaks. It probably occurs in a few additional locations nearby which have not yet been officially vouchered.

<i>Dudleya attenuata</i> Species of succulent

Dudleya attenuata is a species of perennial succulent plant known by the common name taper-tip liveforever, native to Baja California and a small portion of California. A rosette-forming leaf succulent, it has narrow pencil shaped leaves that can often be found covered in a white epicuticular wax. The thin, sprawling stems branch to form the clusters of rosettes, with plants creating a "clump" up to 40 cm wide. The small flowers are white or yellow, with 5 spreading petals. It is a diverse, variable species that extends from the southernmost coast of San Diego County to an area slightly north of the Vizcaino Desert, hybridizing with many other species of Dudleya in its range. Some plants with white or pinkish flowers were referred to as Orcutt's liveforever, referring to a former subspecies split on the basis of the flower color.

Dudleya formosa, known by the common name La Misión liveforever, is a species of perennial succulent plant endemic to the Guadalupe Valley in Baja California. It is characterized by bright green leaves, red floral stems, and pink flowers.

<i>Dudleya abramsii <span style="font-style:normal;">subsp.</span> abramsii</i> Subspecies of succulent plant

Dudleya abramsii subsp. abramsii is a species of succulent plant in the family Crassulaceae known by the common name as Abrams' liveforever. It is a small, delicate plant found growing among rocks, and is characterized by yellow flowers with a red tinge that emerge from May to July. It is native to the southern Sierra Nevada of California and the Peninsular Ranges across both the United States and Mexico.

<i>Dudleya abramsii <span style="font-style:normal;">subsp.</span> affinis</i> Subspecies of plant

Dudleya abramsii subsp. affinis, commonly known as the San Bernardino Mountains Dudleya, is a subspecies of succulent plant endemic to a portion of the San Bernardino Mountains in California. It is a plant with an unbranching stem and elliptical leaves occurring in Green Canyon and the nearby plateaus.

Dudleya cymosasubsp. costatifolia, known commonly as the Pierpoint Springs dudleya or the Pierpoint Springs liveforever, is a species of succulent plant in the family Crassulaceae, narrowly endemic to a locality in Tulare County, California, United States. It is a clumping plant with small rosettes and bright yellow flowers, resembling alpine "cushion plants."

<i>Dudleya cymosa <span style="font-style:normal;">subsp.</span> pumila</i> Subspecies of plant

Dudleya cymosasubsp. pumila, most commonly known as the low canyon dudleya, chalky canyon dudleya or California live-forever, is a species of perennial succulent plant. It has diamond to spoon shaped leaves, sometimes coated with a fine white powder, and in May through July, bright red, orange or yellow flowers adorn the short inflorescence. A leaf succulent primarily found growing in rocky cliffs and slopes, it is endemic to California, and grows in the Transverse Ranges and South Coast Ranges, with some outlying populations. A variable plant, in some localities it is difficult to distinguish from other plants in the genus.

<i>Dudleya cymosa <span style="font-style:normal;">subsp.</span> cymosa</i> Subspecies of succulent perennial plant

Dudleya cymosasubsp. cymosa is a species of succulent perennial plant in the family Crassulaceae native to California. It is the autonymous subspecies for Dudleya cymosa, and is known by the common name canyon liveforever. It is native to the California Coast Ranges, the Sierra Nevada and the Santa Monica Mountains. It is characterized by bright-yellow, orange or red flowers and broad, wide leaves. This plant is commonly found growing on rocky outcrops, talus slopes, and in shaded canyons.

Dudleya virens subsp. extima is a subspecies of succulent plant in the family Crassulaceae commonly known as the Guadalupe green liveforever. It is a rosette-forming leaf succulent, with both green and white waxy foliage. It has white flowers with spreading petals that bloom from May to June. It is a somewhat small plant, continuing a southward trend of decreasing size relative to other Dudleya virens subspecies. This plant is endemic to Guadalupe Island in the eastern Pacific Ocean, which is 241 kilometers off of the Baja California coast. It is very rare, with this plant only surviving on sheer cliff faces and canyons, out of the reach of the former feral goat population rampant on the island. It closely resembles a miniature version of Dudleya virens subsp. virens, but it may be more nearly related to the local Dudleya guadalupensis.

<i>Dudleya cymosa <span style="font-style:normal;">subsp.</span> agourensis</i> Subspecies of flowering plant

Dudleya cymosasubsp. agourensis, commonly known as the Agoura Hills dudleya or Agoura Hills liveforever, is a species of perennial succulent plant. A rare and critically imperiled species from the Santa Monica Mountains in California, it is characterized by glaucous, gray-purple leaves, bright yellow flowers, and ascending bracts. A leaf succulent, it is found growing on west to northwest-facing open, rocky volcanic slopes.

References

  1. "Dudleya cymosa ssp. paniculata". NatureServe Explorer. NatureServe. 2021. Retrieved 17 December 2021.
  2. 1 2 3 4 Nakai, Kei M. (1987). "Some New and Reconsidered California Dudleya (Crassulaceae)". Madroño. 34 (4): 338–339.
  3. "Dudleya cymosa ssp. paniculata". Lady Bird Johnson Wildflower Center. The University of Texas at Austin. 2021-10-29. Archived from the original on 2021-12-17. Retrieved 17 December 2021.
  4. 1 2 Yost, J. M.; Bontrager, M.; McCabe, S. W.; Burton, D.; Simpson, M. G.; Kay, K. M.; Ritter, M. (2013). "Phylogenetic relationships and evolution in Dudleya (Crassulaceae)" (PDF). Systematic Botany. 38 (4): 1096–1104. doi:10.1600/036364413X674760. S2CID   15715233.
  5. 1 2 3 McCabe, Stephen Ward (2012). "Dudleya cymosa subsp. paniculata". Jepson eFlora. Jepson Flora Project (eds.). Archived from the original on 2017-05-08. Retrieved 17 December 2021.
  6. A Flora of Western Middle California. Berkeley, California: Encina Publishing Company. 1901. p. 267. OCLC   663440141.
  7. Rose, Joseph Nelson (1903). "New and noteworthy North American Crassulaceae". Bulletin of the New York Botanical Garden. 3 (9).