La Laguna liveforever | |
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The watercolor illustration by Frederick Andrews Walpole | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Order: | Saxifragales |
Family: | Crassulaceae |
Genus: | Dudleya |
Species: | D. rigida |
Binomial name | |
Dudleya rigida | |
Distribution of Dudleya rigida within Baja California Sur | |
Synonyms | |
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Dudleya rigida is a species of succulent perennial plant in the family Crassulaceae known commonly as the La Laguna liveforever. Characterized by a tall inflorescence with pendant yellowish-red flowers, it is a very rare plant whose existence was doubtful until botanist Reid Moran accidentally re-discovered it. It is endemic to the highest peaks of the Sierra de la Laguna in Baja California Sur, Mexico. [1]
Dudleya rigida has a caudex 1 to 3.5 cm thick, obscured by dried leaves, and branching into clusters with up to 20 rosettes. Rosettes are 6 to 15 cm wide, with 10-25 leaves. Leaves are green or somewhat glaucous, and oblong to triangular-ovate, short-acuminate. The leaves are 5 to 8 cm long, 2.5 to 4 cm wide, and 6 to 10 mm thick. The inflorescence has 8-21 flowers, with a yellow corolla marked in red and a pink to red peduncle. The flowers are pendent, a trait only shared by Dudleya anthonyi and Dudleya pulverulenta .
Compared with Dudleya nubigena , which has an overlapping range, Dudleya rigida has a thicker caudex, oblong leaves, a taller inflorescence, and is green, not farinose. The pollen grains are much larger than D. nubigena, because D.rigida possesses a much higher chromosome count. The plant is 16-ploid with a count of 136 chromosomes. [2]
Joseph Nelson Rose named Dudleya rigida from a living specimen brought from Baja California by a J. E. McClelland. In his treatment, Rose gave no exact locality or date. The plant flowered in Washington by June 1897, but he never elaborated as to why he named it rigida. According to Edward W. Nelson, a J. Ellis McLellan collected birds and mammals in Baja California for the U.S. Biological Survey from March 31 to September 1895, visiting numerous locations of the Baja cape. Reid Moran inferred that based on the name of the collector, rigida must have originated somewhere in the Cape Region or the surrounding islands and mountains. [2]
Pressed specimens were boiled and arranged in a way that obfuscated the true layout of the inflorescence. Fortunately, Frederick Andrews Walpole created a watercolor illustration of a living plant, albeit from a greenhouse collection. Curious about the existence of the plant, which had little documentation since Rose's treatment of the Dudleya genus, Reid Moran, on a journey to the Sierra de la Laguna, rediscovered the plant. Initially confused about the accidental discovery of a Dudleya on the 1900 meter high Cerro la Aguja, Moran recalled the existence of the Walpole plate and made the connection. [2]
Moran managed to obtain more information about McLellan's journey to the cape, as an ornithologist with the USFWS sent him the handwritten notes, which mentioned that he had also explored the area near the Cerro la Aguja. Although McLellan never wrote about the collection of his specimen, Moran was able to confirm that the plants he discovered were the same as Rose's description. Moran did previously identify herbarium specimens of Dudleya rigida as being synonymous with Dudleya nubigena , but after his encounter with the plants in the wild, he changed his mind. [2]
Dudleya rigida is native to the Sierra de la Laguna, growing on rocky outcroppings alongside Agave promontorii , Nolina beldingii, and Myrtillocactus cochal. [2] [3]
Montane species of Dudleya:
Dudleya pulverulenta is a species of perennial succulent plant known by the common names chalk lettuce, chalk dudleya, and chalk liveforever. It is one of the largest Dudleya, with a silvery, waxy rosette that may greatly contrast with its habitat. It is also regarded as one of the most distinctive members of the Dudleya, with one of the most specialized inflorescences in the genus, adapted to hummingbird pollination through its red pendent flowers, the longest corolla, and the highest nectar output. Dudleya pulverulenta has the largest range of all Dudleya, over 1,000 kilometres (620 mi), being found from southern Monterey County in California to the Sierra de San Borja in southern Baja California. It is closely related to Dudleya arizonica, a smaller desert species that tends to lack the specialized floral traits, and Dudleya anthonyi, which differs in a few morphological traits and is restricted to the San Quintín Volcanic Field.
Dudleya greenei is a perennial species of succulent plant known by the common names Greene's liveforever, or Greene's dudleya. It is endemic to the Channel Islands of California, where it grows along the cliffs of four of the eight islands. It is a highly variable plant, presenting with multiple forms and varying levels of ploidy. Taxonomically, this species is an insular segregate of Dudleya caespitosa, and was placed as a stopgap taxon by Reid Moran in his 1951 thesis on the genus. It is characterized by white or green leaf rosettes, loomed over by inflorescences bearing pale yellow to white flowers. It is a member of the subgenus Dudleya, as it cannot be propagated from leaf cuttings, does not grow from a corm, and has tight petals.
Dudleya variegata is a deciduous succulent plant in the family Crassulaceae known by the common names variegated liveforever, variegated dudleya or rarely San Diego Hasseanthus. A cryptic plant that survives part of the year dormant underground from starch reserves in a corm, after sufficient rainfall, leaves will emerge, soon giving way to small inflorescences with yellow star-shaped flowers. It is native to Baja California and adjacent San Diego County in California, where it grows in several habitat types, including chaparral and vernal pools.
Dudleya attenuata is a species of perennial succulent plant known by the common name taper-tip liveforever, native to Baja California and a small portion of California. It is a rosette-forming leaf succulent which has narrow pencil-shaped leaves that can often be found covered in a white epicuticular wax. The thin, sprawling stems branch to form the clusters of rosettes, with plants creating a "clump" up to 40 cm wide. The small flowers are white or yellow, with 5 spreading petals. It is a diverse, variable species that extends from the southernmost coast of San Diego County to an area slightly north of the Vizcaino Desert, hybridizing with many other species of Dudleya in its range. Some plants with white or pinkish flowers were referred to as Orcutt's liveforever, referring to a former subspecies split on the basis of the flower color.
Dudleya ingens is a species of perennial succulent plant in the family Crassulaceae commonly known as the rock liveforever or Baja liveforever. A relatively large member of the genus Dudleya, this species has long green succulent leaves, and in April to June is characterized by pale yellow to white pink-tinged flowers topping tall, reddish inflorescences. It has a stem clothed densely with old, leathery leaves, and the inflorescence may be nodding, with the floral branches bearing the flowers tending to unfurl like the fronds of a fern. It is similar in appearance to Dudleya brittonii, but differs in range and chromosome number. This species is endemic to the state of Baja California in Mexico, being found from Santo Tomás to the southern coast of the state.
Dudleya guadalupensis is a very rare species of succulent perennial plant in the family Crassulaceae commonly known as the Guadalupe liveforever. It is a rosette-forming leaf succulent, with foliage that is variously colored light green, green, and a waxy white. It is characterized by dense leaves that fold over the center in dormancy, a curving, sinuous flower stalk, and white, cup-shaped flowers. It is endemic to the rocks and islets off of Guadalupe Island, an isolated volcanic island in the Pacific Ocean located 241 kilometers off of the coast of Baja California.
Dudleya pauciflora is a species of succulent plant in the stonecrop family known by the common name few-flower liveforever. It is characterized by its small crowded rosettes of narrow leaves and its colorful inflorescence with red-yellow flowers. Found growing on rocky outcrops and cliffs in the high elevation mountains of the Sierra de San Pedro Martir and the Sierra de San Borja, it is endemic to the state of Baja California, Mexico.
Dudleya gatesii, known by the common name as Gates'liveforever, is a species of perennial succulent plant in the family Crassulaceae. It is native to the central desert of the Baja California peninsula, found growing along dry and rocky outcroppings. It is characterized by red inflorescences topped with white flowers, bracts that are often reflexed downwards, and a rosette of dark-green leaves turning reddish.
Dudleya campanulata is a species of perennial succulent plant known by the common name as the Punta Banda liveforever, native to Baja California and endemic to the Punta Banda peninsula, a promontory south of Ensenada that encloses the southern limit of the Bahía de Todos Santos, a deepwater bay. One of many species of Dudleya native to the peninsula and surrounding islands, it is distinguished by its campanulate flowers and its occupation of a narrow habitat that consists of ocean bluffs on the southern end of the Punta Banda, near the well-known blowhole La Bufadora.
Dudleya pachyphytum is an insular succulent plant known by the common name Cedros Island liveforever. It is a member of the genus Dudleya, in the family Crassulaceae. Characterized by thick, blunt leaves covered in a white, powdery wax and adorned by white flowers in bloom, it is regarded as one of the most attractive and charismatic members of its genus. It is endemic to the foggy northern end of Mexico's Cedros Island, occupying an ecological niche shared with the Cedros Island Pine.
Dudleya formosa, known by the common name La Misión liveforever, is a species of perennial succulent plant endemic to the Guadalupe Valley in Baja California. It is characterized by bright green leaves, red floral stems, and pink flowers.
Dudleya rubens is a species of succulent perennial plant in the family Crassulaceae known by the common name as the San Francisco liveforever, native to the mountains of Baja California Sur. It is a rosette-forming plant with waxy leaves, characterized by branching stems and dull red to apricot flowers. It is only found above 500 metres (1,600 ft) in the Sierra de San Francisco and the Sierra de la Giganta ranges in Baja California Sur, primarily on north-facing volcanic slopes.
Dudleya cultrata is a species of perennial succulent in the family Crassulaceae commonly known as the knife-leaved liveforever or the maritime succulent liveforever. This species is characterized by oblong, narrow green leaves and flowers with pale yellow petals that bloom from April to June. Although similar to Dudleya ingens, this species is most often seen growing sympatric with the larger, wax-covered Dudleya anthonyi. It is native to Baja California, occurring on the coast from Punta Colonet and San Quintin to El Rosario.
Dudleya candida is a species of perennial succulent plant in the family Crassulaceae known by the common names Coronados liveforever or chalk rose. It is a rosette-forming, green to white-colored leaf succulent, and in bloom yellow flowers atop red stalks stand above the foliage. It has some visual similarities to the mainland Dudleya brittonii, and has found uses in horticulture as an ornamental plant. It is restricted to the Coronado Islands, an island group off of the extreme northern Baja California coast, visible from the United States.
Dudleya rigidiflora is a very rare species of succulent perennial plant known by the common name Playa Maria liveforever, endemic to the coast of southwestern Baja California.
Dudleya nubigena is a species of succulent plants in the family Crassulaceae. It is a rosette forming perennial with flattish leaves. Endemic to southern Baja California Sur, the species is found in the Sierra de la Laguna and the surrounding lowlands, a small southern portion of the Sierra de la Giganta, and on Isla Espíritu Santo, with a subspecies endemic to Cerralvo Island.
Dudleya albiflora, known by the common name white-flower liveforever, is a species of succulent perennial plant in the family Crassulaceae. It is native to the Baja California Peninsula. This species represents numerous populations with varying chromosome numbers scattered around the peninsula, but all share broad, common morphological traits such as white flowers and narrow leaves.
Dudleya saxosasubsp. aloides is a species of perennial succulent plant in the family Crassulaceae known by the common names desert dudleya or desert savior. It is a rosette-forming species widely distributed throughout the Peninsular Ranges and desert mountains of California in the United States. It is characterized by bright-yellow or greenish-yellow flowers, and can be found in shaded crevices and slopes. Plants in western half of the range may grade into Dudleya lanceolata.
Nolina beldingii is a species of perennial flowering plant in the family Asparagaceae known commonly as the Cape nolina or Belding's beargrass. It is an arborescent monocot growing up to 7 metres (23 ft) high, with fissured bark on a trunk topped with leaf rosettes. The narrow leaves are up to 1.15 m (3.8 ft) long, and are used as thatching by local peoples. This species is endemic to Baja California Sur in Mexico, where it grows only in the highest reaches of the Sierra de la Laguna. It is found primarily in oak forests at elevations over 1,000 m (3,300 ft) along rocky granite outcrops.
Dudleya virens subsp. extima is a subspecies of succulent plant in the family Crassulaceae commonly known as the Guadalupe green liveforever. It is a rosette-forming leaf succulent, with both green and white waxy foliage. It has white flowers with spreading petals that bloom from May to June. It is a somewhat small plant, continuing a southward trend of decreasing size relative to other Dudleya virens subspecies. This plant is endemic to Guadalupe Island in the eastern Pacific Ocean, which is 241 kilometers off of the Baja California coast. It is very rare, with this plant only surviving on sheer cliff faces and canyons, out of the reach of the former feral goat population rampant on the island. It closely resembles a miniature version of Dudleya virens subsp. virens, but it may be more nearly related to the local Dudleya guadalupensis.