Elgin Reptiles is the name given to the Permian and Triassic fossils found in the sandstone deposits in and around the town of Elgin, in Moray, Scotland. They are of historical and scientific importance, and many of the specimens are housed in the Elgin Museum, and some in the Hunterian in Glasgow, and the National Museum of Scotland in Edinburgh. The Elgin Reptiles include the dinosauriform Saltopus elginensis, [1] the dicynodont Gordonia , [2] and the pareiasaur Elginia . [3] There are also many footprints and tail-drags associated with the same Permian and Triassic sandstone deposits. [4] [5]
The sandstone in the Elgin area was originally quarried for building materials. The quarries were where the first reptile fossils were found, and they have continued to yield fossils to this day. [5] The first Elgin Reptile was discovered in 1844, but because it was only a few scales scientists of the time believed it was an Old Red Sandstone fish fossil, [5] which were relatively well-known from sites across Northern Scotland. In 1851 a second indisputably reptilian fossil was found. Fossils of Gordonia were first found by Scottish naturalist Ramsay Heatley Traquair in 1885 and E.T. Newton described several key fossil specimens. [2]
Elgin Museum is Scotland's oldest independent museum. [6] The Elgin Reptiles, footprints, and older fish fossils held by the Elgin Museum are “recognised” by the Scottish Government as a Collection of National Significance due to the major part played by the Museum’s founders and associated geologists in helping answer questions about geological succession that challenged 19th century naturalists. The Museum has many original papers, including letters from Hugh Miller, and correspondence with Charles Darwin. [3]
The land North of Elgin consists of fragments of Permian, Triassic and Jurassic bedrocks on the edge of the Moray Firth Basin, south of the Great Glen Fault. The Elgin Reptiles come from the Permian and Triassic rocks. These are called the Hopeman Sandstone Formation and the Burghead Sandstone Formation, both part of the New Red Sandstone Supergroup. [7] Some marine reptiles have been found in a glacial erratic of Jurassic strata. Most of the sandstones are wind-blown desert dune deposits, but there are some river deposits and the later Jurassic deposits include mudstones. [8] These rocks overly much older Devonian lake deposits from Lake Orcadie. The intervening Carboniferous sediments were eroded before the Permian deposition, leaving a 100 million year gap in the stratigraphy. This unusual sequence of rocks confused 19th century geologists for many years. The Moray Firth Basin has been exploited for offshore oil deposits. Scotland lay at about 20 degrees North in the Late Permian-Early Triassic, and was moving North. [8] [5] Permian, Triassic and Jurassic rocks are rare in Scotland, but some can be found along the Northern edge of the Moray Firth, and in the Inner Hebrides, particularly on the Isle of Skye. The Hopeman Sandstone Formation quarry at Clashach is designated as a SSSI (Site of Special Scientific Importance) for its fossils.
Many of the Elgin Reptile fossils contain no actual bone: they are often voids in the sandstone that used to contain bone, but are now virtually empty. This made them difficult to identify and study. In the 19th century until the late 20th century, the most common method to explore them was to infill the void, then break the surrounding rock. [4] [5] [9] More recently, CT-scanning has been used to see inside the rock and reconstruct the fossils digitally. These can also be printed or cast as models to facilitate study and as museum pieces. Elgin Museum has such a cast of the dicynodont Gordonia . [3] [4] The reptiles of Cutties Hillock, Quarrywood and the Hopeman Sandstones are Late Permian, around 250 million years old, while most other reptile fossils are found in the early Triassic rocks of Lossiemouth, Spynie and Findrassie, and are about 220 million years old. [5] The fossils are “recognised” by the Scottish Government as a Collection of National Significance. [3] Many of the type and significant specimens are on display, with more fossils and archives in the collection at Elgin Museum, available for study and research.
More recent fossil discoveries from Elgin have been footprints, with over 200 found prior to 1997. [4] These range in size from 0.5 cm to 24 cm wide, and most belong to therapsids. [4] [5] Numerous taildrags have also been recovered and studied, many from therapsids. [4]
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Elgin Reptiles: Archosaurs | ||||
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Taxa | Presence | Location | Description | Images |
| Triassic | Lossiemouth, Spynie, and Findrassie Scotland | A 1.2-metre-long (3.9 ft) extinct genus of pseudosuchian originally thought to be the ancestor to carnosaurian dinosaurs, but now known to be more closely related to crocodilians than to dinosaurs. | |
| Triassic | Lossiemouth, Spynie, and Findrassie Scotland | A quadrupedal armoured aetosaur, around 2 m long, with a very small head for its size (25 cm). It had a beak-like face that arched upwards, and is believed to have used this to uproot plants in a similar manner to a modern pig. The peg-like molar teeth would have been suitable for chewing tough vegetation. | |
| Triassic | Lossiemouth, Scotland | This small cursorial archosaur (18 cm long) is of uncertain phylogenetic position. It is possibly the basal-most ornithodiran, the sister-taxon to Pterosauria, or a basal member of Avemetatarsalia that lies outside of Ornithodira. | |
| Triassic | Lossiemouth, Scotland | A pseudosuchian, its phylogenetic position is debated. | |
| Triassic | Lossiemouth, Scotland | Was thought to be Scotland's earliest dinosaur, but research by Professor M. Benton instead identified it as dinosauriform – a forerunner of the dinosaurs. [3] [1] | |
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Elgin Reptiles: Rhynchosaurs | ||||
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Taxa | Presence | Location | Description | Images |
Triassic | Lossiemouth and Spynie, Scotland | A stocky animal around 1.3 metres long, with a beak and heavy teeth suggesting a herbivorous diet. | ||
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Elgin Reptiles: Sphenodontids | ||||
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Taxa | Presence | Location | Description | Images |
Triassic | Lossiemouth, Scotland | It is related to the extant tuatara. | ||
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Elgin Reptiles: Procolophonids | ||||
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Taxa | Presence | Location | Description | Images |
| Triassic | Lossiemouth and Spynie, Scotland | First described by Sir Richard Owen, it was a small, lizard-like animal (27 cm) with a long tail. | |
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Elgin Reptiles: Dicynodonts | ||||
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Taxa | Presence | Location | Description | Images |
| Permian | Cutties Hillock, Scotland | An extinct therapsid. It was formerly assigned to the wastebasket taxon Dicynodon as Dicynodon traquairi. Re-study of the specimen using CT-scanning has allowed this taxon to be more formally described and it was renamed Gordonia traquairi. [4] | |
| Permian | Cutties Hillock, Scotland | An extinct therapsid notable for its unusual square-shaped skull. | |
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Elgin Reptiles: pareiasaurs | ||||
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Taxa | Presence | Location | Description | Images |
| Permian | Cutties Hillock, Scotland | This animal's distinctive skull ornamentation make it instantly recognisable. It has recently been incorporated into the logo for the vertebrate fossil research and preservation group, Pal Alba. [10] |
Synapsida is one of the two major clades of vertebrate animals in the group Amniota, the other being the Sauropsida. The synapsids were the dominant land animals in the late Paleozoic and early Mesozoic, but the only group that survived into the Cenozoic are mammals. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name. The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals.
Therapsida is a clade comprising a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, resulting in a more "standing" quadrupedal posture, as opposed to the lower sprawling posture of many reptiles and amphibians.
Saltopus is a genus of very small bipedal dinosauriform containing the single species Saltopus elginensis from the late Triassic period of Scotland. It is one of the most famous Elgin Reptiles.
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivores that typically bore a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers.
The geology of Scotland is unusually varied for a country of its size, with a large number of different geological features. There are three main geographical sub-divisions: the Highlands and Islands is a diverse area which lies to the north and west of the Highland Boundary Fault; the Central Lowlands is a rift valley mainly comprising Palaeozoic formations; and the Southern Uplands, which lie south of the Southern Uplands Fault, are largely composed of Silurian deposits.
Elginia is an extinct genus of pareiasaurid known from the Late Permian of Scotland and China. It was named for the area around Elgin in Scotland, which has yielded many fossils referred to as the Elgin Reptiles.
The New Red Sandstone, chiefly in British geology, is composed of beds of red sandstone and associated rocks laid down throughout the Permian to the end of the Triassic, that underlie the Jurassic-Triassic age Penarth Group. The name distinguishes it from the Old Red Sandstone which is largely Devonian in age, and with which it was originally confused due to their similar composition.
The Tapinocephalus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the middle Abrahamskraal Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately 2,000 metres (6,600 ft), occur from Merweville and Leeu-Gamka in its southernmost exposures, from Sutherland through to Beaufort West where outcrops start to only be found in the south-east, north of Oudshoorn and Willowmore, reaching up to areas south of Graaff-Reinet. Its northernmost exposures occur around the towns Fraserburg and Victoria West. The Tapinocephalus Assemblage Zone is the second biozone of the Beaufort Group.
The Cistecephalus Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important geological group of the Karoo Supergroup in South Africa. This biozone has outcrops located in the Teekloof Formation north-west of Beaufort West in the Western Cape, in the upper Middleton and lower Balfour Formations respectively from Colesberg of the Northern Cape to east of Graaff-Reinet in the Eastern Cape. The Cistecephalus Assemblage Zone is one of eight biozones found in the Beaufort Group, and is considered to be Late Permian in age.
The Tropidostoma Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the lower Teekloof Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately 240 metres (790 ft), occur from east of Sutherland through to Beaufort West and Victoria West, to areas south of Graaff-Reinet. Its northernmost exposures occur west/north-west of Colesberg. The Tropidostoma Assemblage Zone is the fourth biozone of the Beaufort Group.
The Fremouw Formation is a Triassic-age rock formation in the Transantarctic Mountains of Antarctica. It contains the oldest known fossils of tetrapods from Antarctica, including synapsids, reptiles and amphibians. Fossilized trees have also been found. The formation's beds were deposited along the banks of rivers and on floodplains. During the Triassic, the area would have been a riparian forest at 70–75°S latitude.
Eodicynodon is an extinct genus of dicynodont therapsids, a highly diverse group of herbivorous synapsids that were widespread during the middle-late Permian and early Triassic. As its name suggests, Eodicynodon is the oldest and most primitive dicynodont yet identified, ranging from the middle to late Permian and possessing a mix of ancestral anomodont/therapsid features and derived dicynodont synapomorphies.
Venyukovia is an extinct genus of venyukovioid therapsid, a basal anomodont from the Middle Permian of Russia. The type and sole species, V. prima, is known only by a partial lower jaw with teeth. Venyukovia has often been incorrectly spelt as 'Venjukovia' in English literature. This stems from a spelling error made by Russian palaeontologist Ivan Efremov in 1940, who mistakenly replaced the 'y' with a 'j', which subsequently permeated through therapsid literature before the mistake was caught and corrected. Venyukovia is the namesake for the Venyukovioidea, a group of small Russian basal anomodonts also including the closely related Otsheria, Suminia, Parasuminia and Ulemica, although it itself is also one of the poorest known. Like other venyukovioids, it had large projecting incisor-like teeth at the front and lacked canines, although the remaining teeth are simple compared to some other venyukovioids, but may resemble those of Otsheria.
Geikia is an extinct genus of dicynodont therapsids from the late Permian. The abundance and diversity of dicynodonts during this period, combined with incomplete or inadequately prepared specimens, have led to challenges in determining relationships within this taxon. Only two species, Geikia locusticeps and Geikia elginensis have been assigned to this genus. While this is the currently accepted classification, fossil record limitations have led to repeated debate on the genus assignments of these species.
The Abrahamskraal Formation is a geological formation and is found in numerous localities in the Northern Cape, Western Cape, and the Eastern Cape of South Africa. It is the lowermost formation of the Adelaide Subgroup of the Beaufort Group, a major geological group that forms part of the greater Karoo Supergroup. It represents the first fully terrestrial geological deposits of the Karoo Basin. Outcrops of the Abrahamskraal Formation are found from the small town Middelpos in its westernmost localities, then around Sutherland, the Moordenaarskaroo north of Laingsburg, Williston, Fraserburg, Leeu-Gamka, Loxton, and Victoria West in the Western Cape and Northern Cape. In the Eastern Cape outcrops are known from Rietbron, north of Klipplaat and Grahamstown, and also southwest of East London.
Gordonia is an extinct genus of dicynodont therapsid from the Late Permian of Scotland. Fossils have been found from the Elgin sandstone of Cutties Hillock Sandstone in Elgin, Moray. These are among the many amniote fossils referred to as the Elgin Reptiles. Gordonia was named in 1893 with four species: G. traquairi, G. duffiana, G. huxleyana, and G. juddiana. Currently, the only recognized species is the type G. traquairi. All other species are considered synonyms of the type.
The Hopeman Sandstone Formation is a geologic formation in Scotland. It preserves fossil footprints and body fossils from the Guadalupian Epoch in the Late Permian, to the Early Triassic, It preserves fossils and fossil footprints from various extinct animals such as pareiasaurs and dicynodonts, which are collectively often referred to as the Elgin Reptiles.
Pentasaurus is an extinct genus of dicynodont of the family Stahleckeriidae, closely related to the well known Placerias. It was found in the Lower Elliot Formation of South Africa, dated to the Norian of the Late Triassic period. The genus contains the type and only species, Pentasaurus goggai. Pentasaurus is named after the ichnogenus Pentasauropus, fossil footprints that were originally described from the lower Elliot Formation in 1970 decades before the body fossils of Pentasaurus itself were recognised. Pentasauropus footprints were likely made by dicynodonts, and in South Africa Pentasaurus itself was the likely trackmaker. The name reflects the fact that a large dicynodont was predicted to have existed in the lower Elliot Formation before any body fossils were recognised, and so Pentasaurus was named after its probable footprints. This is a reversal of the more typical occurrence where fossil footprints are named after their presumed trackmakers. The name of the species honours its collector Alfred Brown, nicknamed "Gogga", which means "bug" in Afrikaans.
The geology of Denmark includes 12 kilometers of unmetamorphosed sediments lying atop the Precambrian Fennoscandian Shield, the Norwegian-Scottish Caledonides and buried North German-Polish Caledonides. The stable Fennoscandian Shield formed from 1.45 billion years ago to 850 million years ago in the Proterozoic. The Fennoscandian Border Zone is a large fault, bounding the deep basement rock of the Danish Basin—a trough between the Border Zone and the Ringkobing-Fyn High. The Sorgenfrei-Tornquist Zone is a fault-bounded area displaying Cretaceous-Cenozoic inversion.
The Teekloof Formation is a geological formation that forms part of the Beaufort Group, one of the five geological groups that comprises the Karoo Supergroup in South Africa. The Teekloof Formation is the uppermost formation of Adelaide Subgroup deposits West of 24ºE and contains Middle to Late Permian-aged deposits and four biozones of the Beaufort Group. It overlies the Abrahamskraal Formation. The Teekloof Formation does not underlie other units other than the younger Karoo dolerites and sills that relate to the emplacement of the Early Jurassic Drakensberg Group to the east. Outcrops and exposures of the Teekloof Formation range from Sutherland through the mountain escarpments between Fraserburg and Beaufort West. The northernmost localities of the Teekloof Formation are found by Loxton, Victoria West and Richmond.