Stagonolepis

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Stagonolepis
Temporal range: Carnian
~221–206  Ma
Stagonolepis robertsoni.png
Skeletal restorations of S. robertsoni
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauria
Clade: Pseudosuchia
Order: Aetosauria
Family: Stagonolepididae
Genus: Stagonolepis
Agassiz 1844
Species

Stagonolepis is an extinct genus of stagonolepidid aetosaur known from the Late Triassic (Carnian stage) Hassberge Formation of Germany, the Drawno Beds of Poland, and the Lossiemouth Sandstone of Scotland. Supposed fossils from North and South America have been placed into their own genera, Calyptosuchus and Aetosauroides , respectively. [1] [2]

Contents

Description

Restoration of S. robertsoni Restorations of Stagonolepis robertsoni, after skeletal by Hartman, 2016.jpg
Restoration of S. robertsoni

Stagonolepis robertsoni was about 3 metres (10 ft) long. It was a quadrupedal animal covered in thick armoured scales that ran down the length of its body. A slow-moving browser, it would have used this heavy body armour to repel attacks from contemporary archosauriform carnivores. Stagonolepis had a very small head for its size; it was only 25 centimetres (10 in), accounting for less than 10% of the total body length. It had no teeth in the front of its jaws, but instead had a beak-like tip that arched upwards. This would have allowed it to uproot plants in a similar manner to a modern pig. The peg-like teeth at the back of its mouth would have been suitable for chewing tough vegetation, including horsetails, ferns, and the newly evolved cycads. [3]

A 2018 paper suggested that Stagonolepis olenkae's forelimb morphology is an adaptation for scratch-digging. According to paleontologist Dawid Dróżdż, "S. olenkae might have used its robust forelimbs to break through the compacted soil with its claws and proceed to dig in search of food in softened substrate with the shovel-like expansion at the tip of its snout." [4]

Species

Model of S. olenkae at the Museum of Evolution Warsaw MEPAN aetozaur (stagolepsis).jpg
Model of S. olenkae at the Museum of Evolution Warsaw

Fossil remains of S. robertsoni have been found in Lossiemouth Sandstone of Scotland, while S. olenkae is known from deposits near Krasiejów, Poland. S. olenkae is stratigraphically younger the type species of Stagonolepis, S. robertsoni, [5] although it is questioned by some researchers and described as an example of intraspecific variety (e. g. sexual dimorphism) [6] The genus Aetosauroides from South America has been considered a junior synonym of Stagonolepis by some paleontologists. Two species of Aetosauroides were named, A. scagliai and A. subsulcatus. In 2002, Andrew B. Heckert and Spencer G. Lucas proposed that smaller specimens of both species belong to Stagonolepis robertsoni, and larger specimens to S. wellesi. [7] S. wellesi itself was originally named Calyptosuchus , a stagonolepidid from the Late Triassic Dockum Group of the United States, which was considered to be a species of Stagonolepis by Murray & Long in 1989. However, most of the sequential studies conclude that both Aetosauroides and Calyptosuchus are valid and monotypic genera, the former occurs only in South America and the latter only in the United States. Stagonolepis is restricted to the Carnian stage of Scotland and Poland. [8] [1] [2] [9]

Related Research Articles

<span class="mw-page-title-main">Aetosaur</span> Extinct order of heavily armoured reptiles

Aetosaurs are heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site in which they are found.

<i>Aetosaurus</i> Extinct genus of reptiles

Aetosaurus is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: A. ferratus, the type species from Germany and Italy; A. crassicauda from Germany; and A. arcuatus from eastern North America. Additional specimens referred to Aetosaurus have been found in the Chinle Group of the southwestern United States, and the Fleming Fjord Formation of Greenland. Specimens of Aetosaurus occur in Norian-age strata.

<i>Desmatosuchus</i> Extinct genus of reptiles

Desmatosuchus is an extinct genus of archosaur belonging to the Order Aetosauria. It lived during the Late Triassic.

Calyptosuchus is an extinct genus of aetosaur from the Late Triassic of North America. Like other aetosaurs, it was heavily armored and had a pig-like snout used to uproot plants.

<i>Aetosauroides</i> Extinct genus of reptiles

Aetosauroides is an extinct genus of aetosaur from the Late Triassic of South America. It is one of four aetosaurs known from South America, the others being Neoaetosauroides, Chilenosuchus and Aetobarbakinoides. Three species have been named: the type species A. scagliai, A. subsulcatus and A. inhamandensis. Fossils have been found in the Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina and the Santa Maria Formation in the Paraná Basin in southeastern Brazil. The strata date to the late Carnian and early Norian stages, making Aetosauroides one of the oldest aetosaurs.

<span class="mw-page-title-main">Santa Maria Formation</span> Geologic formation in Brazil

The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age, and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, including the herrerasaurid Staurikosaurus, the basal sauropodomorphs Buriolestes and Saturnalia, and the lagerpetid Ixalerpeton. The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied.

<i>Typothorax</i> Extinct genus of reptiles

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Lucasuchus is an extinct genus of aetosaur. Fossils have been found from the Bull Canyon Formation of the Dockum Group outcropping in the Revuelto Creek locality in Quay County, New Mexico. All specimens date back to the Norian stage of the Late Triassic. The genus was named in 1995 after the American paleontologist Spencer G. Lucas.

<i>Paratypothorax</i> Genus of reptiles

Paratypothorax is an extinct genus of aetosaur, known from a single species, Paratypothorax andressorum. It was a broadly distributed member of the group found in Germany, North America, and possibly parts of Gondwana. The best specimens come from Germany, though for more than a century they were mistakenly considered phytosaur armor. Paratypothorax was a large and wide-bodied typothoracine aetosaur, as well as the namesake of the tribe Paratypothoracisini.

<span class="mw-page-title-main">Aetosaurinae</span> Extinct subfamily of reptiles

Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Aetosaurus than Desmatosuchus. Aetosaurinae currently comprises Aetosaurus, similar forms such as Coahomasuchus and Stenomyti, and the widespread and successful aetosaur clade Typothoracinae.

<span class="mw-page-title-main">Paratypothoracini</span> Extinct clade of reptiles

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<span class="mw-page-title-main">Desmatosuchinae</span> Subfamily of reptiles

Desmatosuchinae is a major subfamily of aetosaurs within the clade Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Desmatosuchus than to Stagonolepis,Aetosaurus, or Paratypothorax.

Aetobarbakinoides is an extinct genus of stagonolepidid aetosaur known from the Late Triassic of Rio Grande do Sul state, southern Brazil. Fossils have been found from the Santa Maria Supersequence of the late Carnian and early Norian stages, making Aetobarbakinoides one of the oldest aetosaurs. The type species, A. brasiliensis, was named in 2012, and is notable for being described primarily by features of the vertebrae; most other aetosaurs are diagnosed by features in bony plates called osteoderms, which are by far the most common material. Although placed in a basal phylogenetic position among aetosaurs, Aetobarbakinoides is closely related to both desmatosuchines and typothoracisines, two derived clades of aetosaurs.

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References

  1. 1 2 Parker, W. G.; Stocker, M. R.; Irmis, R. B. (2008). "A new desmatosuchine aetosaur (Archosauria; Suchia) from the Upper Triassic Tecovas Formation (Dockum Group) of Texas". Journal of Vertebrate Paleontology. 28 (2): 692–701. doi:10.1671/0272-4634(2008)28[692:ANDAAS]2.0.CO;2. ISSN   0272-4634. S2CID   84455880.
  2. 1 2 Desojo, J.B.; Ezcurra, M.D. (2011). "A reappraisal of the taxonomic status of Aetosauroides (Archosauria, Aetosauria) specimens from the Late Triassic of South America and their proposed synonymy with Stagonolepis". Journal of Vertebrate Paleontology. 31 (3): 596–609. Bibcode:2011JVPal..31..596D. doi:10.1080/02724634.2011.572936. S2CID   85418778.
  3. Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 96. ISBN   978-1-84028-152-1.
  4. Dróżdż, Dawid (2018-10-02). "Osteology of a forelimb of an aetosaur Stagonolepis olenkae (Archosauria: Pseudosuchia: Aetosauria) from the Krasiejów locality in Poland and its probable adaptations for a scratch-digging behavior". PeerJ. 6: e5595. doi: 10.7717/peerj.5595 . ISSN   2167-8359. PMC   6173166 . PMID   30310738.
  5. Sulej, Tomasz (2010). "The skull of an early Late Triassic aetosaur and the evolution of the stagonolepidid archosaurian reptiles". Zoological Journal of the Linnean Society. 158 (4): 860–881. doi: 10.1111/j.1096-3642.2009.00566.x .
  6. Antczak, Mateusz (2015-05-22). "Late Triassic aetosaur (Archosauria) from Krasiejów (SW Poland): new species or an example of individual variation?". Geological Journal. 51 (5): 779–788. doi:10.1002/gj.2691. ISSN   0072-1050. S2CID   128685237.
  7. Heckert, A.B.; Lucas, S.G. (2002). "South American occurrences of the Adamanian (Late Triassic: latest Carnian) index taxon Stagonolepis (Archosauria: Aetosauria) and their biochronological significance". Journal of Paleontology. 76 (5): 852–863. doi:10.1666/0022-3360(2002)076<0852:SAOOTA>2.0.CO;2. S2CID   128610620.
  8. Parker, William G. (2007). "Reassessment of the Aetosaur "Desmatosuchus" chamaensis with a reanalysis of the phylogeny of the Aetosauria (Archosauria:Pseudosuchia)" (PDF). Journal of Systematic Palaeontology. 5: 41–68. doi:10.1017/S1477201906001994. S2CID   85826683.
  9. Julia B. Desojo, Martin D. Ezcurra and Edio E. Kischlat (2012). "A new aetosaur genus (Archosauria: Pseudosuchia) from the early Late Triassic of southern Brazil" (PDF). Zootaxa. 3166: 1–33. doi:10.11646/zootaxa.3166.1.1. ISSN   1175-5326.