Aetobarbakinoides | |
---|---|
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | † Aetosauria |
Family: | † Stagonolepididae |
Genus: | † Aetobarbakinoides Desojo et al., 2012 |
Type species | |
†Aetobarbakinoides brasiliensis Desojo et al., 2012 |
Aetobarbakinoides is an extinct genus of stagonolepidid aetosaur known from the Late Triassic of Rio Grande do Sul state, southern Brazil. Fossils have been found from the Santa Maria Supersequence of the late Carnian and early Norian stages, making Aetobarbakinoides one of the oldest aetosaurs. The type species, A. brasiliensis, was named in 2012, and is notable for being described primarily by features of the vertebrae; most other aetosaurs are diagnosed by features in bony plates called osteoderms, which are by far the most common material. Although placed in a basal phylogenetic position among aetosaurs, Aetobarbakinoides is closely related to both desmatosuchines and typothoracisines, two derived clades of aetosaurs. [1]
Aetobarbakinoides is known only from the holotype specimen CPE2 168, a partially articulated partial postcranial skeleton which is housed at the Coleção Municipal in São Pedro do Sul. It was found near the city of São Pedro do Sul, in the center of Rio Grande do Sul, Brazil. It was collected in the Inhamandá locality, from the Hyperodapedon Assemblage Zone, Sequence 2 of the Santa Maria Supersequence (formerly known as Santa Maria Formation) which dates back to the late Carnian and earliest Norian stages of the Late Triassic. CPE2 168 was assigned by Lucas & Heckert (2001) to Stagonolepis robertsoni . It is the first aetosaur to be diagnosed in its original description without osteoderms, the most common material associated with these reptiles. Along with the genus Aetosauroides from Argentina and Brazil and Stagonolepis robertsoni from Scotland, Aetobarbakinoides is among the oldest aetosaurs. Its presence in South America indicates that aetosaurs had a wide geographic distribution early in their history. Aetobarbakinoides is most closely related to Aetosauroides and Neoaetosauroides . Features of its osteoderms suggest a close relationship with aetosaurines, but its vertebrae are more similar to desmatosuchines. [1]
Aetobarbakinoides is known from neck and back vertebrae, ribs, and several limb bones including the scapula, humerus, tibia, and foot bones. Paramedian osteoderms overlying the midline of the back and neck are also known. Most of the distinguishing features of Aetobarbakinoides are found in its vertebrae. Laterally-oriented projections on the neck vertebrae called prezygapophyses are longer and wider than those of many other aetosaurs. In contrast, the prezygapophyses of the vertebrae at the base of the tail are very thin. The humerus and tibia are large relative to the axial skeleton. The vertebrae of Aetobarbakinoides are more similar to those of Desmatosuchus than they are to its closest relatives, Aetosauroides and Neoaetosaurioides. Aetobarbakinoides is a medium-sized aetosaur, it has been estimated to have been about 2 m (6.6 ft) in total length. [1]
A phylogenetic analysis was conducted in the first description of Aetobarbakinoides. Close relatives included Aetosauroides and Neoaetosauroides, both of which are from South America. Aetobarbakinoides was found to be most closely related to a clade including Desmatosuchinae and Typothoracisinae. In most studies, Typothoracisinae is placed in the subfamily Aetosaurinae, while Desmatosuchinae makes up its own subfamily of aetosaurs. This study suggests that Aetosaurinae is paraphyletic, and that aetosaurines represent a grade of early aetosaurs that developed into desmatosuchines and typothoracisines later in the Triassic. The following cladogram simplified after an analysis presented by Julia B. Desojo, Martin D. Ezcurra and Edio E. Kischlat (2012). [1]
Aetosauria |
| ||||||||||||||||||||||||||||||||||||||||||
Aetobarbakinoides was first named by Julia B. Desojo, Martin D. Ezcurra and Edio E. Kischlat in 2012 and the type species is Aetobarbakinoides brasiliensis. The generic name is derived from Greek aetobarbakina meaning "long-legged buzzard" (the common name of Buteo rufinus ) and from Latin oides meaning "form". It refers to its elongated humerus and tibia and the close relation to the "eagle reptile", Aetosaurus . The specific name refers to Brazil, the country in which the holotype was found. [1]
Staurikosaurus is a genus of herrerasaurid dinosaur from the Late Triassic of Brazil, found in the Santa Maria Formation.
Aetosaurs were heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site that they are found in.
Gracilisuchus is an extinct genus of tiny pseudosuchian from the Late Triassic of Argentina. It contains a single species, G. stipanicicorum, which is placed in the clade Suchia, close to the ancestry of crocodylomorphs. Both the genus and the species were first described by Alfred Romer in 1972.
Aetosaurus is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: A. ferratus, the type species from Germany and Italy; A. crassicauda from Germany; and A. arcuatus from eastern North America. Additional specimens referred to Aetosaurus have been found in the Chinle Group of the southwestern United States, and the Fleming Fjord Formation of Greenland. Specimens of Aetosaurus occur in Norian-age strata.
Stagonolepis is an extinct genus of stagonolepidid aetosaur known from the Late Triassic Hassberge Formation of Germany, the Drawno Beds of Poland, the Lossiemouth Sandstone of Scotland, the Chinle Formation of Arizona and Utah and the Bluewater Creek Formation of New Mexico.
Calyptosuchus is an extinct genus of aetosaur from the Late Triassic of North America. Like other aetosaurs, it was heavily armored and had a pig-like snout used to uproot plants.
Aetosauroides is an extinct genus of aetosaur from the Late Triassic of South America. It is one of four aetosaurs known from South America, the others being Neoaetosauroides, Chilenosuchus and Aetobarbakinoides. Three species have been named: the type species A. scagliai, A. subsulcatus and A. inhamandensis. Fossils have been found in the Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina and the Santa Maria Formation in the Paraná Basin in southeastern Brazil. The strata date to the late Carnian and early Norian stages, making Aetosauroides one of the oldest aetosaurs.
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It has a Late Ladinian to Late Carnian age, and is notable for its fossils of early dinosaurs and other dinosauromorphs, including the herrerasaurid Staurikosaurus, the basal sauropodomorphs Buriolestes and Saturnalia, and the lagerpetid Ixalerpeton. It received this name because it was discovered first in the city of Santa Maria, on the central region of Rio Grande do Sul state.
Typothorax is an extinct genus of typothoracisine aetosaur that lived in the Late Triassic. Its remains have been found in North America. Two species are known: T. coccinarum, the type species, and T. antiquum.
Lucasuchus is an extinct genus of aetosaur. Fossils have been found from the Bull Canyon Formation of the Dockum Group outcropping in the Revuelto Creek locality in Quay County, New Mexico. All specimens date back to the Norian stage of the Late Triassic. The genus was named in 1995 after the American paleontologist Spencer G. Lucas.
Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Aetosaurus than to the last common ancestor of Desmatosuchus. The only synapomorphy that diagnoses the clade Aetosaurinae is the medial offset of the dorsal eminences of the paramedian osteoderms. A phylogenetic study in 2012 found Aetosaurinae to be paraphyletic, with Aetosaurus being the basal-most stagonolepidid and aetosaurines like Calyptosuchus, Neoaetosauroides, and the newly described Aetobarbakinoides being successively more derived taxa leading up to a clade containing Desmatosuchinae and Typothoracisinae. Under this phylogeny, most traditional aetosaurines are more closely related to Desmatosuchus than they are to Aetosaurus, and the clade Aetosaurinae can only include Aetosaurus. However, new phylogenies have recovered Typothoracisinae within Aetosaurinae, making Aetosaurinae a valid clade once more.
Desmatosuchinae is one of the two subfamilies of aetosaurs, the other being Aetosaurinae. It is a stem-based taxon defined as all aetosaurs more closely related to Desmatosuchus than the last common ancestor of Desmatosuchus and Stagonolepis. All synapomorphies that diagnose the clade can be found in the osteoderms. These include tongue-and-groove articulations for lateral plates present in dorsal presacral paramedian plates and large spikes on the lateral cervical, dorsal, and caudal plates.
Tarjadia is an extinct genus of erpetosuchid pseudosuchian, distantly related to modern crocodilians. It is known from a single species, T. ruthae, first described in 1998 from the Middle Triassic Chañares Formation in Argentina. Partial remains have been found from deposits that are Anisian-Ladinian in age. Long known mostly from osteoderms, vertebrae, and fragments of the skull, specimens described in 2017 provided much more anatomical details and showed that it was a fairly large predator. Tarjadia predates known species of aetosaurs and phytosaurs, two Late Triassic groups of crurotarsans with heavy plating, making it one of the first heavily armored archosaurs. Prior to 2017, most studies placed it outside Archosauria as a member of Doswelliidae, a family of heavily armored and crocodile-like archosauriforms. The 2017 specimens instead show that it belonged to the Erpetosuchidae.
Archeopelta is an extinct genus of carnivorous archosauriform from the late Middle or early Late Triassic period. It was a 2 m (6 ft) long predator which lived in what is now southern Brazil. Its exact phylogenetic placement within Archosauriformes is uncertain; it was originally classified as a doswelliid, but subsequently it was argued to be an erpetosuchid archosaur.
Doswelliidae is an extinct family of carnivorous archosauriform reptiles that lived in North America and Europe during the Middle to Late Triassic period. Long represented solely by the heavily-armored reptile Doswellia, the family's composition has expanded since 2011, although two supposed South American doswelliids were later redescribed as erpetosuchids. Doswelliids were not true archosaurs, but they were close relatives and some studies have considered them among the most derived non-archosaurian archosauriforms. They may have also been related to the Proterochampsidae, a South American family of crocodile-like archosauriforms.
Stenomyti is an extinct genus of small aetosaur. It contains a single species, Stenomyti huangae, which is known from a skull, postcranial skeleton and other referred material from the Late Triassic Chinle Formation, Eagle Basin of Colorado, United States. Stenomyti is distinguished from other aetosaurs by eight autapomorphies, however its osteoderms are nearly identical to those of Aetosaurus which suggests that osteoderms are not always reliable taxonomic indicators for aetosaurs. Based on its osteoderms, and other shared cranial characters with Aetosaurus, it was suggested that these taxa are closely related and lie outside the clade containing Typothoracisinae and Desmatosuchinae.
Polesinesuchus is an extinct genus of stagonolepidid aetosaur known from the Late Triassic of southern Brazil. Fossils have been found from the Santa Maria Supersequence of the late Carnian and early Norian stages, making Polesinesuchus one of the oldest aetosaurs. It contains a single species, Polesinesuchus aurelioi, the fifth aetosaur species known from South America to date. Anatomical evidence suggests that Polesinesuchus likely represents a juvenile individual of the contemporary Aetosauroides.
Nundasuchus is an extinct genus of crurotarsan, possibly a suchian archosaur related to Paracrocodylomorpha. Remains of this genus are known from the Middle Triassic Manda beds of southwestern Tanzania. It contains a single species, Nundasuchus songeaensis, known from a single partially complete skeleton, including vertebrae, limb elements, osteoderms, and skull fragments.
Martin Dario Ezcurra is an Argentinian born palaeontologist naming many extinct genera such as Aerotitan, Lophostropheus and Powellvenator.
Dynamosuchus is an extinct genus of pseudosuchian archosaurs from the family Ornithosuchidae. It is known from a single species, Dynamosuchus collisensis, which is based on a partial skeleton from the Santa Maria Formation of Brazil. Dynamosuchus is considered a close relative of Venaticosuchus, which is known from the Ischigualasto Formation of Argentina. Ornithosuchids are one of many groups which lived in the Santa Maria and Ischigualasto Formations, which formed at approximately the same time and were ecologically similar. As a large scavenging reptile, Dynamosuchus helps to illuminate the trophic structure of the Santa Maria Formation. It also supports the hypothesis that ornithosuchids had diversified throughout South America by the start of the Carnian, and were not originally endemic to the Ischigualasto-Villa Unión Basin.