Tecovasuchus | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauria |
Clade: | Pseudosuchia |
Order: | † Aetosauria |
Family: | † Stagonolepididae |
Genus: | † Tecovasuchus Martz & Small, 2006 |
Species: | †T. chatterjeei |
Binomial name | |
†Tecovasuchus chatterjeei Martz & Small, 2006 | |
Tecovasuchus is an extinct genus of aetosaur. It is known primarily from osteoderms found from the Tecovas Formation in Texas, which is Late Triassic in age, dating back to the lower Norian. Material is also known from several other localities of the Chinle Group in New Mexico and Arizona, such as older Carnian outcrops and younger Rhaetian outcrops. Specimens of Tecovasuchus have been collected from the Tecovas Formation, the Bluewater Creek Formation, and the Los Esteros Member of the Santa Rosa Formation. [1] [2]
Tecovasuchus was first recognized as a new taxon in 1995, although it was not named until the description of the type species, T. chatterjeei, in 2006. [3] [4] Before its description, specimens of Tecovasuchus were thought to belong to Paratypothorax or a Paratypothorax-like aetosaur. [2] [5] Several features of the osteoderms distinguish Tecovasuchus from other aetosaur genera, including dorsal paramedian osteoderms with strongly thickened and beveled posterior edges and ornamentation consisting of deep pits and radiating grooves, as well as tongue-shaped dorsal and plate-like ventrolateral flanges.
Tecovasuchus is an index taxon for the St. Johnsian sub-LVF (land vertebrate faunachron) of the Adamanian LVF, and the presence of material belonging to the genus helps correlate different Late Triassic localities throughout the southwestern United States. [1]
Aetosaurs are heavily armored reptiles belonging to the extinct order Aetosauria. They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs more closely related to crocodilians than to birds and other dinosaurs. All known aetosaurs are restricted to the Late Triassic, and in some strata from this time they are among the most abundant fossil vertebrates. They have small heads, upturned snouts, erect limbs, and a body ornamented with four rows of plate-like osteoderms. Aetosaur fossil remains are known from Europe, North and South America, parts of Africa, and India. Since their armoured plates are often preserved and are abundant in certain localities, aetosaurs serve as important Late Triassic tetrapod index fossils. Many aetosaurs had wide geographic ranges, but their stratigraphic ranges were relatively short. Therefore, the presence of particular aetosaurs can accurately date a site in which they are found.
Acaenasuchus is an extinct genus of pseudosuchian, endemic to what would be presently be known as Arizona during the Late Triassic, specifically during the Carnian and Norian stages of the Triassic. Acaenasuchus had a stratigraphic range of approximately 11.5 million years. Acaenasuchus is further categorized as one of the type fauna that belong to the Adamanian LVF, based on the fauna of the Blue Mesa Member of the Chinle Petrified Forest Formation of Arizona, where Acaenasuchus was initially discovered.
Typothorax is an extinct genus of typothoracine aetosaur that lived in the Late Triassic. Its remains have been found in North America. Two species are known: T. coccinarum, the type species, and T. antiquum.
The Tecovas Formation is a geological formation in the Texas panhandle and eastern New Mexico. It is one of several formations encompassed by the Late Triassic Dockum Group.
Ebrachosaurus is an extinct genus of aetosaur. It was named after the town of Ebrach, Germany, near an outcrop of the Blasensandstein Formation where the original fossils have been found. Other Blasensandstein fauna include the temnospondyl Metoposaurus and the phytosaur Francosuchus. The genus has often been considered synonymous with the closely related Stagonolepis. The holotype specimen was lost during World War II, so its relationships within Stagonolepididae remain indeterminant.
Adamanasuchus is an extinct genus of aetosaur. Fossils have been found from several localities from the Chinle Group in Arizona and date back to the late Carnian stage of the Late Triassic. The locality from which it was named after also lends its name to the Adamanian LVF.
Rioarribasuchus is a genus of aetosaur. Fossils have been found from the Chinle Formation in Arizona and New Mexico that date back to the upper Late Carnian stage of the Late Triassic.
Lucasuchus is an extinct genus of aetosaur. Fossils have been found from the Bull Canyon Formation of the Dockum Group outcropping in the Revuelto Creek locality in Quay County, New Mexico. All specimens date back to the Norian stage of the Late Triassic. The genus was named in 1995 after the American paleontologist Spencer G. Lucas.
Paratypothorax is an extinct genus of aetosaur, known from a single species, Paratypothorax andressorum. It was a broadly distributed member of the group found in Germany, North America, and possibly parts of Gondwana. The best specimens come from Germany, though for more than a century they were mistakenly considered phytosaur armor. Paratypothorax was a large and wide-bodied typothoracine aetosaur, as well as the namesake of the tribe Paratypothoracisini.
Aetosaurinae is one of the two main clades of aetosaurs, the other being Desmatosuchia. It is a stem-based taxon defined as all aetosaurs more closely related to Aetosaurus than Desmatosuchus. Aetosaurinae currently comprises Aetosaurus, similar forms such as Coahomasuchus and Stenomyti, and the widespread and successful aetosaur clade Typothoracinae.
Colognathus is a genus of extinct reptile from Late Triassic rocks of the southwestern United States. It was described in 1928 from a jaw fragment by Case, who interpreted the new taxon as a fish. The type species is C. obscurus.
Redondasuchus is an extinct genus of aetosaur. It may be a junior synonym of Typothorax coccinarum, another aetosaur. Redondasuchus is a member of the clade Typothoracisinae within the subfamily Aetosaurinae, and lived during the middle Norian stage of the Late Triassic. Material belonging to the genus has been found from the Redonda Formation in east-central New Mexico. The type species, R. reseri, was named in 1991 after having been referred to as a species of Typothorax since 1985. A second species, R. rineharti, was described in 2006.
Doswelliidae is an extinct family of carnivorous archosauriform reptiles that lived in North America and Europe during the Middle to Late Triassic period. Long represented solely by the heavily-armored reptile Doswellia, the family's composition has expanded since 2011, although two supposed South American doswelliids were later redescribed as erpetosuchids. Doswelliids were not true archosaurs, but they were close relatives and some studies have considered them among the most derived non-archosaurian archosauriforms. They may have also been related to the Proterochampsidae, a South American family of crocodile-like archosauriforms.
The Colorado City Formation is a Late Triassic geologic formation in the Dockum Group of Texas, United States. It has previously been known as the Iatan Member, Colorado City Member or 'Pre-Tecovas Horizon'.
The Bluewater Creek Formation is a geologic formation in west-central New Mexico. It preserves fossils dating back to the late Triassic period.
Scutarx is an extinct genus of Aetosauriformes, most commonly regarded by its species name Scutarx deltatylus. Scutarx lived around 230 million years ago during the Carnian and Norian stage of the Late Triassic. Scutarx are “medium sized” paramedian osteoderms belonging to the clade Aetosauria, a heavily armored and more herbivorous cousin of crocodiles.
Apachesuchus is an extinct genus of aetosaur from the Late Triassic of New Mexico. It is only known from several paramedian osteoderms discovered in Quay County in eastern New Mexico. This area belongs to the late Norian-age Quay Member of the Redonda Formation. Unique among aetosaurs, its osteoderms are nearly completely smooth, without strong pits or grooves. The left dorsal paramedian has a relatively high width-to-length ration, suggesting that Apachesuchus is a wide-bodied aetosaur within the clade Typothoracinae.
Land vertebrate faunachrons (LVFs) are biochronological units used to correlate and date terrestrial sediments and fossils based on their tetrapod faunas. First formulated on a global scale by Spencer G. Lucas in 1998, LVFs are primarily used within the Triassic Period, though Lucas later designated LVFs for other periods as well. Eight worldwide LVFs are defined for the Triassic. The first two earliest Triassic LVFs, the Lootsbergian and Nonesian, are based on South African synapsids and faunal assemblage zones estimated to correspond to the Early Triassic. These are followed by the Perovkan and Berdyankian, based on temnospondyl amphibians and Russian assemblages estimated to be from the Middle Triassic. The youngest four Triassic LVFs, the Otischalkian, Adamanian, Revueltian, and Apachean, are based on aetosaur and phytosaur reptiles common in the Late Triassic of the southwestern United States.
Kryphioparma is an extinct genus of aetosaur from the Blue Mesa Member of the Chinle Formation, Arizona. It is the oldest known member of the subfamily Typothoracinae, but currently only known from four isolated and incomplete dorsal osteoderms. Regardless, said osteoderms show a clear mix of features that does not match any other known aetosaur and were thus used as the basis for a new genus and species in 2023. The genus is monotypic, only including a single species, Kryphioparma caerula.