Carnian

Last updated
Carnian
~237 – ~227 Ma
Travenanzes.jpg
Alluvial plain red clays of the Travenanzes Formation, upper Carnian, the Dolomites, northern Italy
Chronology
Etymology
Name formalityFormal
Alternate spelling(s)Karnian
Usage information
Celestial body Earth
Regional usageGlobal (ICS)
Time scale(s) usedICS Time Scale
Definition
Chronological unit Age
Stratigraphic unit Stage
Time span formalityFormal
Lower boundary definition FAD of the Ammonite Daxatina canadensis
Lower boundary GSSP Prati di Stuores, Dolomites, Italy
46°31′37″N11°55′49″E / 46.5269°N 11.9303°E / 46.5269; 11.9303
Lower GSSP ratified2008 [6]
Upper boundary definitionNot formally defined [7]
Upper boundary definition candidatesBase of Stikinoceras kerri ammonoid zone and near FAD of Metapolygnathus echinatus within the M. communisti conodont zones [7]
Upper boundary GSSP candidate section(s)

The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). [8] The Carnian is preceded by the Ladinian and is followed by the Norian. Its boundaries are not characterized by major extinctions or biotic turnovers, but a climatic event (known as the Carnian pluvial episode characterized by substantial rainfall) occurred during the Carnian and seems to be associated with important extinctions or biotic radiations. [9] Another extinction occurred at the Carnian-Norian boundary, ending the Carnian age. [10]

Contents

Stratigraphic definitions

Brotheotrachyceras brotheus from the San Cassiano Formation, Val Badia, Dolomites, Southern Alps. This ammonoid is an index fossil for the lower Carnian Trachyceras Brotheotrachyceras brotheus.jpg
Brotheotrachyceras brotheus from the San Cassiano Formation, Val Badia, Dolomites, Southern Alps. This ammonoid is an index fossil for the lower Carnian

The Carnian was named in 1869 by Mojsisovics. It is unclear if it was named after the Carnic Alps or after the Austrian region of Carinthia (Kärnten in German) [11] or after the Carnia historical region in northeastern Italy.[ citation needed ] The name, however, was first used referring to a part of the Hallstatt Limestone cropping out in Austria. [11]

The base of the Carnian Stage is defined as the place in the stratigraphic record where the ammonite species Daxatina canadensis first appears. The global reference profile for the base is located at the Stuores-Wiesen near Badia in the Val Badia in the region of South Tyrol, Italy. [12]

The top of the Carnian (the base of the Norian) is at the bases of the ammonite biozones of Klamathites macrolobatus or Stikinoceras kerri and the conodont biozones of Metapolygnathus communisti or Metapolygnathus primitius .

Subdivisions

There is no established, standard usage for the Carnian subdivisions, thus, while in some regional stratigraphies a two-substage subdivision is common:

others prefer a three-substage organization of the stage as follows:

Biostratigraphy

In the Tethys domain, the Carnian Stage contains six ammonite biozones:

The Otischalkian land vertebrate faunachron corresponds to the early late Carnian, while the Adamanian land vertebrate faunachron corresponds to the latest Carnian. [13]

Paleogeography and climate

Carnian Earth CARNIAN EARTH.jpg
Carnian Earth

The paleogeography of the Carnian was basically the same as for the rest of the Triassic. Most continents were merged into the supercontinent Pangaea, and there was a single global ocean, Panthalassa. The global ocean had a western branch at tropical latitudes called Paleo-Tethys. The sediments of Paleo-Tethys now crop out in southeastern Europe, in the Middle East, in the Himalayas, and up to the island of Timor. The extreme land-sea distribution led to "mega-monsoons", i.e., an atmospheric monsoon regime more intense than the present one.

As for most of the Mesozoic, there were no ice caps. Climate was mostly arid in the tropics, but an episode of wet tropical climate is documented at least in the Paleo-Tethys. This putative climatic event is called the "Carnian Pluvial Event", its age being between latest early Carnian (Julian) and the beginning of late Carnian (Tuvalian).

Carnian life

In the marine realm, the Carnian saw the first abundant occurrences of calcareous nanoplankton, a morphological group including the coccolithophores.

Invertebrates

There are a few invertebrates which are typical and characteristic of the Carnian. Among molluscs, the ammonoid genus Trachyceras is exclusive to the lower Carnian (i.e., Julian of the two-substages subdivision, see above). The family Tropitidae and the genus Tropites appear at the base of the upper Carnian (Tuvalian). The bivalve genus Halobia, a bottom-dweller of deep sea environments, differentiated from Daonella at the beginning of this age. Scleractinian coral reefs, i.e., reefs with corals of the modern type, became relatively common for the first time in the Carnian.

Vertebrates

The earliest unequivocal dinosaurs, such as those from the Ischigualato Formation (e.g. Herrerasaurus and Eoraptor ) and those from the Santa Maria Formation (e.g. Staurikosaurus and Buriolestes ) originated during the Carnian, around 230 Ma.

In this stage the archosaurs became the dominant faunas in the world, evolving into groups such as the phytosaurs, rhynchosaurs, aetosaurs, and rauisuchians. The first dinosaurs (and the pterosaur Carniadactylus) also appeared in this stage, and though at the time they were small and insignificant, they diversified rapidly and would dominate the fauna for the rest of the Mesozoic. On the other hand, the therapsids, which included the ancestors of mammals, decreased in both size and diversity, and would remain relatively small until the extinction of the dinosaurs.

Conodonts were present in Triassic marine sediments. Paragondolella polygnathiformis appeared at the base of the Carnian Stage, and is considered a characteristic species. A partial list of Carnian vertebrates is given below. Many Carnian vertebrates are found in Santa Maria Formation rocks of the Paleorrota geopark.

Classic localities and Lagerstätten

The lower Carnian fauna of the San Cassiano Formation (Dolomites, northern Italy) has been studied since the 19th century. Fossiliferous localities are many, and are distributed mostly in the surroundings of Cortina d'Ampezzo and in the high Badia Valley, near the village of San Cassiano, after which the formation was named. This fauna is extremely diverse, including ammonoids, gastropods, bivalves, echinoderms, calcareous sponge, corals, brachiopods, and a variety of less common fossils. A collection of this fauna is exposed in the "Museo delle Regole", a museum in Cortina d'Ampezzo.

The Ischigualasto Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina yielded a very important vertebrate association, including the oldest dinosaurian assemblage.

The Lagerstätte of the Madygen Formation in Kyrgyzstan has provided over 20,000 fossil insects, vertebrates and flora.

Notable Formations

* Tentatively assigned to the Carnian; age estimated primarily via terrestrial tetrapod biostratigraphy (see Triassic land vertebrate faunachrons)

Related Research Articles

The Triassic is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.

The Lopingian is the uppermost series/last epoch of the Permian. It is the last epoch of the Paleozoic. The Lopingian was preceded by the Guadalupian and followed by the Early Triassic.

The Rhaetian is the latest age of the Triassic Period or the uppermost stage of the Triassic System. It was preceded by the Norian and succeeded by the Hettangian. The base of the Rhaetian lacks a formal GSSP, though candidate sections include Steinbergkogel in Austria and Pignola-Abriola in Italy. The end of the Rhaetian is more well-defined. According to the current ICS system, the Rhaetian ended 201.4 ± 0.2 Ma.

<span class="mw-page-title-main">Anisian</span> Stage of the Triassic

In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from 247.2 million years ago until 242 million years ago. The Anisian Age succeeds the Olenekian Age and precedes the Ladinian Age.

The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 to 72.1 million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian.

The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya and 83.6 ± 0.7 mya. The Santonian is preceded by the Coniacian and is followed by the Campanian.

The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma. The Hettangian follows the Rhaetian and is followed by the Sinemurian.

In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma. The Kimmeridgian follows the Oxfordian and precedes the Tithonian.

<span class="mw-page-title-main">Wuchiapingian</span> Eighth stage of the Permian

In the geologic timescale, the Wuchiapingian or Wujiapingian is an age or stage of the Permian. It is also the lower or earlier of two subdivisions of the Lopingian Epoch or Series. The Wuchiapingian spans the time between 259.51 and 254.14 million years ago (Ma). It was preceded by the Capitanian and followed by the Changhsingian.

In the geologic time scale, the Changhsingian or Changxingian is the latest age or uppermost stage of the Permian. It is also the upper or latest of two subdivisions of the Lopingian Epoch or Series. The Changhsingian lasted from 254.14 to 251.9 million years ago (Ma). It was preceded by the Wuchiapingian and followed by the Induan.

<span class="mw-page-title-main">Middle Triassic</span> Second epoch of the Triassic period

In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between 247.2 Ma and 237 Ma. It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages.

The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between 237 Ma and 201.4 Ma. It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian ages.

<span class="mw-page-title-main">Induan</span> First age of the Early Triassic epoch

The Induan is the first age of the Early Triassic epoch in the geologic timescale, or the lowest stage of the Lower Triassic series in chronostratigraphy. It spans the time between 251.9 Ma and 251.2 Ma. The Induan is sometimes divided into the Griesbachian and the Dienerian subages or substages. The Induan is preceded by the Changhsingian and is followed by the Olenekian.

<span class="mw-page-title-main">Ladinian</span> Age in the Middle Triassic

The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between 242 Ma and ~237 Ma. The Ladinian was preceded by the Anisian and succeeded by the Carnian.

<span class="mw-page-title-main">Trachyceratidae</span> Family of molluscs (fossil)

The Trachyceratidae is an extinct family of ceratitid ammonoid cephalopods.

The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to 208.5 million years ago. It was preceded by the Carnian and succeeded by the Rhaetian.

The Carnian pluvial episode (CPE), often called the Carnian pluvial event, was an interval of major change in global climate that was synchronous with significant changes in Earth's biota both in the sea and on land. It occurred during the latter part of the Carnian Stage, a subdivision of the late Triassic period, and lasted for perhaps 1-2 million years. The CPE represents a significant episode in the evolution and diversification of many taxa that are important today, among them some of the earliest dinosaurs, lepidosaurs, pterosaurs and true mammals. In the marine realm it saw the first appearance among the microplankton of coccoliths and dinoflagellates, with the latter linked to the rapid diversification of scleractinian corals through the establishment of symbiotic zooxanthellae within them. The CPE also saw the extinction of many aquatic invertebrate species, especially among the ammonoids, bryozoa, and crinoids.

The Pignola-Abriola section is a ~63 m long stratigraphic sequence of cherty limestones deposited in the Lagonegro Basin during the latest Norian and the early Rhaetian Stages. The main outcrop is on the western side of Mount Crocetta along the SP5 road connecting the villages of Pignola and Abriola. A smaller outcrop, overlapping the central part of the main section, is located near a former railway tunnel, few meters below the road level. The Pignola-Abriola section has been recently proposed as GSSP of the Rhaetian Stage.

Land vertebrate faunachrons (LVFs) are biochronological units used to correlate and date terrestrial sediments and fossils based on their tetrapod faunas. First formulated on a global scale by Spencer G. Lucas in 1998, LVFs are primarily used within the Triassic Period, though Lucas later designated LVFs for other periods as well. Eight worldwide LVFs are defined for the Triassic. The first two earliest Triassic LVFs, the Lootsbergian and Nonesian, are based on South African synapsids and faunal assemblage zones estimated to correspond to the Early Triassic. These are followed by the Perovkan and Berdyankian, based on temnospondyl amphibians and Russian assemblages estimated to be from the Middle Triassic. The youngest four Triassic LVFs, the Otischalkian, Adamanian, Revueltian, and Apachean, are based on aetosaur and phytosaur reptiles common in the Late Triassic of the southwestern United States.

References

  1. ↑ Widmann, Philipp; Bucher, Hugo; Leu, Marc; et al. (2020). "Dynamics of the Largest Carbon Isotope Excursion During the Early Triassic Biotic Recovery". Frontiers in Earth Science. 8 (196): 196. Bibcode:2020FrEaS...8..196W. doi: 10.3389/feart.2020.00196 .
  2. ↑ McElwain, J. C.; Punyasena, S. W. (2007). "Mass extinction events and the plant fossil record". Trends in Ecology & Evolution. 22 (10): 548–557. doi:10.1016/j.tree.2007.09.003. PMID   17919771.
  3. ↑ Retallack, G. J.; Veevers, J.; Morante, R. (1996). "Global coal gap between Permian–Triassic extinctions and middle Triassic recovery of peat forming plants". GSA Bulletin. 108 (2): 195–207. Bibcode:1996GSAB..108..195R. doi:10.1130/0016-7606(1996)108<0195:GCGBPT>2.3.CO;2 . Retrieved 2007-09-29.
  4. ↑ Payne, J. L.; Lehrmann, D. J.; Wei, J.; Orchard, M. J.; Schrag, D. P.; Knoll, A. H. (2004). "Large Perturbations of the Carbon Cycle During Recovery from the End-Permian Extinction". Science. 305 (5683): 506–9. Bibcode:2004Sci...305..506P. doi:10.1126/science.1097023. PMID   15273391. S2CID   35498132.
  5. ↑ Ogg, James G.; Ogg, Gabi M.; Gradstein, Felix M. (2016). "Triassic". A Concise Geologic Time Scale: 2016. Elsevier. pp. 133–149. ISBN   978-0-444-63771-0.
  6. ↑ Mietto, Paolo; Manfrin, Stefano; Preto, Nereo; Rigo, Manuel; Roghi, Guido; Furin, Stefano; Gianolla, Piero; Posenato, Renato; Muttoni, Giovanni; Nicora, Alda; Buratti, Nicoletta; Cirilli, Simonetta; Spötl, Christoph; Ramezani, Jahandar; Bowring, Samuel (September 2012). "The Global Boundary Stratotype Section and Point (GSSP) of the Carnian Stage (Late Triassic) at Prati Di Stuores/Stuores Wiesen Section (Southern Alps, NE Italy)" (PDF). Episodes. 35 (3): 414–430. doi:10.18814/epiiugs/2012/v35i3/003 . Retrieved 13 December 2020.
  7. 1 2 3 4 "Global Boundary Stratotype Section and Point". International Commission of Stratigraphy. Retrieved 23 December 2020.
  8. ↑ "Ics-chart".
  9. ↑ Dal Corso, Jacopo; Bernardi, Massimo; Sun, Yadong; Song, Haijun; Seyfullah, Leyla J.; Preto, Nereo; Gianolla, Piero; Ruffell, Alastair; Kustatscher, Evelyn; Roghi, Guido; Merico, Agostino (2020). "Extinction and dawn of the modern world in the Carnian (Late Triassic)". Science Advances. 6 (38): eaba0099. Bibcode:2020SciA....6...99D. doi:10.1126/sciadv.aba0099. PMC   7494334 . PMID   32938682.
  10. ↑ Onoue, Tetsuji; Zonneveld, John-Paul; Orchard, Michael J.; Yamashita, Misa; Yamashita, Katsuyuki; Sato, Honami; Kusaka, Soichiro (1 January 2016). "Paleoenvironmental changes across the Carnian/Norian boundary in the Black Bear Ridge section, British Columbia, Canada". Palaeogeography, Palaeoclimatology, Palaeoecology . 441: 721–733. Bibcode:2016PPP...441..721O. doi:10.1016/j.palaeo.2015.10.008 . Retrieved 24 January 2023.
  11. 1 2 Gradstein FM, Ogg JG, Schmitz MD, Ogg GM, eds. (2012). The Geologic Timescale 2012 (volume 1). Elsevier. p. 687. ISBN   978-0-44-459390-0.
  12. ↑ The GSSP was firstly proposed by Broglio Loriga et al. (1999) and established by Mietto et al. (2012)
  13. ↑ Heckert, A.B. (2004). "Late Triassic microvertebrates from the lower Chinle Group (Otischalkian-Adamanian: Carnian), southwestern USA". New Mexico Museum of Natural History and Science Bulletin. 27: 1. Retrieved 17 November 2021.

Bibliography

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