Santa Maria Formation | |
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Stratigraphic range: Carnian ~ | |
Type | Geological formation |
Unit of | Rosário do Sul Group |
Sub-units | Passo das Tropas & Alemoa Members |
Underlies | Caturrita Formation |
Overlies | Sanga do Cabral Formation |
Location | |
Coordinates | 29°41′42″S53°47′43″W / 29.695042°S 53.795403°W |
Region | Rio Grande do Sul |
Country | Brazil |
Extent | Paraná Basin |
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age [3] [4] [5] (Late Triassic), [6] [7] and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, [8] [9] including the herrerasaurid Staurikosaurus , the basal sauropodomorphs Buriolestes and Saturnalia, and the lagerpetid Ixalerpeton . [10] [11] [12] [13] The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied.
The Santa Maria Formation makes up the majority of the Santa Maria Supersequence, which extends through the entire Late Triassic. The Santa Maria Supersequence is divided into four geological sequences, separated from each other by short unconformities. The first two of these sequences (Pinheiros-Chiniquá and Santa Cruz sequences) lie entirely within the Santa Maria Formation, while the third (the Candelária sequence) is shared with the overlying Norian-age Caturrita Formation. The fourth and youngest sequence (the Mata sequence) is equivalent to the Rhaetian-age Mata Sandstone. [7]
The oldest sequence in the formation is the Pinheiros-Chiniquá Sequence (latest Ladinian-earliest Carnian, ~237 Ma), which is biostratigraphically equivalent to the Dinodontosaurus Assemblage Zone. It is followed by the shorter Santa Cruz Sequence (early Carnian-middle Carnian, ~236 Ma), [3] biostratigraphically equivalent to the Santacruzodon Assemblage Zone. [7]
The final sequence, which is only partially present within the formation, is the Candelária Sequence (middle Carnian-latest Carnian, ~233-228 Ma). [4] [14] [5] [6] [15] The lower portion of this sequence, coinciding with the upper part of the Santa Maria Formation, is equivalent to the Hyperodapedon Assemblage Zone.The Hyperodapedon Assemblage Zone is itself subdivided into Hyperodapedon Acme Zone (most of the zone, where the rhynchosaur Hyperodapedon is widely reported) and Exaeretodon Zone (restricted to about three known and sampled localities, where rhynchsaurs are almost completely absent, but the traversodontid cynodont Exaeretodon is widely reported). [16] [7] These subdivisions are also known as Lower and Upper Hyperodapedon Assemblage Zone, respectively. [15] [16]
U-Pb radiometric dating of Cerro da Alemoa (the type locality of Saturnalia tupiniquim ) in the Upper portion of the Santa Maria Formation found an estimated age of 233.23±0.73 million years ago, putting that locality 1.5 million years older than the Ischigualasto Formation and younger than Los Chañares Formation. The Santa Maria and Ischigualasto formations are approximately equal as having the earliest dinosaur localities. [4] [9] [17] [18] [19]
Most of the information below was included on a revision of the Triassic faunal successions of the Paraná Basin, southern Brazil by Schultz et al. (2020). [7]
Temnospondyls of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Compsocerops [20] | C. sp. [21] | Restinga Sêca | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | A partial skull | A chigutisaurid temnospondyl |
Temnospondyls of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Ptychoceratodus [5] | P. cf. phillipsi | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | A ptychoceratodontid lungfish |
Dicynodonts of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Dinodontosaurus | D.pedroanum | São Pedro do Sul, Candelária, Dona Francisca, Bom Retiro, Vale Verde, Novo Cabrais, Bom Retiro do Sul, Dilermando de Aguiar | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial and postcranial remains | A kannemeyeriiform dicynodont | |
Stahleckeria | S.potens | Dona Francisca, Candelária | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial and postcranial remains | A stahleckeriid dicyndont | |
Cynognathians of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Exaeretodon [22] | E. major | São Pedro do Sul | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial and postcranial remains | A traversodontid cynodont | |
E. riograndensis | Agudo, Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon Zone | Cranial and postcranial remains | A traversodontid cynodont | ||
Gomphodontosuchus [23] | G. brasiliensis | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Cranial remains | A traversodontid cynodont | |
Luangwa | L.sudamericana | Vera Cruz, Candelária | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial remains | A traversodontid cynodont | |
Massetognathus | M. ochagavie | Bom Retiro, Vale Verde, Dona Francisca | Lower and Middle Santa Maria Formation, Alemoa Member, Santa Cruz nad Pinheiros-Chiniquá Sequences, Santacruzodon Assemblage Zone and Dinodontosaurus Assemblage Zone | Cranial and postcranial remains | A traversodontid cynodont | |
Menadon | M.besairiei | Santa Cruz | Middle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage Zone | Cranial remains | A traversodontid cynodont | |
Protuberum [24] | P. cabralensis | Paraíso do Sul, Novo Cabrais | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Partial articulated skeleton including skull, vertebrae and ribs | A traversodontid cynodont | |
Santacruzodon [25] | S. hopsoni [25] | Santa Cruz | Middle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage Zone | Cranial remains [25] | A traversodontid cynodont [25] | |
Santagnathus [26] | S. mariensis | Hyperodapedon Assemblage Zone | Numerous cranial and postcranial remains | A traversodontid cynodont | ||
Scalenodon | S.ribeiroae | Agudo | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial remains | A traversodontid cynodont | |
Traversodon | T.stahleckeri | São Pedro do Sul | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial remains | A traversodontid cynodont | |
Probainognathians of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Alemoatherium [27] | A. huebneri | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | A partial dentary | A prozostrodontian cynodont | |
Aleodon | A.cromptoni | Vale Verde, Candelária, Bom Retiro do Sul, Novo Cabrais | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial and postcranial remains | A chiniquodontid cynodont | |
Bonacynodon | B.schultzi | Candelária | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial remains | A probainognathid cynodont | |
Candelariodon [28] | C. barberenai | Candelária | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | A partial mandible having some complete teeth | A basal probainognathian cynodont | |
Charruodon [29] | C.tetracuspidatus | Candelária | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | A partial dentary | A basal probainognathian cynodont [30] | |
Chiniquodon [23] | C. theotonicus | São Pedro do Sul | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Several cranial remains | A chiniquodontid cynodont | |
C. sp. | Santa Cruz | Middle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage Zone | Cranial and postcranial remains | A chiniquodontid cynodont | ||
Protheriodon | P.estudianti | Dona Francisca | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial remains | A basal probainognathian cynodont | |
Prozostrodon [31] | P. brasiliensis | Santa Maria, São João do Polêsine | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Cranial remains | A prozostrodontian cynodont | |
Santacruzgnathus [32] | S. abdalai | Santa Cruz | Middle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage Zone | Cranial remains | A prozostrodontian cynodont | |
Therioherpeton [33] | T. cargnini | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Cranial remains | A prozostrodontian cynodont | |
Trucidocynodon [34] | T. riograndensis | Agudo | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon Zone | Cranial and postcranial elements of an almost complete skeleton and an additional large skull | An ecteniniid cynodont | |
Pseudosuchians of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Aetobarbakinoides [35] | A. brasiliensis | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Partial postcranial skeleton | A aetosaur | |
Aetosauroides [36] | A. scagliai | Santa Maria, São João do Polêsine, São Pedro do Sul | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | An almost complete skull and incomplete postcranial elements | An aetosaur | |
Archeopelta | A. arborensis | São Pedro do Sul | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial and postcranial elements | An erpetosuchid | |
Dagasuchus [2] | D. santacruzensis | Santa Cruz do Sul | Middle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage Zone | An ilium and a pair of ischia | A loricatan | |
Decuriasuchus [37] | D. quartacolonia | Dona Francisca | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Assemblage of several individuals, which together preserved virtually all the skeleton | A loricatan | |
Dynamosuchus [38] | D. collisensis | Agudo | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon Zone | Semi-articulated partial skull and disarticulated fragmentary postcranial skeleton | An ornithosuchid | |
Pagosvenator [39] | P. candelariensis | Candelária | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Poorly preserved skull and cervical elements | An erpetosuchid | |
Parvosuchus | P. aurelioi | Linha Várzea 2, Pinheiros-Chiniquá. | Dinodontosaurus Assemblage Zone. | A largely articulated skeleton, skull, and lower jaws. | A gracilisuchid | |
"Polesinesuchus" [40] | "P. aurelioi" | São João do Polêsine | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Several postcranial elements | An immature aetosaur, possible synonym of Aetosauroides scagliai [41] | |
Prestosuchus [42] [43] [44] | P. chiniquensis "P. loricatus""Karamuru vorax" | Candelária, Dona Francisca, São Pedro do Sul, and possibly Vale Verde ("K. vorax") | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Virtually all the skeleton is known | A loricatan | |
Procerosuchus [45] | P. celer | São Pedro do Sul | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Postcranial elements | A loricatan | |
Rauisuchus [46] | R. tiradentes | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Incomplete skeleton including skull elements, several vertebrae and partial hindlimb | A loricatan | |
Unnamed loricatan [47] | - | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Specimen UFSM 11617, an almost complete right maxilla | A loricatan | |
Avemetatarsalians of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Amanasaurus [48] | A. nesbitti | Restinga Sêca | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Two partial femora | A silesaurid | |
Bagualosaurus [49] | B. agudoensis | Agudo | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon Zone | Almost complete skull with lower jaws, several vertebrae, partial pelvic girdle, partial hindlimbs | A basal sauropodomorph | |
Buriolestes [50] [51] | B. schultzi | São João do Polêsine | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Parts of the skull, vertebrae, left forelimb, and left hindlimb | The basalmost sauropodomorph | |
Gamatavus [52] | G. antiquus | Dilermando de Aguiar | Lower Santa Maria Formation ('Picada do Gama' site), Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | A partial right ilium; a partial left femur and four incomplete vertebrae found in association with the holotype may belong to Gamatavus, but they were not assigned | One of the oldest known silesaurids from South America | |
Gnathovorax [53] | G. cabreirai | São João do Polêsine | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | A complete semi-articulated skeleton, part of a multiataxic fossil assemblage, including two cynodonts and two rhynchosaurs | A herrerasaurid | |
Gondwanax [54] | G. paraisensis | Paraíso do Sul | Lower Santa Maria Formation ('Linha Várzea 2' site), Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Associated remains possibly belonging to multiple individuals, including cervical, dorsal, sacral, and caudal vertebrae, a partial pelvic girdle, and a femur | One of the oldest known silesaurids from South America | |
Ixalerpeton [50] | I. polesinensis | São João do Polêsine | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Partial skull including a partial maxilla and both dentaries, most of the presacral vertebrae, two sacral vertebrae, several caudal verterae and associated chevrons, scapula and humerus, femora of at least two individuals, and tibia | A basal lagerpetid | |
Nhandumirim [55] | N. waldsangae | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Some vertebrae, partial pelvic girdle, partial hindlimb | A juvenile saturnaliid sauropodomorph [56] [57] [58] [59] [60] | |
Pampadromaeus [61] | P. barberenai | Agudo | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon Zone | Almost complete skull and lower jaws, several vertebrae, partial pectoral girdle, partial forelimb, partial pelvic girdle, partial hindlimb | A basal sauropodomorph | |
Saturnalia [13] | S. tupinquim | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Three partial skeleton, including partial skull and lower jaw, complete cervical, dorsal and partial caudal vertebral series, pectoral girdle, partial forelimb, pelvic girdle, partial hindlimb | A basal sauropodomorph | |
Spondylosoma [62] | S. absconditum | São Pedro do Sul | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Several postcranial remains | An aphanosaurian | |
Staurikosaurus [10] [11] | S. pricei | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Partial postcranial skeleton with mandible | A basal herrerasaurid | |
Venetoraptor [63] | V. gassenae | São João do Polêsine | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Partially articulated skeleton consisting of an incomplete skull, cervical, dorsal, and caudal vertebrae, a partial pelvic girdle, and elements of the arms and legs | A lagerpetid. | |
Unnamed dinosauromorph [64] | - | Dona Francisca | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Specimen CAPPA/UFSM 0282, a complete, but poorly preserved right femur | The oldest dinosauromorph from South America | |
Unnamed herrerasaurid [65] | - | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Specimen UFSM 11330, comprised by partial cranial and postcranial remains | The largest Triassic dinosaur skeleton found in Rio Grande do Sul state so far | |
Unnamed lagerpetid [9] | - | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Specimen UFSM 11611 (partial left femur) | A basal lagerpetid distinct from Ixalerpeton | |
Unnamed silesaurid [9] [66] | - | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Specimen UFSM 11579, comprised by cranial and postcranial remains | A silesaurid | |
Other reptiles of the Santa Maria Formation | ||||||
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Genus | Species | Location | Stratigraphic position | Material | Notes | Images |
Barberenasuchus [67] | B. brasiliensis | Novo Cabrais | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Poorly preserved skull and axis vertebra | An archosauriform of uncertain affinities | |
Brasinorhynchus [68] | B. mariantensis | Bom Retiro do Sul | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | A complete skull, atlas, axis, third neck vertebra | A stenaulorhynchine rhynchosaur archosauromorph | |
Candelaria | C. barbouri | Dona Francisca, Candelária, Novo Cabrais | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | Cranial and postcranial remains | An owenetiid procolophonoid | |
Cerritosaurus [69] | C. binsfeldi | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Badly preserved skull and postcranial elements | A proterochampsid archosauriform | |
Clevosaurus | C. hadroprodon | Candelária | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon Zone | Cranial remains | A clevosaurid rhynchocephalian | |
Hyperodapedon [70] [15] | H. sanjuanensis H. mariensis? H. huenei | Santa Maria, Agudo, Candelária, São João do Polêsine, Venâncio Aires, Restinga Sêca, São Pedro do Sul | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone | Complete skulls, and most postcranial elements | A hyperodapedontine rhynchosaur archosauromorph | |
Kuruxuchampsa [71] | K. dornellesi | Middle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage Zone | A proterochampsid archosauriform. Previously referred to Chanaresuchus [72] | |||
Macrocephalosaurus | M. mariensis | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone | Skull and front part of the torso | A hyperodapedontine rhynchosaur archosauromorph. Sometimes referred to Hyperodapedon | ||
Pinheirochampsa [71] | P. rodriguesi | Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone | A proterochampsid archosauriform | |||
Proterochampsa | P. nodosa | Candelária | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Skull and partial postcranial elements | A proterochampsid archosauriform | |
Rhadinosuchus [73] | R. gracilis | Santa Maria | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme Zone | Badly preserved skull and postcranial elements | A proterochampsid archosauriform | |
Stenoscelida [74] | S.aurantiacus | Agudo | Upper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon Zone | Almost complete and well preserved hind limb | A proterochampsid archosauriform | |
Unnamed proterochampsid [75] | - | Unknown | Santa Maria Formation, Alemoa Member | Badly preserved skull | A proterochampsid archosauriform | |
Probainognathidae is an extinct family of insectivorous cynodonts which lived in what is now South America during the Middle to Late Triassic. The family was established by Alfred Romer in 1973 and includes two genera, Probainognathus from the Chañares Formation of Argentina and Bonacynodon from the Dinodontosaurus Assemblage Zone of Brazil. Probainognathids were closely related to the clade Prozostrodontia, which includes mammals and their close relatives.
Eoraptor is a genus of small, lightly built, basal sauropodomorph dinosaur. One of the earliest-known dinosaurs and one of the earliest sauropodomorphs, it lived approximately 231 to 228 million years ago, during the Late Triassic in Western Gondwana, in the region that is now northwestern Argentina. The type and only species, Eoraptor lunensis, was first described in 1993, and is known from an almost complete and well-preserved skeleton and several fragmentary ones. Eoraptor had multiple tooth shapes, which suggests that it was omnivorous.
Unaysaurus is a genus of unaysaurid sauropodomorph herbivore dinosaur. Discovered in southern Brazil, in the geopark of Paleorrota, in 1998, and announced in a press conference on Thursday, December 3, 2004, it is one of the oldest dinosaurs known. It is closely related to plateosaurid dinosaurs found in Germany, which indicates that it was relatively easy for species to spread across the giant landmass of the time, the supercontinent of Pangaea.
Herrerasauridae is a family of carnivorous dinosaurs, possibly basal to either theropods or even all of saurischians, or even their own branching from Dracohors, separate from Dinosauria altogether. They are among the oldest known dinosaurs, first appearing in the fossil record around 233.23 million years ago, before becoming extinct by the end of the Carnian stage. Herrerasaurids were relatively small-sized dinosaurs, normally no more than 4 metres (13 ft) long, although the holotype specimen of "Frenguellisaurus ischigualastensis" is thought to have reached around 6 meters long. The best known representatives of this group are from South America, where they were first discovered in the 1930s in relation to Staurikosaurus and 1960s in relation to Herrerasaurus. A nearly complete skeleton of Herrerasaurus ischigualastensis was discovered in the Ischigualasto Formation in San Juan, Argentina, in 1988. Less complete possible herrerasaurids have been found in North America and Africa, and they may have inhabited other continents as well.
Lewisuchus is a genus of archosaur that lived during the Late Triassic. As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs. Lewisuchus was about 1 metre (3.3 ft) long. Fossils have been found in the Chañares Formation of Argentina. It exhibited osteoderms along its back.
Exaeretodon is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. E. argentinus is from the Carnian-age Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina. E. major and E. riograndensis are from the Carnian-age portion of the Santa Maria Formation of the Paraná Basin in southeastern Brazil. E. statisticae is from the Carnian-age Lower Maleri Formation of India.
The Ischigualasto Formation is a Late Triassic geological formation in the Ischigualasto-Villa Unión Basin of southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation dates to the late Carnian and early Norian stages of the Late Triassic, according to radiometric dating of ash beds.
The Caturrita Formation is a rock formation found in Rio Grande do Sul, Brazil. Its sediments were deposited in the Paraná Basin. The formation is from the Upper Triassic and forms part of the Santa Maria Supersequence in the upper section of the Rosário do Sul Group.
Saturnaliidae is a family of basal sauropodomorph dinosaurs found in Brazil, Argentina and possibly Zimbabwe. It is not to be confused with Saturnalidae, a family of radiolarian protists.
Gomphodontosuchinae is a subfamily of Triassic traversodontid cynodonts. It includes the genera Gomphodontosuchus, Exaeretodon, Santagnathus, Siriusgnathus, Menadon, Proexaeretodon,Protuberum, Ruberodon and Scalenodontoides.
Pampadromaeus is an extinct genus of basal sauropodomorph dinosaurs known from the Late Triassic (Carnian) Santa Maria Formation of the Paraná Basin in Rio Grande do Sul, southern Brazil.
Botucaraitherium is an extinct genus of prozostrodontian cynodonts from the early Norian Riograndia Assemblage Zone in the Candelária Formation of the Paraná Basin in southeastern Brazil. It is known from a single type species, Botucaraitherium belarminoi. The genus name is derived from the Botucaraí Hill, which dominates the landscape of Candelária, Rio Grande do Sul. The species epithet honors Belarmino Stefanello, a volunteer at the Museu Municipal Aristides Carlos Rodrigues, who found the fossil.
Buriolestes is a genus of early sauropodomorph dinosaurs from the Late Triassic Santa Maria Formation of the Paraná Basin in southern Brazil. It contains a single species, B. schultzi, named in 2016. The type specimen was found alongside a specimen of the lagerpetid dinosauromorph Ixalerpeton.
Macrocollum is a genus of unaysaurid sauropodomorph dinosaur that lived during the Late Triassic period in what is now Brazil. It is one of the oldest dinosaurs known.
Nhandumirim is a genus of basal sauropodomorph dinosaur from the Carnian age of Late Triassic Brazil. It is currently considered a saturnaliid sauropodomorph. The type and only species, Nhandumirim waldsangae, is known from a single immature specimen including vertebrae, a chevron, pelvic material, and a hindlimb found in the Santa Maria Formation in Rio Grande do Sul.
The Candelária Formation, in other literature also referred to as Candelária Sequence, is a sedimentary formation of the Santa Maria Group in the Paraná Basin in Rio Grande do Sul, southeastern Brazil. The formation dates to the Carnian of the Late Triassic, locally referred to as Tuvalian, from 231.4 to approximately 222 Ma.
Erythrovenator is a genus of basal theropod dinosaurs from the Late Triassic of Rio Grande so Sul, Brazil. The genus contains a single species, Erythrovenator jacuiensis.
Bonacynodon is an extinct genus of cynodonts that lived in what is now southern Brazil during the Triassic period. The genus is monotypic, containing only the type species Bonacynodon schultzi. B. schultzi is known from two specimens, consisting of two partial skulls and some badly preserved parts of the postcranium. Both specimens were recovered from the Pinheiros-Chiniquá Sequence, part of the Santa Maria Supersequence of the Paraná Basin. This sequence preserves a faunal association known as the Dinodontosaurus Assemblage Zone, which contains numerous other species of cynodonts, dicynodonts and reptiles. Bonacynodon was a small, likely insectivorous cynodont, whose length has been estimated at around 30 centimetres (12 in). It can be distinguished from other cynodonts by its large, serrated (saw-like) canine teeth. Together with the genus Probainognathus of Argentina, it made up the family Probainognathidae, one of the earliest-diverging lineages of the clade Probainognathia. It was a fairly close relative of mammals, the only group of cynodonts alive today.
Gamatavus is a genus of silesaurid dinosauriform from the Middle Triassic Santa Maria Formation of Dilermando de Aguiar Municipality, Brazil. The genus contains a single species, G. antiquus, known from a partial ilium. Gamatavus represents one of the oldest silesaurids known from South America, alongside the roughly coeval Gondwanax.
Gondwanax is an extinct genus of silesaurid dinosauriform from the Triassic Pinheiros-Chiniquá Sequence of Brazil. The genus contains a single species, G. paraisensis, known from a partial skeleton. Gondwanax represents one of the oldest known dinosauromorphs, and, alongside the roughly coeval Gamatavus, one of the oldest South American silesaurs. While the possession of two sacral vertebrae characterizes more basal "silesaurid" taxa, Gondwanax has three—the oldest occurrence of this trait in the fossil record.
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