Santa Maria Formation

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Santa Maria Formation
Stratigraphic range: Carnian
~237–228  Ma
Schoenstatt Outcrop.png
The Schoenstatt Outcrop of the Santa Maria Formation, which is the type locality of Dagasuchus santacruzensis . Photographed around 2006. [1] [2]
Type Geological formation
Unit of Rosário do Sul Group
Sub-unitsPasso das Tropas & Alemoa Members
Underlies Caturrita Formation
Overlies Sanga do Cabral Formation
Location
Coordinates 29°41′42″S53°47′43″W / 29.695042°S 53.795403°W / -29.695042; -53.795403
RegionBandeira do Rio Grande do Sul.svg  Rio Grande do Sul
CountryFlag of Brazil.svg  Brazil
Extent Paraná Basin
Santa Maria Formation. Source: UFSM FormacaoX.PNG
Santa Maria Formation. Source: UFSM

The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age [3] [4] [5] (Late Triassic), [6] [7] and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, [8] [9] including the herrerasaurid Staurikosaurus , the basal sauropodomorphs Buriolestes and Saturnalia, and the lagerpetid Ixalerpeton . [10] [11] [12] [13] The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied.

Contents

The Santa Maria Formation makes up the majority of the Santa Maria Supersequence, which extends through the entire Late Triassic. The Santa Maria Supersequence is divided into four geological sequences, separated from each other by short unconformities. The first two of these sequences (Pinheiros-Chiniquá and Santa Cruz sequences) lie entirely within the Santa Maria Formation, while the third (the Candelária sequence) is shared with the overlying Norian-age Caturrita Formation. The fourth and youngest sequence (the Mata sequence) is equivalent to the Rhaetian-age Mata Sandstone. [7]

The oldest sequence in the formation is the Pinheiros-Chiniquá Sequence (latest Ladinian-earliest Carnian, ~237 Ma), which is biostratigraphically equivalent to the Dinodontosaurus Assemblage Zone. It is followed by the shorter Santa Cruz Sequence (early Carnian-middle Carnian, ~236 Ma), [3] biostratigraphically equivalent to the Santacruzodon Assemblage Zone. [7]

The final sequence, which is only partially present within the formation, is the Candelária Sequence (middle Carnian-latest Carnian, ~233-228 Ma). [4] [14] [5] [6] [15] The lower portion of this sequence, coinciding with the upper part of the Santa Maria Formation, is equivalent to the Hyperodapedon Assemblage Zone.The Hyperodapedon Assemblage Zone is itself subdivided into Hyperodapedon Acme Zone (most of the zone, where the rhynchosaur Hyperodapedon is widely reported) and Exaeretodon Zone (restricted to about three known and sampled localities, where rhynchsaurs are almost completely absent, but the traversodontid cynodont Exaeretodon is widely reported). [16] [7] These subdivisions are also known as Lower and Upper Hyperodapedon Assemblage Zone, respectively. [15] [16]

U-Pb radiometric dating of Cerro da Alemoa (the type locality of Saturnalia tupiniquim ) in the Upper portion of the Santa Maria Formation found an estimated age of 233.23±0.73 million years ago, putting that locality 1.5 million years older than the Ischigualasto Formation and younger than Los Chañares Formation. The Santa Maria and Ischigualasto formations are approximately equal as having the earliest dinosaur localities. [4] [9] [17] [18] [19]

Vertebrate paleofauna

Most of the information below was included on a revision of the Triassic faunal successions of the Paraná Basin, southern Brazil by Schultz et al. (2020). [7]

Temnospondyls

Temnospondyls of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Compsocerops [20] C. sp. [21] Restinga SêcaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneA partial skullA chigutisaurid temnospondyl
Compsocerops1DB.jpg

Fish

Temnospondyls of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Ptychoceratodus [5] P. cf. phillipsiUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneA ptychoceratodontid lungfish

Synapsids

Dicynodonts

Dicynodonts of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Dinodontosaurus D.pedroanumSão Pedro do Sul, Candelária, Dona Francisca, Bom Retiro, Vale Verde, Novo Cabrais, Bom Retiro do Sul, Dilermando de AguiarLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial and postcranial remainsA kannemeyeriiform dicynodont Dinodontosaurus1DB.jpg
Stahleckeria S.potensDona Francisca, CandeláriaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial and postcranial remainsA stahleckeriid dicyndont Stahleckeria1DB.jpg

Cynognathians

Cynognathians of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Exaeretodon [22] E. majorSão Pedro do SulLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial and postcranial remainsA traversodontid cynodont
E. riograndensisAgudo, Santa MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon ZoneCranial and postcranial remainsA traversodontid cynodont
Exaeretodon BW.jpg
Gomphodontosuchus [23] G. brasiliensisSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneCranial remainsA traversodontid cynodont
Luangwa L.sudamericanaVera Cruz, CandeláriaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial remainsA traversodontid cynodont
Massetognathus M. ochagavieBom Retiro, Vale Verde, Dona FranciscaLower and Middle Santa Maria Formation, Alemoa Member, Santa Cruz nad Pinheiros-Chiniquá Sequences, Santacruzodon Assemblage Zone and Dinodontosaurus Assemblage ZoneCranial and postcranial remainsA traversodontid cynodont
Menadon M.besairieiSanta CruzMiddle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage ZoneCranial remainsA traversodontid cynodont
Protuberum [24] P. cabralensisParaíso do Sul, Novo CabraisLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZonePartial articulated skeleton including skull, vertebrae and ribsA traversodontid cynodont
Santacruzodon [25] S. hopsoni [25] Santa CruzMiddle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage ZoneCranial remains [25] A traversodontid cynodont [25]
Santagnathus [26] S. mariensis Hyperodapedon Assemblage ZoneNumerous cranial and postcranial remainsA traversodontid cynodont
Scalenodon S.ribeiroaeAgudoLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial remainsA traversodontid cynodont
Traversodon T.stahleckeriSão Pedro do SulLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial remainsA traversodontid cynodont

Probainognathians

Probainognathians of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Alemoatherium [27] A. huebneriSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneA partial dentaryA prozostrodontian cynodont Alemoatherium jaw diagram.png
Aleodon A.cromptoniVale Verde, Candelária, Bom Retiro do Sul, Novo CabraisLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial and postcranial remainsA chiniquodontid cynodont
Bonacynodon B.schultziCandeláriaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial remainsA probainognathid cynodont Bonacynodon witout visible ears.png
Candelariodon [28] C. barberenaiCandeláriaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneA partial mandible having some complete teethA basal probainognathian cynodont
Charruodon [29] C.tetracuspidatusCandeláriaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneA partial dentaryA basal probainognathian cynodont [30]
Chiniquodon [23] C. theotonicusSão Pedro do SulLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneSeveral cranial remainsA chiniquodontid cynodont Chiniquodon NT small.jpg
C. sp.Santa CruzMiddle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage ZoneCranial and postcranial remainsA chiniquodontid cynodont
Protheriodon P.estudiantiDona FranciscaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial remainsA basal probainognathian cynodont
Prozostrodon [31] P. brasiliensisSanta Maria, São João do PolêsineUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneCranial remainsA prozostrodontian cynodont
Santacruzgnathus [32] S. abdalaiSanta CruzMiddle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage ZoneCranial remainsA prozostrodontian cynodont
Therioherpeton [33] T. cargniniSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneCranial remainsA prozostrodontian cynodont
Trucidocynodon [34] T. riograndensisAgudoUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon ZoneCranial and postcranial elements of an almost complete skeleton and an additional large skullAn ecteniniid cynodont
Trucidocynodon riograndensis.jpg

Reptiles

Pseudosuchians

Pseudosuchians of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Aetobarbakinoides [35] A. brasiliensisSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZonePartial postcranial skeletonA aetosaur
Aetosauroides [36] A. scagliaiSanta Maria, São João do Polêsine, São Pedro do SulUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneAn almost complete skull and incomplete postcranial elementsAn aetosaur
Aetosauroides scagliai life restoration.jpg
Archeopelta A. arborensisSão Pedro do SulLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial and postcranial elementsAn erpetosuchid
Dagasuchus [2] D. santacruzensisSanta Cruz do SulMiddle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage ZoneAn ilium and a pair of ischiaA loricatan
Decuriasuchus [37] D. quartacoloniaDona FranciscaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneAssemblage of several individuals, which together preserved virtually all the skeletonA loricatan
Decuriasuchus.jpg
Dynamosuchus [38] D. collisensisAgudoUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon ZoneSemi-articulated partial skull and disarticulated fragmentary postcranial skeletonAn ornithosuchid
Dynamosuchus Castro life restoration.png
Pagosvenator [39] P. candelariensisCandeláriaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZonePoorly preserved skull and cervical elementsAn erpetosuchid
Parvosuchus P. aurelioiLinha Várzea 2, Pinheiros-Chiniquá.Dinodontosaurus Assemblage Zone.A largely articulated skeleton, skull, and lower jaws.A gracilisuchid
Parvosuchus skull.png
"Polesinesuchus" [40] "P. aurelioi"São João do PolêsineUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneSeveral postcranial elementsAn immature aetosaur, possible synonym of Aetosauroides scagliai [41]
Prestosuchus [42] [43] [44] P. chiniquensis
"P. loricatus""Karamuru vorax"
Candelária, Dona Francisca, São Pedro do Sul, and possibly Vale Verde ("K. vorax")Lower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneVirtually all the skeleton is knownA loricatan
Prestosuchus-chiniquensis (2).jpg
Procerosuchus [45] P. celerSão Pedro do SulLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZonePostcranial elementsA loricatan
Rauisuchus [46] R. tiradentesSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneIncomplete skeleton including skull elements, several vertebrae and partial hindlimbA loricatan
Rauisuchus tiradentes.jpg
Unnamed loricatan [47] -Santa MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneSpecimen UFSM 11617, an almost complete right maxilla A loricatan

Avemetatarsalians

Avemetatarsalians of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Amanasaurus [48] A. nesbittiRestinga SêcaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneTwo partial femoraA silesaurid
Bagualosaurus [49] B. agudoensisAgudoUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon ZoneAlmost complete skull with lower jaws, several vertebrae, partial pelvic girdle, partial hindlimbsA basal sauropodomorph
Bagualosaurus NT.jpg
Buriolestes [50] [51] B. schultziSão João do PolêsineUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneParts of the skull, vertebrae, left forelimb, and left hindlimbThe basalmost sauropodomorph
Buriolestes schultzi restoration.png
Gamatavus [52] G. antiquus Dilermando de AguiarLower Santa Maria Formation ('Picada do Gama' site), Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneA partial right ilium; a partial left femur and four incomplete vertebrae found in association with the holotype may belong to Gamatavus, but they were not assignedOne of the oldest known silesaurids from South America
Gnathovorax [53] G. cabreiraiSão João do PolêsineUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneA complete semi-articulated skeleton, part of a multiataxic fossil assemblage, including two cynodonts and two rhynchosaursA herrerasaurid
Gnathovorax cabreirai J..jpg
Gondwanax [54] G. paraisensisParaíso do SulLower Santa Maria Formation ('Linha Várzea 2' site), Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneAssociated remains possibly belonging to multiple individuals, including cervical, dorsal, sacral, and caudal vertebrae, a partial pelvic girdle, and a femurOne of the oldest known silesaurids from South America
Ixalerpeton [50] I. polesinensisSão João do PolêsineUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZonePartial skull including a partial maxilla and both dentaries, most of the presacral vertebrae, two sacral vertebrae, several caudal verterae and associated chevrons, scapula and humerus, femora of at least two individuals, and tibiaA basal lagerpetid
Ixalerpeton models.jpg
Nhandumirim [55] N. waldsangaeSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneSome vertebrae, partial pelvic girdle, partial hindlimbA juvenile saturnaliid sauropodomorph [56] [57] [58] [59] [60]
Pampadromaeus [61] P. barberenaiAgudoUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon ZoneAlmost complete skull and lower jaws, several vertebrae, partial pectoral girdle, partial forelimb, partial pelvic girdle, partial hindlimbA basal sauropodomorph
Pampadromaeus life.png
Saturnalia [13] S. tupinquimSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneThree partial skeleton, including partial skull and lower jaw, complete cervical, dorsal and partial caudal vertebral series, pectoral girdle, partial forelimb, pelvic girdle, partial hindlimbA basal sauropodomorph
Saturnalia tupiniquim Saturnalia tupiniquim.jpg
Saturnalia tupiniquim
Spondylosoma [62] S. absconditumSão Pedro do SulLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneSeveral postcranial remainsAn aphanosaurian
Staurikosaurus [10] [11] S. priceiSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZonePartial postcranial skeleton with mandibleA basal herrerasaurid
Staurikosaurus new NT.jpg
Venetoraptor [63] V. gassenaeSão João do PolêsineUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZonePartially articulated skeleton consisting of an incomplete skull, cervical, dorsal, and caudal vertebrae, a partial pelvic girdle, and elements of the arms and legsA lagerpetid.
Unnamed dinosauromorph [64] -Dona FranciscaLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneSpecimen CAPPA/UFSM 0282, a complete, but poorly preserved right femur The oldest dinosauromorph from South America
Unnamed herrerasaurid [65] -Santa MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneSpecimen UFSM 11330, comprised by partial cranial and postcranial remainsThe largest Triassic dinosaur skeleton found in Rio Grande do Sul state so far
Gnathofodaofodao.png
Unnamed lagerpetid [9] -Santa MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneSpecimen UFSM 11611 (partial left femur)A basal lagerpetid distinct from Ixalerpeton
Unnamed silesaurid [9] [66] -Santa MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneSpecimen UFSM 11579, comprised by cranial and postcranial remainsA silesaurid

Other reptiles

Other reptiles of the Santa Maria Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Barberenasuchus [67] B. brasiliensisNovo CabraisUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZonePoorly preserved skull and axis vertebraAn archosauriform of uncertain affinities
Brasinorhynchus [68] B. mariantensisBom Retiro do SulLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneA complete skull, atlas, axis, third neck vertebraA stenaulorhynchine rhynchosaur archosauromorph
Candelaria C. barbouriDona Francisca, Candelária, Novo CabraisLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneCranial and postcranial remainsAn owenetiid procolophonoid
Cerritosaurus [69] C. binsfeldiSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneBadly preserved skull and postcranial elementsA proterochampsid archosauriform
Cerritosaurus.jpg
Clevosaurus C. hadroprodonCandeláriaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon ZoneCranial remainsA clevosaurid rhynchocephalian
Clevosaurus restoration.jpg
Hyperodapedon [70] [15] H. sanjuanensis

H. mariensis?

H. huenei

Santa Maria, Agudo, Candelária, São João do Polêsine, Venâncio Aires, Restinga Sêca, São Pedro do SulUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage ZoneComplete skulls, and most postcranial elementsA hyperodapedontine rhynchosaur archosauromorph
Hyperodapedon NT small.jpg
Kuruxuchampsa [71] K. dornellesiMiddle Santa Maria Formation, Alemoa Member, Santa Cruz Sequence, Santacruzodon Assemblage ZoneA proterochampsid archosauriform. Previously referred to Chanaresuchus [72]
Kuruxuchampsa.jpg
Macrocephalosaurus M. mariensisUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage ZoneSkull and front part of the torsoA hyperodapedontine rhynchosaur archosauromorph. Sometimes referred to Hyperodapedon
Pinheirochampsa [71] P. rodriguesiLower Santa Maria Formation, Alemoa Member, Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage ZoneA proterochampsid archosauriform
Pinheirochampsa.jpg
Proterochampsa P. nodosaCandeláriaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneSkull and partial postcranial elementsA proterochampsid archosauriform
Proterochampsa.jpg
Rhadinosuchus [73] R. gracilisSanta MariaUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Hyperodapedon Acme ZoneBadly preserved skull and postcranial elementsA proterochampsid archosauriform
Rhadinosuchus.jpg
Stenoscelida [74] S.aurantiacusAgudoUpper Santa Maria Formation, Alemoa Member, Candelária Sequence, Hyperodapedon Assemblage Zone, Exaeretodon ZoneAlmost complete and well preserved hind limbA proterochampsid archosauriform
Stenoscelida.jpg
Unnamed proterochampsid [75] -UnknownSanta Maria Formation, Alemoa MemberBadly preserved skullA proterochampsid archosauriform

Formations

Formacoes Geologicas Paleorrota.jpg
Geological formations in Rio Grande do Sul:

See also

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Buriolestes is a genus of early sauropodomorph dinosaurs from the Late Triassic Santa Maria Formation of the Paraná Basin in southern Brazil. It contains a single species, B. schultzi, named in 2016. The type specimen was found alongside a specimen of the lagerpetid dinosauromorph Ixalerpeton.

<i>Macrocollum</i> Extinct genus of dinosaurs

Macrocollum is a genus of unaysaurid sauropodomorph dinosaur that lived during the Late Triassic period in what is now Brazil. It is one of the oldest dinosaurs known.

<i>Nhandumirim</i> Genus of reptiles (fossil)

Nhandumirim is a genus of basal sauropodomorph dinosaur from the Carnian age of Late Triassic Brazil. It is currently considered a saturnaliid sauropodomorph. The type and only species, Nhandumirim waldsangae, is known from a single immature specimen including vertebrae, a chevron, pelvic material, and a hindlimb found in the Santa Maria Formation in Rio Grande do Sul.

<span class="mw-page-title-main">Candelária Formation, Paraná Basin</span> Sedimentary formation in Brazil

The Candelária Formation, in other literature also referred to as Candelária Sequence, is a sedimentary formation of the Santa Maria Group in the Paraná Basin in Rio Grande do Sul, southeastern Brazil. The formation dates to the Carnian of the Late Triassic, locally referred to as Tuvalian, from 231.4 to approximately 222 Ma.

Erythrovenator is a genus of basal theropod dinosaurs from the Late Triassic of Rio Grande so Sul, Brazil. The genus contains a single species, Erythrovenator jacuiensis.

<i>Bonacynodon</i> Extinct genus of cynodonts

Bonacynodon is an extinct genus of cynodonts that lived in what is now southern Brazil during the Triassic period. The genus is monotypic, containing only the type species Bonacynodon schultzi. B. schultzi is known from two specimens, consisting of two partial skulls and some badly preserved parts of the postcranium. Both specimens were recovered from the Pinheiros-Chiniquá Sequence, part of the Santa Maria Supersequence of the Paraná Basin. This sequence preserves a faunal association known as the Dinodontosaurus Assemblage Zone, which contains numerous other species of cynodonts, dicynodonts and reptiles. Bonacynodon was a small, likely insectivorous cynodont, whose length has been estimated at around 30 centimetres (12 in). It can be distinguished from other cynodonts by its large, serrated (saw-like) canine teeth. Together with the genus Probainognathus of Argentina, it made up the family Probainognathidae, one of the earliest-diverging lineages of the clade Probainognathia. It was a fairly close relative of mammals, the only group of cynodonts alive today.

Gamatavus is a genus of silesaurid dinosauriform from the Middle Triassic Santa Maria Formation of Dilermando de Aguiar Municipality, Brazil. The genus contains a single species, G. antiquus, known from a partial ilium. Gamatavus represents one of the oldest silesaurids known from South America, alongside the roughly coeval Gondwanax.

<i>Gondwanax</i> Genus of silesaurid dinosauromorphs

Gondwanax is an extinct genus of silesaurid dinosauriform from the Triassic Pinheiros-Chiniquá Sequence of Brazil. The genus contains a single species, G. paraisensis, known from a partial skeleton. Gondwanax represents one of the oldest known dinosauromorphs, and, alongside the roughly coeval Gamatavus, one of the oldest South American silesaurs. While the possession of two sacral vertebrae characterizes more basal "silesaurid" taxa, Gondwanax has three—the oldest occurrence of this trait in the fossil record.

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