Ixalerpeton Temporal range: Carnian ~ | |
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Skeletal diagram of Ixalerpeton polesinensis, known remains depicted in white and light grey, unknown (in dark grey) is reconstructed after other lagerpetids such as Lagerpeton and Dromomeron . | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Family: | † Lagerpetidae |
Genus: | † Ixalerpeton Cabreira et al., 2016 |
Species: | †I. polesinensis |
Binomial name | |
†Ixalerpeton polesinensis Cabreira et al., 2016 | |
Ixalerpeton (meaning "leaping reptile") is a genus of lagerpetid avemetatarsalian containing one species, I. polesinensis. It lived in the Late Triassic of Brazil alongside the sauropodomorph dinosaur Buriolestes . [1]
The holotype specimen of Ixalerpeton, numbered ULBRA-PVT059, consists of parts from the skull, vertebral column, and all four limbs. The specimen comes from the Carnian Santa Maria Formation of Brazil, and it was found alongside two individuals of Buriolestes as well as a set of femora belonging to second individual of Ixalerpeton. The genus name of Ixalerpeton combines the Greek words ixalos ("leaping") and erpeton ("reptile"), and the species name polesinensis references the town of São João do Polêsine, where the dig site is located. [1]
Ixalerpeton was similar to other lagerpetids (namely Dromomeron and Lagerpeton ) in having long hindlimbs with well-developed muscle attachments on the femur; in particular, its fourth trochanter was quite large and formed a crest (which is unlike Dromomeron). [1] However, the last few dorsal vertebrae of Ixalerpeton do not have the forward-inclining neural spines of Lagerpeton (which were associated with the latter's hopping, or saltatory, lifestyle). [1] [2] In addition to the enlarged fourth trochanter, a suite of other traits differentiate Ixalerpeton from all previously-described lagerpetids; there is an antitrochanter on the ilium; the end of the shaft of the ischium is tall; there is no ambiens process on the pubis; the medial condyle on the femur is relatively flat at the front end but sharply angled at the back end; and the back face of the top end of the tibia has a deep groove.
The head and forelimbs found with Ixalerpeton are the first of these elements that have been found among lagerpetids. [3] Unlike dinosauriforms, the posttemporal fenestra at the back of the skull is large and unreduced; there is an extra bone, the postfrontal, bordering the eye socket; there is no supratemporal fossa, which is an indentation found on the frontal bone in dinosauriforms; and the glenoid cavity on the scapula, where the scapula-humerus joint is located, faces slightly sideways instead of backwards. On the other hand, there is an anterior tympanic recess on the braincase, and the deltopectoral crest on the humerus is long, both of which are common among basal dinosauromorphs. [1]
A 2016 phylogenetic analysis found that Ixalerpeton was the closest relative of Dromomeron . The phylogenetic tree recovered is partially reproduced below. [1]
Eucrocopoda |
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Similar to the Ischigualasto [4] and Chinle [5] Formations, the Santa Maria Formation preserves both Ixalerpeton (a non-dinosaur avemetatarsalian) and Buriolestes (a dinosaur). This indicates that dinosaurs did not rapidly replace related groups once they had evolved. [1]
Eoraptor is a genus of small, lightly built, basal sauropodomorph dinosaur. One of the earliest-known dinosaurs and one of the earliest members of the sauropod family, it lived approximately 231 to 228 million years ago, during the Late Triassic in Western Gondwana, in the region that is now northwestern Argentina. The type and only species, Eoraptor lunensis, was first described in 1993, and is known from an almost complete and well-preserved skeleton and several fragmentary ones. Eoraptor had multiple tooth shapes, which suggests that it was omnivorous. Eoraptor was 1.5 feet tall and 3 feet long.
Sauropodomorpha is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The prosauropods, which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic until their decline and extinction at the end of the Cretaceous.
Staurikosaurus is a genus of herrerasaurid dinosaur from the Late Triassic of Brazil, found in the Santa Maria Formation.
Herrerasaurus is likely a genus of saurischian dinosaur from the Late Triassic period. This genus was one of the earliest dinosaurs from the fossil record. Its name means "Herrera's lizard", after the rancher who discovered the first specimen in 1958 in South America. All known fossils of this carnivore have been discovered in the Ischigualasto Formation of Carnian age in northwestern Argentina. The type species, Herrerasaurus ischigualastensis, was described by Osvaldo Reig in 1963 and is the only species assigned to the genus. Ischisaurus and Frenguellisaurus are synonyms.
Lagosuchus is an extinct genus of avemetatarsalian archosaur from the Late Triassic of Argentina. The type species of Lagosuchus, Lagosuchus talampayensis, is based on a small partial skeleton recovered from the early Carnian-age Chañares Formation. The holotype skeleton of L. talampayensis is fairly fragmentary, but it does possess some traits suggesting that Lagosuchus was a probable dinosauriform, closely related to dinosaurs.
Marasuchus is a genus of basal dinosauriform archosaur which is possibly synonymous with Lagosuchus. Both genera lived during the Late Triassic in what is now La Rioja Province, Argentina. Marasuchus contains a single species, Marasuchus lilloensis.
Lagerpeton is a genus of lagerpetid avemetatarsalian, comprising a single species, L. chanarensis. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, Lagerpeton's anatomy is somewhat incompletely known, with fossil specimens accounting for the pelvic girdle, hindlimbs, posterior presacral, sacral and anterior caudal vertebrae. Skull and shoulder material has also been described.
Teleocrater is a genus of avemetatarsalian archosaur from the Middle Triassic Manda Formation of Tanzania. The name was coined by English paleontologist Alan Charig in his 1956 doctoral dissertation, but was only formally published in 2017 by Sterling Nesbitt and colleagues. The genus contains the type and only species T. rhadinus. Uncertainty over the affinities of Teleocrater have persisted since Charig's initial publication; they were not resolved until Nesbitt et al. performed a phylogenetic analysis. They found that Teleocrater is most closely related to the similarly enigmatic Yarasuchus, Dongusuchus, and Spondylosoma in a group that was named the Aphanosauria. Aphanosauria was found to be the sister group of the Ornithodira, the group containing dinosaurs and pterosaurs.
Dinosauromorpha is a clade of avemetatarsalian archosaurs that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, with later formulations specifically excluding pterosaurs from the group. Birds are the only dinosauromorphs which survive to the present day.
Dromomeron is a genus of lagerpetid avemetatarsalian which lived around 220 to 211.9 ± 0.7 million years ago. The genus contains species known from Late Triassic-age rocks of the Southwestern United States and northwestern Argentina. It is described as most closely related to the earlier Lagerpeton of Argentina, but was found among remains of true dinosaurs like Chindesaurus, indicating that the first dinosaurs did not immediately replace related groups.
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age, and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, including the herrerasaurid Staurikosaurus, the basal sauropodomorphs Buriolestes and Saturnalia, and the lagerpetid Ixalerpeton. The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied.
Asilisaurus ; from Swahili, asili, and Greek, σαυρος is an extinct genus of silesaurid archosaur. The type species is Asilisaurus kongwe.Asilisaurus fossils were uncovered in the Manda Beds of Tanzania and date back to the early Carnian, making it one of the oldest known members of the Avemetatarsalia. It was the first non-dinosaurian dinosauriform recovered from Africa. The discovery of Asilisaurus has provided evidence for a rapid diversification of avemetatarsalians during the Middle Triassic, with the diversification of archosaurs during this time previously only documented in pseudosuchians.
Lagerpetidae is a family of basal avemetatarsalians. Though traditionally considered the earliest-diverging dinosauromorphs, fossils described in 2020 suggest that lagerpetids may instead be pterosauromorphs. Lagerpetid fossils are known from the Late Triassic of Argentina, Arizona, Brazil, Madagascar, New Mexico, and Texas. They were typically small, although some lagerpetids, like Dromomeron gigas and a specimen from the Santa Rosa Formation attributed to Dromomeron sp., were able to get quite large. Lagerpetid fossils are rare; the most common finds are bones of the hindlimbs, which possessed a number of unique features.
Eodromaeus is an extinct genus of probable basal theropod dinosaurs from the Late Triassic of Argentina. Like many other of the earliest-known dinosaurs, it hails from the Carnian-age Ischigualasto Formation, within the Ischigualasto-Villa Unión Basin of northwestern Argentina. Upon its discovery, it was argued to be one of the oldest true theropods, supplanting its contemporary Eoraptor, which was reinterpreted as a basal sauropodomorph.
Diodorus is a genus of silesaurid dinosauromorph that lived during the Late Triassic in what is now Morocco. Fossils were discovered in the Timezgadiouine Formation of the Argana Basin, and were used to name the new genus and species Diodorus scytobrachion. The genus name honors the mythological king Diodorus and the ancient historian Diodorus Siculus; the specific name is ancient Greek for 'leathery arm' and also honors the mythographer Dionysius Scytobrachion. The holotype specimen is a partial dentary bone (front of the lower jaw), and assigned specimens include isolated teeth, two humeri (upper arm bones), a metatarsal (foot bone), and femur (thigh bone).
Ignotosaurus is an extinct genus of silesaurid dinosauriform known from the Late Triassic (Carnian) Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina. It was therefore contemporary with early dinosaurs such as Herrerasaurus, and lived in the same place.
Buriolestes is a genus of early sauropodomorph dinosaurs from the Late Triassic Santa Maria Formation of the Paraná Basin in southern Brazil. It contains a single species, B. schultzi, named in 2016. The type specimen was found alongside a specimen of the lagerpetid dinosauromorph Ixalerpeton.
Nhandumirim is a genus of basal sauropodomorph dinosaur from the Carnian age of Late Triassic Brazil. It is currently considered a saturnaliid sauropodomorph. The type and only species, Nhandumirim waldsangae, is known from a single immature specimen including vertebrae, a chevron, pelvic material, and a hindlimb found in the Santa Maria Formation in Rio Grande do Sul.
Kongonaphon is an extinct genus of lagerpetid avemetatarsalians from the Middle to Late Triassic of Madagascar. It contains a single species, Kongonaphon kely, which is known from a fragmentary partial skeleton. This fossil hails from the late Ladinian or early Carnian-age "basal Isalo II beds". As the first lagerpetid found in Africa, Kongonaphon extends the range of the family significantly. It possessed a combination of features from various other lagerpetids, but developed particularly long and slender leg bones. Kongonaphon is also the first lagerpetid for which fossils of the snout and teeth are known. It was likely an insectivore based on the shape and texture of its teeth.
Incertovenator is an extinct genus of archosauriform reptile, likely an archosaur, of uncertain affinities. Its unstable position is a result of possessing a number features found in both the bird-line avemetatarsalian archosaurs and the crocodylian-line pseudosuchians. The type and only known species is I. longicollum, which is known from single specimen discovered in the Late Triassic Ischigualasto Formation of Argentina. Incertovenator is known almost entirely by its vertebral column. This indicates that it had a relatively long neck, leading to its uncertain classification due to the convergent evolution of elongated neck vertebrae in both avemetatarsalian and pseudosuchian archosaurs.