Procerosuchus

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Procerosuchus
Temporal range: Carnian
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauria
Clade: Pseudosuchia
Clade: Loricata
Genus: Procerosuchus
Huene, 1942
Type species
Procerosuchus celer
Huene, 1942

Procerosuchus is an extinct genus of loricatan archosaur. Fossils have been collected from the Late Triassic Santa Maria Formation in Geopark of Paleorrota, Rio Grande do Sul, Brazil, which is Carnian in age. [1] The genus was first described by the German paleontologist Friedrich von Huene in 1942. [2]

Contents

Classification

Initially, Procerosuchus was regarded as a stagonolepidid along with the genera Rauisuchus and Prestosuchus . [2] Later, it was reassigned by Huene to the family Rauisuchidae. Alfred Sherwood Romer first considered Procerosuchus to be a possible ornithosuchid, but later assigned it to the family Prestosuchidae, which he constructed in 1966. [3] [4] In 1972, Romer assigned Procerosuchus as a possible member of the family Proterochampsidae. [5] Krebs (1976) considered it to be a rauisuchid, as did Chatterjee (1985) and Carroll (1988). [6] [7] [8]

Procerosuchus has been suggested to be member of the subfamily Rauisuchinae and the tribe Rauisuchini. [9] However, the genus has not yet been included in any phylogenetic analyses of rauisuchians, and its classification remains uncertain. The alpha taxonomy of rauisuchians is still debated (the order itself is now considered paraphyletic) and the anatomy of many taxa, including Procerosuchus, has not yet been thoroughly described. [10] Procerosuchus does not seem to belong to a recently identified monophyletic grouping of rauisuchians termed Clade X (and now known as Poposauroidea), which includes poposaurids and ctenosauriscids. [11]

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<span class="mw-page-title-main">Alfred Romer</span> American paleontologist

Alfred Sherwood Romer was an American paleontologist and biologist and a specialist in vertebrate evolution.

<span class="mw-page-title-main">Rauisuchia</span> Informal group of Triassic archosaurs with pillar-erect posture

"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.

<i>Postosuchus</i> Genus of reptiles

Postosuchus, meaning "Crocodile from Post", is an extinct genus of rauisuchid reptiles comprising two species, P. kirkpatricki and P. alisonae, that lived in what is now North America during the Late Triassic. Postosuchus is a member of the clade Pseudosuchia, the lineage of archosaurs that includes modern crocodilians. Its name refers to Post Quarry, a place in Texas where many fossils of the type species, P. kirkpatricki, were found. It was one of the apex predators of its area during the Triassic, larger than the small dinosaur predators of its time. It was a hunter which probably preyed on large bulky herbivores like dicynodonts and many other creatures smaller than itself.

<i>Arizonasaurus</i> Extinct genus of reptiles

Arizonasaurus was a ctenosauriscid archosaur from the Middle Triassic. Arizonasaurus is found in the Middle Triassic Moenkopi Formation of northern Arizona. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. The taxon has a large sailback formed by elongated neural spines of the vertebrae. The type species, Arizonasaurus babbitti, was named by Samuel Paul Welles in 1947.

<i>Teratosaurus</i> Extinct genus of reptiles

Teratosaurus is a genus of rauisuchians known from the Triassic Stubensandstein of Germany. It is estimated to be 6.2 meters (20.35 ft) long.

Spinosuchus is an extinct genus of trilophosaurid allokotosaur from the Late Triassic of Texas, southern United States. It has been assigned to a variety of groups over its history, from coelophysid dinosaur to pseudosuchian to uncertain theropod dinosaur and to Proterosuchidae. This uncertainty is not unusual, given that it was only known from a poorly preserved, wall-mounted, partial vertebral column of an animal that lived in a time of diverse, poorly known reptile groups. However, newly collected material and recent phylogenetic studies of early archosauromorphs suggest that it represents an advanced trilophosaurid very closely related to Trilophosaurus.

<span class="mw-page-title-main">Prestosuchidae</span> Extinct family of reptiles

Prestosuchidae is a polyphyletic grouping of carnivorous archosaurs that lived during the Triassic. They were large active terrestrial apex predators, ranging from around 2.5 to 7 metres in length. They succeeded the Erythrosuchidae as the largest archosaurs of their time. While resembling erythrosuchids in size and some features of the skull and skeleton, they were more advanced in their erect posture and crocodile-like ankle, indicating more efficient gait. "Prestosuchids" flourished throughout the whole of the middle, and the early part of the late Triassic, and fossils are so far known from Europe, India, Africa (Tanzania), Argentina, and Paleorrota in Brazil. However, for a long time experts disagree regarding the phylogenetic relationships of the group, what genera should be included, and whether indeed the "Prestosuchidae" constitute a distinct family.

<i>Erythrosuchus</i> Extinct genus of reptiles

Erythrosuchus is an extinct genus of archosauriform reptiles from the early Triassic of South Africa. Remains have been found from the Cynognathus Assemblage Zone of the Beaufort Group in the Karoo of South Africa.

<i>Garjainia</i> Extinct genus of reptiles

Garjainia is an extinct genus of erythrosuchid archosauriform reptile from the Olenekian of Russia and South Africa. It was approximately 1.5–2 metres (4.9–6.6 ft) long. It contained two species, Garjainia prima from the Yarengian/Yarkenskian Supergorizont of Russia, and Garjainia madiba from the Burgersdorp Formation of South Africa. "Vjuskovia triplicostata", a name assigned to some erythrosuchid fossils from Russia, has been synonymized with Garjainia prima.

<i>Proterosuchus</i> Extinct genus of reptiles from the Early Triassic of South Africa

Proterosuchus is an extinct genus of archosauriform reptiles that lived during the Early Triassic. It contains three valid species: the type species P. fergusi and the referred species P. alexanderi and P. goweri. All three species lived in what is now South Africa. The genus was named in 1903 by the South African paleontologist Robert Broom. The genus Chasmatosaurus is a junior synonym of Proterosuchus.

<i>Batrachotomus</i> Genus of reptiles

Batrachotomus is a genus of prehistoric archosaur. Fossils of this animal have been found in southern Germany and dated from the Ladinian stage of the Middle Triassic period, around 242 to 237 million years ago. Batrachotomus was described by palaeontologist David J. Gower 22 years after its discovery.

Yarasuchus is an extinct genus of avemetatarsalian archosaur that lived during the Anisian stage of the Middle Triassic of India. The genus was named and described in 2005 from a collection of disarticulated but fairly complete fossil material found from the Middle Triassic Yerrapalli Formation. The material is thought to be from two individuals, possibly three, with one being much more complete and articulated than the other. The type and only species is Y. deccanensis. Yarasuchus was a quadruped roughly 2–2.5 metres (6.6–8.2 ft) long, with an elongated neck and tall spines on its vertebrae. Unlike other quadrupedal Triassic reptiles, the limbs and shoulders of Yarasuchus were slender, and more like those of ornithodirans.

<i>Dyoplax</i> Extinct genus of reptiles

Dyoplax is an extinct genus of pseudosuchian archosaur, possibly an erpetosuchid. Fossils have been found from the type locality within the upper Schilfsandstein Formation in Stuttgart, Germany. The holotype specimen was a natural cast of a nearly complete skeleton that lacked only parts of the tail and limb bones.

<i>Fasolasuchus</i> Extinct genus of reptiles

Fasolasuchus is an extinct genus of loricatan. Fossils have been found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina that date back to the Norian stage of the Late Triassic, making it one of the last "rauisuchians" to have existed before the order became extinct at the end of the Triassic.

Fenhosuchus is an extinct genus of archosauriform. The holotype, IVPP V 2697, and referred materials have been found in the Hsishihwa locality at Wuhsiang, China, from the Upper Ermaying Formation. The locality dates back to the Anisian stage of the Middle Triassic. The genus was named after the Fen River in Shanxi Province from which specimens were found. It may prove to be a chimera being composed of material from several different animals. Some material were believed to represent a rauisuchid. The calcaneum of Fenhosuchus seems to belong to an erythrosuchid or other basal archosauriform. Much of the material of the tarsal bones seem to be similar to those of the genus Shansisuchus. According to Nesbitt (2009) the assessment of Gower (2000) was correct, the holotype is a mix of Shansisuchus remains and a possible fragment from a paracrocodylomorph or a dinosauriform. Thus, Fenhosuchus cannot be considered a rauisuchian.

<i>Heptasuchus</i> Extinct genus of reptiles

Heptasuchus is an extinct genus of loricatan pseudosuchian known from the Middle or Late Triassic upper Chugwater Group of Wyoming, United States. It contains a single species, Heptasuchus clarki, the first formally recognized "rauisuchian" or loricatan pseudosuchian from North America.

Notochampsa is an extinct genus of protosuchian crocodyliform. Fossils have been found from the lower Clarens Formation of the Karoo Supergroup in South Africa, dating back to the Pliensbachian stage of the Early Jurassic. Notochampsa comes from a period of relative fossil scarcity, and is the youngest known occurrence of a crocodylomorph from the Karoo Basin of South Africa.

<i>Platyognathus</i> Extinct genus of reptiles

Platyognathus is an extinct genus of protosuchian crocodyliform. Fossils are known from the Early Jurassic Lower Lufeng Formation in Yunnan, China and belong to the type and only species, P. hsui.

Stagonosuchus is an extinct genus of paracrocodylomorph, possibly a loricatan synonymous with Prestosuchus or a poposauroid. Fossils have been found from the Late Triassic Manda Formation in Tanzania that are Anisian in age.

<i>Youngosuchus</i> Extinct genus of reptiles

Youngosuchus is an extinct genus of archosaur from the Middle Triassic of China. The type species is Y. sinensis. Y. sinensis was first described in 1973 as a new species of the erythrosuchid Vjushkovia. In 1985, it was reassigned as its own genus of rauisuchid. A 1992 study supported the original classification of Youngosuchus sinensis as an erythrosuchid, but more recent studies classify it as a "rauisuchian"-grade loricatan archosaur completely unrelated to Vjushkovia, which is most likely a synonym of Garjainia.

References

  1. Gower, D. J. (2000). "Rauisuchian archosaurs (Reptilia:Diapsida): An overview" (PDF). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 218 (3): 447–488. doi:10.1127/njgpa/218/2000/447.
  2. 1 2 Huene, F. v. (1942). "Ergebnisse der Sauriergrabungen in Südbrasilien 1928/29". Die fossilen Reptilien des südamerikanischen Gondwanalandes. München: C. H. Beck. p. 332.
  3. Romer, A. S. (1956). Osteology of Reptiles. Chicago: University of Chicago Press. p. 772. ISBN   0-89464-985-X.
  4. Romer, A. S. (1966). Vertebrate Paleontology (3rd ed.). Chicago: University of Chicago Press. pp.  468. ISBN   0-226-72488-3.
  5. Romer, A. S. (1972). "The Chañares (Argentina) Triassic Reptile Fauna. XVI. Thecodont Classification". Breviora. 395: 1–24.
  6. Kuhn, O. (1976). "Pseudosuchia". In Kuhn, O. (ed.). Handbuch der Paläoherpetologie (13th ed.). Stuttgart: G. Fischer Verlag. pp. 40–98. ISBN   0-89574-191-1.
  7. Chatterjee, S. (1985). "Postosuchus, a new thecodontian reptile from the Triassic of Texas and the origin of tyrannosaurs". Philosophical Transactions of the Royal Society B . 309 (1139): 395–460. Bibcode:1985RSPTB.309..395C. doi:10.1098/rstb.1985.0092.
  8. Carroll, R. L. (1988). Vertebrate Paleontology and Evolution . New York: W. H. Freeman and Company. pp.  1–698. ISBN   0-7167-1822-7.
  9. Sennikov, A. G. (1995). "Ranniye Tekodont Vostochnoy Yevropy". Trudy Paleontologicheskogo Instituta. 263. Moscow: 1–141.
  10. Gower, D. J.; Schoch, R. R. (2009). "Postcranial anatomy of the rauisuchian archosaur Batrachotomus kupferzellensis" (PDF). Journal of Vertebrate Paleontology. 29 (1): 103–122. doi:10.1080/02724634.2009.10010365. S2CID   196611289.
  11. Nesbitt, S. J. (2005). "Osteology of the Middle Triassic pseudosuchian archosaur Arizonasaurus babbitti". Historical Biology. 17 (1–4): 19–47. doi:10.1080/08912960500476499. S2CID   84326151.