Sphenosuchia Temporal range: Late Triassic - Late Jurassic, | |
---|---|
Life restoration of Hesperosuchus agilis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Pseudosuchia |
Clade: | Crocodylomorpha |
Informal group: | † Sphenosuchia von Huene, 1942 |
Genera | |
Synonyms | |
|
Sphenosuchia is a suborder of basal crocodylomorphs that first appeared in the Triassic and occurred into the Middle Jurassic. Most were small, gracile animals with an erect limb posture. They are now thought to be ancestral to crocodyliforms, a group which includes all living crocodilians.
The earliest known members of the group (i.e. Hesperosuchus ) are early Norian in age, found in the Blue Mesa Member of the Chinle Formation. Only one sphenosuchian is currently known from the Middle Jurassic, Junggarsuchus , from the Junggar Basin (Shishugou Formation) of China during either the Bathonian or the Callovian (~165 Ma) age, [1] and the Hallopodidae are known from the Late Jurassic of North America. [2]
The monophyly of the group is debated, although several synapomorphies characterize the clade, including extremely slender limbs, a compact carpus and an elongate coracoid process.
In 2002, Clark and Sues found a possible sphenosuchian clade of Dibothrosuchus , Sphenosuchus , and possibly Hesperosuchus and Saltoposuchus , with several other genera in unresolved positions ( Kayentasuchus , Litargosuchus , Pseudhesperosuchus , and Terrestrisuchus ). [3] More recently, however, Clark et al. (2004) argued for the paraphyly of the group, contending that morphological characters were secondarily lost in more highly derived crocodylomorphs. [1] Further analysis and study is required before the group's monophyly is resolved with certainty — a perfect phylogenetic analysis is, at present, impossible due to a paucity of fossil remains demonstrating phylogenetically informative characters.
Below is a cladogram modified from Nesbitt (2011). [4] Sphenosuchians are marked by the green bracket.
| Sphenosuchians |
Genus | Status | Age | Location | Unit | Notes | Images |
---|---|---|---|---|---|---|
Valid | Early Jurassic | China | Lower Lufeng Series | |||
Valid | Late Triassic | USA | Newark Supergroup | |||
Valid | Late Triassic (Carnian) | Germany | Schilfsandstein Formation | A possible sphenosuchian; [5] alternatively, it could be an erpetosuchid. [6] | ||
Valid | Late Triassic (Carnian) | USA | Chinle Formation | |||
Valid | Middle Jurassic | China | Shishugou Formation | |||
Valid | Early Jurassic (Sinemurian - Pliensbachian) | USA | Kayenta Formation | |||
Valid | Early Jurassic | South Africa | Elliot Formation | |||
Nomen dubium | Late Triassic | USA | Dockum Group | An indeterminate sphenosuchian known only from undiagnostic vertebrae [7] | ||
Valid | Early Jurassic | China | Lower Lufeng Series | |||
Valid | Late Triassic (Norian) | Argentina | Los Colorados Formation | |||
Valid | Late Triassic | USA | Redonda Formation | |||
Valid | Late Triassic (Norian) | Germany | Löwenstein Formation | |||
Valid | Early Jurassic | South Africa | Elliot Formation | |||
Valid. | Late Triassic | UK | ||||
Valid | Late Triassic | Argentina | Ischigualasto Formation | |||
Archosauria is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term, which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives; and Avemetatarsalia, which includes birds and their extinct relatives.
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large, carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, phytosaurs, aetosaurs, and crocodylomorphs.
Phytosaurs are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. and are sometimes referred to as parasuchians. Phytosauria, Parasuchia, Parasuchidae, and Phytosauridae have often been considered equivalent groupings containing the same species. Some recent studies have offered a more nuanced approach, defining Parasuchidae and Phytosauridae as nested clades within Phytosauria as a whole. Phytosaurs were long-snouted and heavily armoured, bearing a remarkable resemblance to modern crocodilians in size, appearance, and lifestyle, as an example of convergence or parallel evolution. The name "phytosaur" means "plant reptile", as the first fossils of phytosaurs were mistakenly thought to belong to plant eaters.
Saltoposuchus is an extinct genus of small, long-tailed crocodylomorph reptile (Sphenosuchia), from the Norian of Europe. The name translated means "leaping foot crocodile". It has been proposed that Terrestrisuchus gracilis and Saltoposuchus connectens represent different ontogenetic stages of the same genus. Saltoposuchus was commonly referred to in popular literature as the ancestor to dinosaurs, however, recent scientific research show that this is not the case.
Hallopus was a prehistoric reptile, named in 1877 as a species of Nanosaurus and classified as a genus by O. C. Marsh in 1881 from the Late Jurassic Morrison Formation. Today though the animal is thought to be a pseudosuchian more closely related to crocodilians. It was redescribed as a sphenosuchian crocodylomorph in 1970, now thought to be a paraphyletic group. It was a quite small animal, reaching a length of 1 m (3.3 ft) with long and slender limbs. Macelognathus, a similarly slender-proportioned crocodylomorph to Hallopus, may be synonymous with it.
Poposauridae is a family of large carnivorous archosaurs which lived alongside dinosaurs during the Late Triassic. They were around 2.5 to 5 metres long. Poposaurids are known from fossil remains from North and South America. While originally believed to be theropod dinosaurs, cladistic analysis has shown them to be more closely related to crocodiles.
Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Despite Pseudosuchia meaning "false crocodiles", the name is a misnomer as true crocodilians are now defined as a subset of the group.
Rauisuchidae is a group of large predatory Triassic archosaurs. Some disagreement exists over which genera should be included in the Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed whether these should even be thought of as separate valid families. Rauisuchids occurred throughout much of the Triassic, and may have first occurred in the Early Triassic if some archosaurian taxa such as Scythosuchus and Tsylmosuchus are considered to be within the family.
Prestosuchidae is a polyphyletic grouping of carnivorous archosaurs that lived during the Triassic. They were large active terrestrial apex predators, ranging from around 2.5 to 7 metres in length. They succeeded the Erythrosuchidae as the largest archosaurs of their time. While resembling erythrosuchids in size and some features of the skull and skeleton, they were more advanced in their erect posture and crocodile-like ankle, indicating more efficient gait. "Prestosuchids" flourished throughout the whole of the middle, and the early part of the late Triassic, and fossils are so far known from Europe, India, Africa (Tanzania), Argentina, and Paleorrota in Brazil. However, for a long time experts disagree regarding the phylogenetic relationships of the group, what genera should be included, and whether indeed the "Prestosuchidae" constitute a distinct family.
Terrestrisuchus is an extinct genus of very small early crocodylomorph that was about 76 centimetres (30 in) long. Fossils have been found in Wales and Southern England and date from near the very end of the Late Triassic during the Rhaetian, and it is known by type and only known species T. gracilis. Terrestrisuchus was a long-legged, active predator that lived entirely on land, unlike modern crocodilians. It inhabited a chain of tropical, low-lying islands that made up southern Britain, along with similarly small-sized dinosaurs and abundant rhynchocephalians. Numerous fossils of Terrestrisuchus are known from fissures in limestone karst which made up the islands it lived on, which formed caverns and sinkholes that preserved the remains of Terrestrisuchus and other island-living reptiles.
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction.
Litargosuchus is a sphenosuchian crocodylomorph, a basal member of the crocodylomorph clade from the Early Jurassic of South Africa. Its genus name Litargosuchus is derived from Greek meaning "fast running crocodile" and its species name leptorhynchus refers to its gracile snout. Litargosuchus, along with all of South Africa's crocodylomorph taxa, are confined to the upper Elliot Formation (UEF) in South Africa.
Dromicosuchus is an extinct genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It was found in Upper Triassic rocks of North Carolina, United States, and is known from a nearly complete skull and partial skeleton. This specimen is unusual in that it was found beneath the skeleton of a larger rauisuchian and has apparent bite damage, suggesting that it was attacked by the larger carnivore before both died and were buried together.
Pedeticosaurus is an extinct genus of crocodylomorph from the Clarens Formation of South Africa. The type species Pedeticosaurus leviseuri was named by Egbert Cornelis Nicolaas van Hoepen in 1915 on the basis of a mold of a mostly complete skeleton found in a quarry near Rosendal, Free State. The mold preserves most of the right half of the skeleton including the skull, ribs, dorsal vertebrae, forelimbs and hindlimbs, but not the tail. It is currently housed in the National Museum in Bloemfontein and cataloged as NMQR606.
Junggarsuchus is an extinct genus of sphenosuchian crocodylomorph from the Middle or Late Jurassic period of China. The type and only species is J. sloani. The generic name of Junggarsuchus comes from the Junggar Basin, where the fossil was found, and the Greek word "souchos" meaning crocodile. The specific name, "sloani" is in honor of C. Sloan, who is credited with finding the holotype.
Phyllodontosuchus is a genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It is known from a skull and jaws from Lower Jurassic rocks of Yunnan, China. Phyllodontosuchus is unusual because some of its teeth were leaf-shaped, like those of some herbivorous dinosaurs, and it does not appear to have been a strict carnivore like most other crocodylomorphs.
Dyoplax is an extinct genus of pseudosuchian archosaur, possibly an erpetosuchid. Fossils have been found from the type locality within the upper Schilfsandstein Formation in Stuttgart, Germany. The holotype specimen was a natural cast of a nearly complete skeleton that lacked only parts of the tail and limb bones.
Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as Postosuchus and Prestosuchus. More specifically, Loricata includes Crocodylomorpha and most "rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs.
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing Poposaurus and Crocodylus niloticus. Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic.
Erpetosuchidae is an extinct family of pseudosuchian archosaurs. Erpetosuchidae was named by D. M. S. Watson in 1917 to include Erpetosuchus. It includes the type species Erpetosuchus granti from the Late Triassic of Scotland, Erpetosuchus sp. from the Late Triassic of eastern United States and Parringtonia gracilis from the middle Middle Triassic of Tanzania; the group might also include Dyoplax arenaceus from the Late Triassic of Germany, Archeopelta arborensis and Pagosvenator candelariensis from Brazil and Tarjadia ruthae from Argentina.