Elliot Formation

Last updated
Elliot Formation
Stratigraphic range: Norian-Pliensbachian
~220–190  Ma
Elliot Formation Caves in 'Matalane Valley, Leribe, Lesotho - panoramio.jpg
Elliot Formation caves in the Matalane Valley, Leribe, Lesotho
Type Geological formation
Unit of Stormberg Group
Sub-unitsUpper Elliot, Lower Elliot
Underlies Clarens Formation
Overlies Molteno Formation
Thicknessup to 500 m (1,600 ft)
Lithology
Primary Mudstone, sandstone
Other Siltstone, conglomerate
Location
Coordinates 30°30′S27°24′E / 30.5°S 27.4°E / -30.5; 27.4
Approximate paleocoordinates 44°06′S1°54′W / 44.1°S 1.9°W / -44.1; -1.9
Region Eastern Cape, Free State, Mafeteng, Maseru, Quthing, Qacha's Nek & Mohale's Hoek
CountryFlag of Lesotho.svg  Lesotho
Flag of South Africa.svg  South Africa
Type section
Named for Elliot, Eastern Cape
South Africa relief location map.svg
Blue pog.svg
Elliot Formation (South Africa)

The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, southern Free State, and in the eastern KwaZulu-Natal provinces of South Africa. Outcrops and exposures are also found in several localities in Lesotho such as Qacha's Neck, Hill Top, Quthing, and near the capital, Maseru. The Elliot Formation is further divided into the lower (LEF) and upper (UEF) Elliot formations to differentiate significant sedimentological differences between these layers. The LEF is dominantly Late Triassic (Norian-Hettangian) in age while the UEF is mainly Early Jurassic (Sinemurian-Pliensbachian) and is tentatively regarded to preserve a continental record of the Triassic-Jurassic boundary in southern Africa. [1] This geological formation is named after the town of Elliot in the Eastern Cape, and its stratotype locality is located on the Barkly Pass, 9 km north of the town. [2] [3] [4]

Contents

Geology

The Elliot Formation unconformably overlies the Molteno Formation and is conformably overlain by the Clarens Formation. Due to the reddish colour of the rocks, the Elliot Formation is colloquially referred to as the “Red Beds” in older geologic literature.

The Elliot Formation is dominated by mudstones and siltstones that can be finely laminated. However, the internal structures in the mudstones are often not visible due to locally poor laminations. Calcareous nodules are also found in the mudstone layers and become more frequent up section into the UEF. The mudstones range in colour from greyish purple red in the LEF and turn a more brick red colour with more mature palaeosols in the UEF. Localized intraformational pebble conglomerates that comprise intrabasinal clasts that comprise mud chips, quartzite pebbles, pedogenic nodules, and fossil bone fragments only occur in the UEF. The lower and upper Elliot formations both contain sandstones but they vary in their internal geometries. The sandstones of the LEF mainly comprise laterally accreting channel deposits that are multi-story and contain trough, low angle, and planar, cross-bedding. Ripple cross laminations with good horizontal lamination are also present. In the UEF, sandstone beds are single story and mainly reflect downstream accretion channel geometries and are more tabular in appearance. Common internal sedimentary structures of UEF sandstones are planar, low angle cross-bedding, horizontal and ripple-cross laminations.

The LEF was deposited in a fluvio-lacustrine environment where rivers were more perennial and formed meandering channel geometries, as evidenced by the presence of lateral accretion. However, this depositional environment changed at the onset of the UEF deposits where evidence of shallower river channels, longer periods of floodplain stasis (mature palaeosols) and flash flood events (pedogenic nodule conglomerates) shows that the climate became more arid. [5] [6] [7] [8]

Correlation

The Elliot Formation is currently considered to correlate chronostratigraphically with geological formations of the Bodibeng Sandstone of the Tuli Basin in Botswana, the Omingonde Formation of the Etjo Basin in Namibia, and the Chinle Formation of the Colorado Plateau in Utah, United States. [9] [10] [11] [12] [13] [14]

Paleontology

The Elliot Formation is well known for its diverse dinosaur fossils. The most common dinosaur species is of the sauropodomorph species Massospondylus carinatus . [15] [16] Other species include Blikanasaurus cromptoni , Aardonyx celestae , Euskelosaurus browni , Antetonitrus ingenipes , Pulanesaura eocollum , and the largest sauropodomorph yet found, Ledumahadi mafube . [17] [18] [19] [20] [21] [22] [23] [24] Fossilised Massospondylus eggs, some with the fossilized remains of embryos intact, have been recovered from UEF deposits in the Golden Gate Highlands National Park. [25] [26] Euskelosaurus fossils are more common in the LEF while Massospondylus are only found in the UEF. The basal ornithischian dinosaurs, Heterodontosaurus tucki , Lesothosaurus diagnosticus , Abrictosaurus consors , and Lycorhinus angustidens have also been recovered from the UEF. [27] [28] In addition this formation has yielded various crocodylomorph species, [29] namely Litargosuchus leptorhynchus , Sphenosuchus acutus and Orthosuchus stormbergi . [30] [31] [32] [33] A large theropod dinosaur, Dracovenator regenti , has been found in the UEF. [34] Synapsids from the formation include the dicynodont Pentasaurus goggai [35] the tritheledontid cynodont Elliotherium kersteni [36] [37] and the mammaliaform Megazostrodon rudnerae . [38] More recent vertebrate fossil finds near the town Qhemegha in the Eastern Cape have yielded possible fossil material of a poposauroid pseudosuchian. The mudstones of the LEF sometimes yield petrified wood, fossil plant matter, crustaceans, fishes, and turtles while the sandstones of the upper Elliot Formation more often contain various trace fossils. These include vertebrate trackways of basal ornithischian dinosaurs found in the Leribe, Mafeteng, and Mohales Hoek Districts of Lesotho. Possible trackways of the dicynodont Pentasaurus have been found on Morobong Hill in the Mohales Hoek District of Lesotho. [39] [40] [41] [42]

Dinosaurs

Ornithischians

Ornithischians of the Elliot Formation
GenusSpeciesLocationStratigraphyNotesImages
Abrictosaurus A. consors
  • Upper Elliot
A heterodontosaurid, formerly species of Lycorhinus Abrictosaurus dinosaur.png
Eocursor E. parvus
  • Upper Elliot
A basal ornithischian Eocursor BW.jpg
Fabrosaurus F. australis
  • Upper Elliot
A basal ornithischian, nomen dubium
Heterodontosaurus H. tucki
  • Upper Elliot
A heterodontosaurid Heterodontosaurus restoration.jpg
Lesothosaurus L. diagnosticus
  • Upper Elliot
A basal neornithischian or thyreophoran, formerly species of Fabrosaurus Lesothosaurus diagnosticus.png
Lycorhinus L. angustidens
  • Upper Elliot
A heterodontosaurid
Pegomastax P. africana
  • Upper Elliot
A heterodontosaurid Pegomastax africana reconstruction.jpg

Sauropodomorphs

Sauropodomorphs of the Elliot Formation
GenusSpeciesLocationStratigraphyNotesImages
Aardonyx A. celestae
  • Upper Elliot
A derived sauropodomorph Aardonyx NT.jpg
Antetonitrus A. ingenipes
  • Upper Elliot
A derived sauropodomorph or basal sauropod Antetonitrus reconstruction.jpg
Arcusaurus A. pereirabdalorum
  • Spion Kop Heelbo
  • Upper Elliot
A basal sauropodomorph, known from juveniles
Gryponyx G. africanus
  • Upper Elliot
A massospondylid, potentially dubious
Blikanasaurus B. cromptoni
  • Blikana Mountain
  • Lower Elliot
A derived sauropodomorph or basal sauropod
Eucnemesaurus E. fortis
  • Lower Elliot
A possible riojasaurid
E. entaxonis
Euskelosaurus E. browni
  • Lower Elliot
A plateosaurid, potentially dubious
Ignavusaurus I. rachelis
  • Upper Elliot
A massospondylid, known from a juvenile, potential synonym of Massospondylus
Kholumolumo
  • Lower Elliot
A basal sauropodiform
Ledumahadi L. mafubeA derived sauropodomorph or basal sauropod, largest in the formation Ledumahadi NT.jpg
Massospondylus M. carinatus
  • Upper Elliot
A massospondylid, most common fossil found Massospondylus reconstruction.png
M. kaalae
  • Upper Elliot
A massospondylid, differs from M. carinatus in cranial features
Melanorosaurus M. readi
  • Thaba 'Nyama
  • Lower Elliot
A derived sauropodomorph Melanorosaurus life restoration 2018.jpg
Meroktenos M. thabanensis
  • Thabana Morena
  • Lower Elliot
A derived sauropodomorph, formerly a species of Melanorosaurus
Plateosauravus P. cullingworthi
  • Lower Elliot
A basal sauropodomorph, formerly specimens of Euskelosaurus
Pulanesaura P. eocollum
  • Upper Elliot
A derived sauropodomorph or basal sauropod
Sefapanosaurus S. zastronensis
  • Upper or Lower Elliot
A derived sauropodomorph

Theropods

Theropods of the Elliot Formation
GenusSpeciesLocationStratigraphyNotesImages
Dracovenator D. regenti
  • Upper Drumbo Farm
  • Upper Elliot
A dilophosaurid Dracovenator regenti.jpg
Megapnosaurus Megapnosaurus rhodesiensis
  • Upper Elliot
A coelophysid Coelophysis rhodesiensis.JPG

Suchians

Crocodylomorphs

Crocodylomorphs of the Elliot Formation
GenusSpeciesLocationStratigraphyNotesImages
Basutodon B. ferox
  • Lower Elliot
A dubious suchian
Litargosuchus L. leptorhynchus
  • Eagles Crag Farm
  • Upper Elliot
A basal crocodylomorph Litargosuchus leptorhynchus.jpg
Orthosuchus O. stormbergi
  • Upper Elliot
A possible protosuchid
Rauisuchian
  • Lower Elliot
A indeterminate large "Rauisuchian"
Sphenosuchus S. acutus
  • Upper Elliot
A large basal crocodylomorph

Synapsids

Synapsids of the Elliot Formation
GenusSpeciesLocationStratigraphyNotesImages
Elliotherium E. kersteni
  • Lower Elliot
A tritheledontid cynodont
Megazostrodon M. rudnerae
  • Upper Elliot
A megazostrodontid cynodont Megazostrodon sp. Natural History Museum - London.jpg
Pentasaurus P. goggai
  • Lower Elliot
A stahleckeriid dicynodont
Scalenodontoides S. macrodontes
  • Lower Elliot
A very large traversodontid cynodont

Related Research Articles

<i>Lesothosaurus</i> Extinct genus of ornithischian dinosaur

Lesothosaurus is a monospecific genus of ornithischian dinosaur that lived during the Early Jurassic in what is now South Africa and Lesotho. It was named by paleontologist Peter Galton in 1978, the name meaning "lizard from Lesotho". The genus has only one valid species, Lesothosaurus diagnosticus. Lesothosaurus is one of the most completely-known early ornithischians, based on numerous skull and postcranial fossils from the Upper Elliot Formation. It had a simpler tooth and jaw anatomy than later ornithischians, and may have been omnivorous in some parts of the year.

<i>Melanorosaurus</i> Sauropodomorph dinosaur genus from the Late Triassic period

Melanorosaurus is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. A herbivore from South Africa, it had a large body and sturdy limbs, suggesting it moved about on all fours. Its limb bones were massive and heavy like the limb bones of true sauropods.

<i>Abrictosaurus</i> Extinct genus of dinosaur from the early Jurassic of southern Africa

Abrictosaurus is a genus of heterodontosaurid dinosaur that lived during the Early Jurassic in what is now in parts of southern Africa such as Lesotho and South Africa. It was a bipedal herbivore or omnivore and was one of the most basal heterodontosaurids. It was approximately 1.2 metres (3.9 ft) long and weighed between 0.68 and 3 kilograms.

<i>Massospondylus</i> Sauropodomorph dinosaur genus from Early Jurassic South Africa and Botswana

Massospondylus was a genus of sauropodomorph dinosaur from the Early Jurassic. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. Fossils have since been found at other locations in South Africa, Lesotho, and Zimbabwe. Material from Arizona's Kayenta Formation, India, and Argentina has been assigned to the genus at various times, but the Arizonan and Argentinian material are now assigned to other genera.

<i>Antetonitrus</i> Sauropod dinosaur genus from Early Jurassic South Africa

Antetonitrus is a genus of sauropod dinosaur found in the Early Jurassic Elliot Formation of South Africa. The only species is Antetonitrus ingenipes. As one of the oldest known sauropods, it is crucial for the understanding of the origin and early evolution of this group. It was a quadrupedal herbivore, like all of its later relatives, but shows primitive adaptations to use the forelimbs for grasping, instead of purely for weight support.

<i>Blikanasaurus</i> Extinct genus of dinosaur from the late Triassic of South Africa

Blikanasaurus is a genus of sauropodomorph dinosaur from the late Triassic of South Africa. The generic name Blikanasaurus is derived from Greek, meaning "lizard from Blikana". The species name cromptoni is taken from the surname of A.W. "Fuzz" Crompton, an American paleontologist who led numerous field expeditions in Elliot Formation outcrop localities in South Africa. Blikanasaurus is only known from partial hindlimb bones that were recovered from the lower Elliot Formation (LEF) in the Eastern Cape.

<i>Gryponyx</i> Extinct genus of dinosaur from early Jurassic South Africa

Gryponyx is an extinct genus of massopod sauropodomorph known from southern Free State, central South Africa.

<i>Eocursor</i> Extinct genus of dinosaur from early Jurassic South Africa

Eocursor is genus of basal ornithischian dinosaur that lived in what is now South Africa during the Early Jurassic. Remains of this animal have been found in the Upper Elliot Formation and it is among the most completely known early ornithischians, shedding new light on the origin of the group.

<span class="mw-page-title-main">Clarens Formation</span> Geological formation of the Stormberg Group in southern Africa

The Clarens Formation is a geological formation found in several localities in Lesotho and in the Free State, KwaZulu-Natal, and Eastern Cape provinces in South Africa. It is the uppermost of the three formations found in the Stormberg Group of the greater Karoo Supergroup rocks and represents the final phase of preserved sedimentation of the Karoo Basin.

<span class="mw-page-title-main">Massospondylidae</span> Extinct family of dinosaurs

Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and Antarctica during the Late Triassic to the Early Jurassic periods. Several dinosaurs have been classified as massospondylids over the years. The largest cladistic analysis of early sauropodomorphs, which was presented by Apaldetti and colleagues in November 2011, found Adeopapposaurus, Coloradisaurus, Glacialisaurus, Massospondylus, Leyesaurus and Lufengosaurus to be massospondylids. This result supports many previous analyses that tested fewer taxa. However, this analysis found the two recently described North American massopods, Sarahsaurus and Seitaad, and the South African Ignavusaurus to nest outside Massospondylidae, as opposed to some provisional proposals. Earlier in 2011, Pradhania, a sauropodomorph from India, was tested for the first time in a large cladistic analysis and was found to be a relatively basal massospondylid. Mussaurus and Xixiposaurus may also be included within Massospondylidae.

<span class="mw-page-title-main">Forest Sandstone</span> Geologic formation in Southern Africa

The Forest Sandstone is a geological formation in southern Africa, dating to roughly between 200 and 190 million years ago and covering the Hettangian to Sinemurian stages of the Jurassic Period in the Mesozoic Era. As its name suggests, it consists mainly of sandstone.

Notochampsa is an extinct genus of protosuchian crocodyliform. Fossils have been found from the lower Clarens Formation of the Karoo Supergroup in South Africa, dating back to the Pliensbachian stage of the Early Jurassic. Notochampsa comes from a period of relative fossil scarcity, and is the youngest known occurrence of a crocodylomorph from the Karoo Basin of South Africa.

<span class="mw-page-title-main">Molteno Formation</span> Triassic geological formation in the Stormberg Group in Lesotho and South Africa

The Molteno Formation is a geological formation found in several localities in Lesotho and South Africa. It lies mainly south of Maseru, near Burgersdorp, Aliwal North, Dordrecht, Molteno, and Elliot. It extends as far north as Matatiele in the Eastern Cape. The formation's localities lie along the Drakensberg Mountains in Kwazulu-Natal, and near Ladybrand in the Free State of South Africa. The Molteno Formation is the lowermost of the three formations in the Stormberg Group of the greater Karoo Supergroup. The Molteno Formation represents the initial phase of preserved sedimentation of the Stormberg Group.

Ignavusaurus is a genus of basal sauropodomorph dinosaur that lived during the Early Jurassic in what is now Lesotho. Its fossils were found in the Upper Elliot Formation which is probably Hettangian in age. It was described on the basis of a partial, well preserved articulated skeleton. The type species, I. rachelis, was described in 2010 by Spanish palaeontologist F. Knoll.

<span class="mw-page-title-main">Stormberg Group</span> Triassic/Jurassic geological group in the Karoo Supergroup in South Africa

The Stormberg Group is one of the four geological groups that comprises the Karoo Supergroup in South Africa. It is the uppermost geological group representing the final phase of preserved sedimentation of the Karoo Basin. The Stormberg Group rocks are considered to range between Lower Triassic (Olenekian) to Lower Jurassic (Pliensbachian) in age. These estimates are based on means of geological dating including stratigraphic position, lithostratigraphic and biostratigraphic correlations, and palynological analyses.

<i>Pegomastax</i> Extinct genus of dinosaurs

Pegomastax is a genus of heterodontosaurid dinosaur that lived during the Early Jurassic of South Africa. The only known specimen was discovered in a 1966–1967 expedition in Transkei District of Cape Province, but was not described until 2012 when Paul Sereno named it as the new taxon Pegomastax africana. The genus name is derived from the Greek for "strong jaw", and the species name describes the provenance of Africa; it was originally spelled africanus, was corrected to africana to align with the gender of the genus name.

<i>Pentasaurus</i> Genus of dicynodont therapsid from the late Triassic of South Africa

Pentasaurus is an extinct genus of dicynodont of the family Stahleckeriidae, closely related to the well known Placerias. It was found in the Lower Elliot Formation of South Africa, dated to the Norian of the Late Triassic period. The genus contains the type and only species, Pentasaurus goggai. Pentasaurus is named after the ichnogenus Pentasauropus, fossil footprints that were originally described from the lower Elliot Formation in 1970 decades before the body fossils of Pentasaurus itself were recognised. Pentasauropus footprints were likely made by dicynodonts, and in South Africa Pentasaurus itself was the likely trackmaker. The name reflects the fact that a large dicynodont was predicted to have existed in the lower Elliot Formation before any body fossils were recognised, and so Pentasaurus was named after its probable footprints. This is a reversal of the more typical occurrence where fossil footprints are named after their presumed trackmakers. The name of the species honours its collector Alfred Brown, nicknamed "Gogga", which means "bug" in Afrikaans.

<span class="mw-page-title-main">Lessemsauridae</span> Extinct clade of dinosaurs

Lessemsauridae is a clade of early sauropodiform dinosaurs that lived in the Triassic and Jurassic of Argentina, South Africa and possibly Lesotho. A phylogenetic analysis performed by Apaldetti and colleagues in 2018 recovered a new clade of sauropodiforms uniting Lessemsaurus, Antetonitrus, and Ingentia which they named Lessemsauridae. It is a node-based taxon, defined as all descendants of the most recent common ancestor of Lessemsaurus sauropoides and Antetonitrus ingenipes. Depending on the definition of Sauropoda, Lessemsauridae is either one of the most basal sauropod taxa, or a sister taxon of Sauropoda. An additional member of the clade was named later in 2018, Ledumahadi. A 2021 study by Pol and colleagues also assigned the genera Kholumolumo and Meroktenos to the group.

<i>Ngwevu</i> Sauropodomorph dinosaur genus from Early Jurassic South Africa

Ngwevu is a genus of massospondylid sauropodomorph dinosaur from the Lower Jurassic of South Africa. The genus contains one species, Ngwevu intloko.

References

  1. Bordy, Emese M.; Abrahams, Miengah; Sharman, Glenn R.; Viglietti, Pia A.; Benson, Roger B.J.; McPhee, Blair W.; Barrett, Paul M.; Sciscio, Lara; Condon, Daniel; Mundil, Roland; Rademan, Zandri (April 2020). "A chronostratigraphic framework for the upper Stormberg Group: Implications for the Triassic-Jurassic boundary in southern Africa". Earth-Science Reviews. 203: 103120. doi:10.1016/j.earscirev.2020.103120. ISSN   0012-8252. S2CID   213646670.
  2. Bordy, Emese M.; John Hancox, P.; Rubidge, Bruce S. (2004-03-01). "Fluvial style variations in the Late Triassic–Early Jurassic Elliot formation, main Karoo Basin, South Africa". Journal of African Earth Sciences. 38 (4): 383–400. Bibcode:2004JAfES..38..383B. doi:10.1016/j.jafrearsci.2004.02.004. ISSN   1464-343X.
  3. "A description of the sedimentology and palaeontology of the Late Triassic–Early Jurassic Elliot Formation in Lesotho". WIReDSpace Wits. 2004-09-01. ISSN   0078-8554.
  4. Bordy, E.M.; Eriksson, P. (2015-09-01). "Lithostratigraphy of the Elliot Formation (Karoo Supergroup), South Africa". South African Journal of Geology. 118 (3): 311–316. doi:10.2113/gssajg.118.3.311. hdl: 2263/55739 . ISSN   1012-0750.
  5. Tankard, Anthony; Welsink, Herman; Aukes, Peter; Newton, Robert; Stettler, Edgar (2009-09-01). "Tectonic evolution of the Cape and Karoo basins of South Africa". Marine and Petroleum Geology. 26 (8): 1379–1412. doi:10.1016/j.marpetgeo.2009.01.022. ISSN   0264-8172.
  6. "Meander channel, point bar, crevasse splay and aeolian deposits from the Elliot Formation in Barkly Pass, North-eastern Cape". South African Journal of Geology. 83 (1). 1980-01-01. ISSN   1012-0750.
  7. Bordy, E.M.; Eriksson, P. (2015-09-01). "Lithostratigraphy of the Elliot Formation (Karoo Supergroup), South Africa". South African Journal of Geology. 118 (3): 311–316. doi:10.2113/gssajg.118.3.311. hdl: 2263/55739 . ISSN   1012-0750.
  8. Sciscio, Lara; De Kock, Michiel; Bordy, Emese; Knoll, Fabien (2017-11-01). "Magnetostratigraphy across the Triassic-Jurassic boundary in the main Karoo Basin". Gondwana Research. 51: 177–192. Bibcode:2017GondR..51..177S. doi:10.1016/j.gr.2017.07.009. hdl: 10261/338607 . ISSN   1342-937X.
  9. Attridge, J.; Crompton, A. W.; Jenkins, Farish A. (June 1985). "The Southern African Liassic prosauropodMassospondylusdiscovered in North America". Journal of Vertebrate Paleontology. 5 (2): 128–132. doi:10.1080/02724634.1985.10011850. ISSN   0272-4634.
  10. Eriksson, P.; Bordy, E. M. (September 2015). "Lithostratigraphy of the Elliot Formation (Karoo Supergroup), South Africa". South African Journal of Geology. 118 (3): 311–316. doi:10.2113/gssajg.118.3.311. hdl: 2263/55739 . ISSN   1012-0750.
  11. Smith, R. M.H.; Swart, R. (2002-06-01). "Changing Fluvial Environments and Vertebrate Taphonomy in Response to Climatic Drying in a Mid-Triassic Rift Valley Fill: The Omingonde Formation (Karoo Supergroup) of Central Namibia". PALAIOS. 17 (3): 249–267. Bibcode:2002Palai..17..249S. doi:10.1669/0883-1351(2002)017<0249:CFEAVT>2.0.CO;2. ISSN   0883-1351. S2CID   130316395.
  12. Abdala, Fernando; Smith, Roger M. H. (2009-09-12). "A Middle Triassic cynodont fauna from Namibia and its implications for the biogeography of Gondwana". Journal of Vertebrate Paleontology. 29 (3): 837–851. doi:10.1671/039.029.0303. ISSN   0272-4634. S2CID   129096876.
  13. Walker, A. D. (1990-10-29). "A revision of Sphenosuchus acutus Haughton, a crocodylomorph reptile from the Elliot Formation (late Triassic or early Jurassic) of South Africa". Phil. Trans. R. Soc. Lond. B. 330 (1256): 1–120. doi:10.1098/rstb.1990.0185. ISSN   0962-8436.
  14. Tucker, Maurice E.; Benton, Michael J. (1982-12-01). "Triassic environments, climates and reptile evolution". Palaeogeography, Palaeoclimatology, Palaeoecology. 40 (4): 361–379. Bibcode:1982PPP....40..361T. doi: 10.1016/0031-0182(82)90034-7 . ISSN   0031-0182.
  15. Sues, H.D., Reisz, R.R., Hinic, S. and Raath, M.A., 2004. On the skull of Massospondylus carinatus Owen, 1854 (Dinosauria: Sauropodomorpha) from the Elliot and Clarens formations (Lower Jurassic) of South Africa. Annals of Carnegie Museum, 73(4), pp.239–257.
  16. Chinsamy, Anusuya. "Bone histology and growth trajectory of the prosauropod dinosaur Massospondylus carinatus Owen | Request PDF". ResearchGate. Retrieved 2018-11-14.
  17. Galton, Peter M.; Heerden, Jacques (April 1998). "Anatomy of the prosauropod dinosaurBlikanasaurus cromptoni (Upper Triassic, South Africa), with notes on the other tetrapods from the lower Elliot Formation". Paläontologische Zeitschrift. 72 (1–2): 163–177. doi:10.1007/bf02987824. ISSN   0031-0220. S2CID   128464155.
  18. Yates, Adam M. (2003). "A definite prosauropod dinosaur from the Lower Elliot Formation (Norian: Upper Triassic) of South Africa". Palaeontologia Africana. ISSN   0078-8554.
  19. Barrett, Paul M. (2004-09-01). "Sauropodomorph dinosaur diversity in the upper Elliot Formation (Massospondylus range zone: Lower Jurassic) of South Africa: research letter". South African Journal of Science. 100 (9–10). ISSN   0038-2353.
  20. McPhee, Blair; Bordy, Emese; Sciscio, Lara; Choiniere, Jonah (2017). "The sauropodomorph biostratigraphy of the Elliot Formation of southern Africa: Tracking the evolution of Sauropodomorpha across the Triassic–Jurassic boundary". Acta Palaeontologica Polonica. 62 (3). doi: 10.4202/app.00377.2017 . ISSN   0567-7920.
  21. Hancox, P. John; Rubidge, Bruce S.; Yates, Adam M. (1 September 2004). "First record of a sauropod dinosaur from the upper Elliot Formation (Early Jurassic) of South Africa: research letter". South African Journal of Science. 100 (9): 504–506.
  22. McPhee, Blair W.; Yates, Adam M.; Choiniere, Jonah N.; Abdala, Fernando (2014-04-25). "The complete anatomy and phylogenetic relationships of Antetonitrus ingenipes (Sauropodiformes, Dinosauria): implications for the origins of Sauropoda". Zoological Journal of the Linnean Society. 171 (1). doi: 10.1111/zoj12127 . ISSN   1096-3642.
  23. McPhee, B. W.; Benson RBJ; Botha-Brink, J.; Bordy, E. M.; Choiniere, J. N. (2018-10-08). "A Giant Dinosaur from the Earliest Jurassic of South Africa and the Transition to Quadrupedality in Early Sauropodomorphs". Current Biology. 28 (19): 3143–3151.e7. doi: 10.1016/j.cub.2018.07.063 . ISSN   0960-9822. PMID   30270189.
  24. Sander, P. M.; Lallensack, J. N. (2018-10-08). "Dinosaurs: Four Legs Good, Two Legs Bad". Current Biology. 28 (19): R1160–R1163. doi: 10.1016/j.cub.2018.08.025 . ISSN   0960-9822. PMID   30300605.
  25. Kitching, James (1979). "Preliminary report on a clutch of six dinosaurian eggs from the Upper Triassic Elliot Formation, Northern Orange Free State". Palaeontologia Africana. 22: 41–45.
  26. Reisz, Robert R.; Evans, David C.; Sues, Hans-Dieter; Scott, Diane (2010-12-02). "Embryonic skeletal anatomy of the sauropodomorph dinosaurMassospondylusfrom the Lower Jurassic of South Africa". Journal of Vertebrate Paleontology. 30 (6): 1653–1665. doi:10.1080/02724634.2010.521604. ISSN   0272-4634. S2CID   84599991.
  27. Knoll, Fabien (2002-07-08). "Nearly complete skull ofLesothosaurus(Dinosauria: Ornithischia) from the Upper Elliot Formation (Lower Jurassic: Hettangian) of Lesotho". Journal of Vertebrate Paleontology. 22 (2): 238–243. doi:10.1671/0272-4634(2002)022[0238:ncsold]2.0.co;2. ISSN   0272-4634. S2CID   85790601.
  28. Butler, Richard J. (October 2005). "The 'fabrosaurid' ornithischian dinosaurs of the Upper Elliot Formation (Lower Jurassic) of South Africa and Lesotho". Zoological Journal of the Linnean Society. 145 (2): 175–218. doi: 10.1111/j.1096-3642.2005.00182.x . ISSN   1096-3642.
  29. Clark, James M.; Sues, Hans-Dieter (2002-08-21). "Two new basal crocodylomorph archosaurs from the Lower Jurassic and the monophyly of the Sphenosuchia". Zoological Journal of the Linnean Society. 136 (1): 77–95. doi: 10.1046/j.1096-3642.2002.00026.x . ISSN   1096-3642.
  30. Walker, A. D. (1990-10-29). "A revision of Sphenosuchus acutus Haughton, a crocodylomorph reptile from the Elliot Formation (late Triassic or early Jurassic) of South Africa". Phil. Trans. R. Soc. Lond. B. 330 (1256): 1–120. doi:10.1098/rstb.1990.0185. ISSN   0962-8436.
  31. Nash, Diane (2009-08-20). "A crocodile from the Upper Triassic of Lesotho". Journal of Zoology. 156 (2): 163–179. doi:10.1111/j.1469-7998.1968.tb05927.x. ISSN   0952-8369.
  32. Nash, D.S., 1975. The morphology and relationships of a crocodilian, Orthosuchus stormbergi, from the Upper Triassic of Lesotho. South African Museum.
  33. Dollman, K. N.; Viglietti, P. A.; Choiniere, J. N. (2017-10-12). "A new specimen of Orthosuchus stormbergi (Nash 1968) and a review of the distribution of Southern African Lower Jurassic crocodylomorphs". Historical Biology. 31 (5): 653–664. doi:10.1080/08912963.2017.1387110. ISSN   0891-2963. S2CID   134134524.
  34. "A new theropod dinosaur from the Early Jurassic of South Africa and its implication for the early evolution of theropods | Request PDF". ResearchGate. Retrieved 2018-11-16.
  35. F., Kammerer, Christian (2018). "The first skeletal evidence of a dicynodont from the lower Elliot Formation of South Africa". Palaeontologia Africana. ISSN   2410-4418.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  36. Hancox, P. J.; Sidor, C. A. (March 2006). "Elliotherium Kersteni, A New Tritheledontid from the Lower Elliot Formation (Upper Triassic) of South Africa". Journal of Paleontology. 80 (2): 333–342. CiteSeerX   10.1.1.557.9156 . doi:10.1666/0022-3360(2006)080[0333:EKANTF]2.0.CO;2. ISSN   0022-3360. S2CID   130003909.
  37. Fernando, Abdala; Ross, Damiani; Adam, Yates; Johann, Neveling (2007). "A non-mammaliaform cynodont from the Upper Triassic of South Africa: a therapsid Lazarus taxon?". Palaeontologia Africana. ISSN   0078-8554.
  38. E, Gow, Chris (1986-06-30). "A new skull of Megazostrodon (Mammalia, Triconodonta) from the Elliot Formation (Lower Jurassic) of Southern Africa". Palaeontologia Africana. ISSN   0078-8554.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  39. Anderson, J.M.; Anderson, H.M.; Cruickshank, A.R.I. (1998). "Late Triassic ecosystems of the Molteno Lower Elliot biome of southern Africa" (PDF). Palaeontology. 41 (3): 387–421.
  40. Smith, R.; Kitching, J. (1997-06-01). "Sedimentology and vertebrate taphonomy of the Tritylodon Acme Zone: a reworked palaeosol in the Lower Jurassic Elliot Formation, Karoo Supergroup, South Africa". Palaeogeography, Palaeoclimatology, Palaeoecology. 131 (1–2): 29–50. Bibcode:1997PPP...131...29S. doi:10.1016/S0031-0182(96)00143-5. ISSN   0031-0182.
  41. Knoll, Fabien (January 2005). "The tetrapod fauna of the Upper Elliot and Clarens formations in the main Karoo Basin (South Africa and Lesotho)". Bulletin de la Société Géologique de France. 176 (1): 81–91. doi:10.2113/176.1.81. ISSN   0037-9409.
  42. Sciscio, Lara (December 2015). Position of the Triassic-Jurassic boundary in South Africa and Lesotho: a multidisciplinary approach aimed at improving the chronostratigraphy and biostratigraphy of the Elliot Formation, Stormberg Group (Thesis). University of Cape Town via OpenUCT.