This article relies largely or entirely on a single source .(September 2020) |
Australochelys Temporal range: Early Jurassic, | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Rhaptochelydia |
Family: | † Australochelyidae |
Genus: | † Australochelys Gaffney & Kitching, 1994 |
Type species | |
Australochelys africanus Gaffney & Kitching, 1994 |
Australochelys is an extinct genus of rhaptochelydian turtle. It is known from one species, A. africanus, that came from the Elliot Formation of South Africa. The holotype of Australochelys consists of only a skull and a fragment of the carapace, which shows both primitive and derived features. Like Proganochelys , Australochelys has large orbits and a ventral basioccipital tubercle, but like derived turtles such as casichelydians, a group containing Cryptodira and Pleurodira, it possesses a sutured basipterygoidal attachment, and a middle ear region partially enclosed laterally. [1] These characteristics show that Australochelys is more closely related to casichelydians than to Proganochelys, and together with the former, it makes up Rhaptochelydia. The skull of Australochelys shows that an advanced hearing mechanism of turtles evolved before the appearance of modern turtles. [1]
Australochelys africanus was named in 1994 by Eugene S. Gaffney and James W. Kitching. The generic name comes from the Greek australos, meaning "south" and chelys, "turtle". The specific name is derived from the Greek africanus, meaning "from Africa". [1]
The holotype of Australochelys includes a skull and a fragment of the carapace. The skull shares features with both the primitive turtle Proganochelys and Casichelydia, a group of derived turtles. The skull shows an advanced hearing mechanism, a feature thought to have evolved with modern turtles. [1]
Many features were found among Australochelys that function to distinguish it from other turtles. Below are the characteristics found by Gaffney and Kitching in 1995 that are unique among all turtles: [1]
Below are features that are only found in turtles between Australochelys and Proganochelys, but also present in tetrapods: [1]
Below are characteristics found in Australochelys and Casichelydia but not in Proganochelys: [1]
Australochelys shows a strange mix of basal and derived characteristics. This led Gaffney and Kitching to classify it in two new groups, Rhaptochelydia and Australochelyidae. Rhaptochelydia includes Australochelyidae and Casichelydia, a group including the derived groups Cryptodira and Pleurodira. [1]
Below is a simple cladogram found by Gaffney and Kitching in 1995 demonstrating the relationships of Australochelys: [1]
Testudines |
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Australochelys is from the Early Jurassic Elliot Formation, a formation that dates around 190 to 180 million years ago. This makes it the oldest african turtle, a record that was previously held by a fragmentary specimen from the Gokwe Formation, which was dated to the Late Jurassic. Other than the fragmentary possible remains, the oldest definite specimens date to the Early Cretaceous of Africa, which shows about a 60 million year gap between themselves and Australochelys. The holotype of Australochelys was found in a Tritylodon horizon, in the middle section of the formation. [1]
Podocnemididae is a family of pleurodire (side-necked) turtles, once widely distributed. Most of its 20 genera and 30 species are now extinct. Seven of its eight surviving species are native to South America: the genus Peltocephalus, with only one species ; and the genus Podocnemis, with six living species of South American side-necked river turtles. There is also one genus native to Madagascar: Erymnochelys, the Madagascan big-headed turtle, whose single species E. madagascariensis.
Proganochelys is an extinct, primitive stem-turtle that has been hypothesized to be the sister taxon to all other turtles creating a monophyletic group, the Casichelydia. Proganochelys was named by Georg Baur in 1887 as the oldest turtle in existence at the time. The name Proganochelys comes from the Greek word ganos meaning brightness, combined with prefix pro, and Greek base chelys meaning "turtle". Proganochelys is believed to have been around 1 meter in size and herbivorous in nature. Proganochelys was known as the most primitive stem-turtle for over a century, until the novel discovery of Odontochelys in 2008. Odontochelys and Proganochelys share unique primitive features that are not found in Casichelydia, such as teeth on the pterygoid and vomer and a plate-like coracoid.
Elginia is an extinct genus of pareiasaurid known from the Late Permian of Scotland and China. It was named for the area around Elgin in Scotland, which has yielded many fossils referred to as the Elgin Reptiles.
The Pleurodira are one of the two living suborders of turtles, the other being the Cryptodira. The division between these two suborders represents a very deep evolutionary divide between two very different types of turtles. The physical differences between them, although anatomical and largely internal, are nonetheless significant, and the zoogeographic implications of them are substantial. The Pleurodira are known more commonly as the side-necked turtles and the name Pleurodira quite literally translates to side neck, whereas the Cryptodira are known as hidden-necked turtles. The Pleurodira turtles are currently restricted to freshwater habitats in the Southern Hemisphere, largely to Australia, South America, and Africa. Within the Pleurodira, three living families are represented: Chelidae, also known as the Austro-South American side-necked turtles, the Pelomedusidae, also known as the African mud terrapins, and the Podocnemididae, also known as the American side-neck river turtles. However, they are a cosmopolitan clade across the Cretaceous and Cenozoic, and even occurred in marine environments around the world.
Metoposaurus meaning "front lizard" is an extinct genus of stereospondyl temnospondyl amphibian, known from the Late Triassic of Germany, Italy, Poland, and Portugal. This mostly aquatic animal possessed small, weak limbs, sharp teeth, and a large, flat head. This highly flattened creature mainly fed on fish, which it captured with its wide jaws lined with needle-like teeth. Metoposaurus was up to 3 m long and weighed about 450 kg. Many Metoposaurus mass graves have been found, probably from creatures that grouped together in drying pools during drought.
Nicrosaurus (/nɛkroʊˈsɔrəs/) is an extinct genus of phytosaur reptile existing during the Late Triassic period. Although it looked like a crocodile, it was not closely related to these creatures, instead being an example of parallel evolution. The main difference between Nicrosaurus and modern crocodiles is the position of the nostrils – Nicrosaurus's nostrils, or external nares, were placed directly in front of the forehead, whereas in crocodiles, the nostrils are positioned on the end of the snout. A 2013 study has also found that illium of Nicrosaurus is quite distinctive from all other phytosaurs.
Dracovenator is a genus of neotheropod dinosaur that lived approximately 201 to 199 million years ago during the early part of the Jurassic Period in what is now South Africa. Dracovenator was a medium-sized, moderately-built, ground-dwelling, bipedal carnivore, that could grow up to an estimated 7 m (23.0 ft) long. Its type specimen was based on only a partial skull that was recovered.
Paracryptodira is an extinct group of reptiles in the clade Testudinata, known from the Jurassic to Paleogene of North America and Europe. Initially treated as a suborder sister to Cryptodira, they were then thought to be a very primitive lineage inside the Cryptodira according to the most common use of the latter taxon. They are now often regarded as late-diverging stem-turtles, lying outside the clade formed by Cryptodira and Pleurodira. The paracryptodires are said to have phylogenic relationships, noted as primary subclades, within the Baenidae and Pleurosternidae. Within each subclade, lies many biodiverse turtles that are continuously being investigated and added to the fossil record. Paracryptodires are divided into 3 main groups, Compsemydidae, known from the Late Jurassic to Paleocene of North America and Europe, Pleurosternidae, known from the Late Jurassic to Early Cretaceous of North America and Europe, and Baenidae, known from the Early Cretaceous to Eocene of North America. The latter two groups are more closely related to each other than to Compsemys, forming the clade Baenoidea.
Leptopleuron is an extinct genus of procolophonid that lived in the dry lands during the late Triassic in Elgin of northern Scotland and was the first to be included in the clade of Procolophonidae. First described by English paleontologist and biologist Sir Richard Owen, Leptopleuron is derived from two Greek bases, leptos for "slender" and pleuron for "rib," describing it as having slender ribs. The fossil is also known by a second name, Telerpeton, which is derived from the Greek bases tele for "far off" and herpeton for "reptile." In Scotland, Leptopleuron was found specifically in the Lossiemouth Sandstone Formation. The yellow sandstone it was located in was poorly lithified with wind coming from the southwest. The environment is also described to consist of barchan dunes due to the winds, ranging up to 20 m tall that spread during dry phases into flood plains. Procolophonoids such as Leptopleuron were considered an essential addition to the terrestrial ecosystem during the Triassic.
Labidosaurikos is a genus of extinct captorhinid anapsid reptile that lived around 279 to 272 million years ago during Kungurian age of the lower Permian. The American Paleontologist John Willis Stovall first described Labidosaurikos in 1950, naming it "Labidosaurus like" for the striking similarity of the holotype skull of his specimen to the cranial anatomy of another captorhinid Labidosaurus hamatus. Labidosaurus or generally called "lipped lizard" is another genus of the family Captorhinidae whose name is derived from the Greek "forceps lizard" based on τσιμπίδα and σαυρος ("lizard")
Eileanchelys is an extinct genus of primitive turtle from the Middle Jurassic (Bathonian) period some 164 million years ago of Britain. Only one species is recorded, Eileanchelys waldmani. It is the best-represented turtle from the Middle Jurassic, because of the amount of specimens that can be assigned to it. The turtle is also one of the oldest turtles ever found to be aquatic, and might represent a milestone in turtle evolution.
Biseridens is an extinct genus of anomodont therapsid, and one of the most basal anomodont genera known. Originally known from a partial skull misidentified as an 'eotitanosuchian' in 1997, another well-preserved skull was found in the Xidagou Formation, an outcropping in the Qilian Mountains of Gansu, China, in 2009 that clarified its relationships to anomodonts, such as the dicynodonts.
Progalesaurus is an extinct genus of galesaurid cynodont from the early Triassic. Progalesaurus is known from a single fossil of the species Progalesaurus lootsbergensis, found in the Lystrosaurus Assemblage Zone of the Balfour Formation. Close relatives of Progalesaurus, other galesaurids, include Galesaurus and Cynosaurus. Galesaurids appeared just before the Permian-Triassic extinction event, and disappeared from the fossil record in the Middle-Triassic.
Kayentachelys is an extinct genus of turtle known only from the "silty facies" of the Lower Jurassic Kayenta Formation in northeastern Arizona on the lands of the Navajo Nation.
Jesairosaurus is an extinct genus of early archosauromorph reptile known from the Illizi Province of Algeria. It is known from a single species, Jesairosaurus lehmani. Although a potential relative of the long-necked tanystropheids, this lightly-built reptile could instead be characterized by its relatively short neck as well as various skull features.
Nebulasaurus is an extinct genus of basal eusauropod dinosaur known from the early Middle Jurassic Zhanghe Formation of Yunnan Province, China. It is known only from the holotype braincase LDRC-v.d.1. A phylogenetic analysis found Nebulasaurus to be a sister taxon to Spinophorosaurus from the Middle Jurassic of Africa. This discovery is significant paleontologically because it represents a clade of basal eusauropods previously unknown from Asia.
Khunnuchelys was a genus of trionychine turtle from the Late Cretaceous of Asia. Three species are known, K. erinhotensis, the type species, K. kizylkumensis, and K. lophorhothon. K. erinhotensis is known from the Iren Dabasu Formation in China from the late Turonian until the middle Campanian. K. kizylkumensis is known from the late Turonian Bissekty Formation of Uzbekistan. The third species, described in 2013 by Danilov et al., is known from the early to middle Campanian aged Bostobe Formation of Kazakhstan.
Carnufex is an extinct genus of crocodylomorph suchian from the Late Triassic of North America. The genus was first described in 2015 by Zanno et al., who named the binomial Carnufex carolinensis, meaning "Carolina butcher". Two specimens are known, the holotype skull and skeleton NCSM 21558, and the referred humerus NCSM 21623. The specimens are from the Carnian-age Pekin Formation, which dates to 231 million years ago. Based on the holotype, Carnufex would have been about 3 m (9.8 ft) long and 1.5 m (4.9 ft) tall, although it may have gotten larger due to the holotype not being fully grown.
Mesochelydia is a clade within Pantestudines, more inclusive than Perichelydia, but less than Testudinata. The clade is known from the Early Jurassic to the Present, and contains all Jurassic representatives of Testudinata aside from Australochelys. The ancestral condition for Mesochelydia is thought to be aquatic, as opposed to terrestrial for Testudinata. They are distinguished from more basal testudinatans by the presence of the following characters: strap like pectoral girdle, supramarginals absent, reduced posterior entoplastral process, eleven pairs of peripherals, elongate processus interfenestralis, paired basioccipital tubercles, fully formed cavum tympani and antrum postoticum, single vomer, confluent external nares, lacrimals and supratemporals absent.
Chinatichampsus is an extinct genus of crocodilian from the Devil's Graveyard Formation of Texas, specifically the Dalquest Desert Research Site. It is a monotypic genus, containing only the type species Chintanichampsus wilsonorum. A single specimen, TMM 45911–1, was first discovered in 2010. Chinatichampsus is the most basal Eocene caimanine, dating to between 42.8 and 41.5 million years ago, and is considered to be more basal than Protocaiman.