Sandownidae

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Sandownidae
Temporal range: Barremian–Paleocene
Sandownia.jpg
Skull of Sandownia harrisi
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pantestudines
Clade: Testudinata
Clade: Perichelydia
Clade: Thalassochelydia
Family: Sandownidae
Tong & Meylan, 2013
Genera [1]

Sandownidae is a family of extinct marine turtles from the Cretaceous and Paleogene distributed around the Atlantic Ocean and adjacent areas. The family is defined as all taxa closer to the type genus Sandownia than to Pelomedusa , Testudo , Solnhofia , Eurysternum , Plesiochelys , Thalassemys or Protostega , a definition that encompasses the previous concept of the clade while also excluding it from being synonymous with other clades of modern or extinct marine turtles. Sandownidae may be within the larger clade Angolachelonia, defined as inclusive of Angolachelys and Solnhofia, sister to the entirely Late Jurassic marine group Thalassochelydia, although the concepts of the clades may shift with further phylogenetic analysis. [1]

Contents

Genera

Angolachelys mbaxi Angolachelys mbaxi cropped.jpg
Angolachelys mbaxi

Phylogeny

After Evers and Benson, 2018:

Testudinata

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Angolachelonia is a clade of extinct turtles from the Late Jurassic to Paleogene of Eurasia. The group is defined as all taxa derived from the ancestor of the type genus Angolachelys and Solnhofia, a definition that could potentially encompass a clade of entirely marine turtles. Angolachelonia was originally inclusive of only Solnhofia, Angolachelys and Sandownia when originally conceived by Octavio Mateus and colleagues in 2009, but later phylogenetic analyses by Serjocha Evers and Roger Benson in 2018 unites the family Sandownidae, including Angolachelys and Sandownia among other taxa, with the entirely Late Jurassic clade Thalassochelydia, where Solnhofia may be a basal member. While the placement of Solnhofia is weak and the clade that Angolachelonia represents may change with further analysis, the clade of Sandownidae and Thalassochelydia is well-supported, and does not collapse despite the uncertain evolutionary history of the group. Three alternative potential origins of Angolachelonia sensu Evers and Benson are shown below.

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Compsemydidae is an extinct family of turtles, likely belonging to the clade Paracryptodira. The earliest undisputed member is Tongemys from the Berriasian age of the Early Cretaceous; two Late Jurassic genera have also sometimes been included in the group, but may alternatively be members of the family Pleurosternidae. The genus Compsemys survived the Cretaceous–Paleogene extinction event and lasted until the Thanetian age of the Paleocene.

<span class="mw-page-title-main">Platychelyidae</span> Extinct family of turtles

Platychelyidae is an extinct family of pan-pleurodiran turtles, known from the Late Jurassic and Early Cretaceous of Europe, South America, North America, and the Caribbean. It represents the oldest known clade of stem-pleurodires. All known members have been found in marine or coastal deposits. Despite this, their limb morphology suggests that they were not adapted for open marine conditions, but were likely inhabitants of shallow water environments, including brackish and saline waters, and they likely never inhabited environments more marine than lagoons. Their tolerance for saline environments likely aided their dispersal during the breakup of Pangea. Unlike modern pleurodires, which retract their necks to the sides, Platychelys retracted its neck inwards, similar to modern cryptodire turtles. Platychelys is strongly morphologically similar to mata mata and snapping turtles, suggesting that it had a similar ecology as a ram or suction feeder.

References

  1. 1 2 Evers, S.W.; Benson, R.B.J. (2018). "A new phylogenetic hypothesis of turtles with implications for the timing and number of evolutionary transitions to marine lifestyles in the group". Palaeontology. 62 (2): 93–134. doi:10.1111/pala.12384. S2CID   134736808.