An anapsid is an amniote whose skull lacks one or more skull openings near the temples. Traditionally, the Anapsida are considered the most primitive subclass of amniotes, the ancestral stock from which Synapsida and Diapsida evolved, making anapsids paraphyletic. It is, however, doubtful that all anapsids lack temporal fenestra as a primitive trait, and that all the groups traditionally seen as anapsids truly lacked fenestra.
Carettochelyidae is a family of cryptodiran turtles belonging to the Trionychia. It contains only a single living species, the pig-nosed turtle native to New Guinea and Northern Australia. Stem-group carettochelyids are known from the Cretaceous of Asia, with the family being widely distributed across North America, Europe, Asia and Africa during much of the Cenozoic.
The Cryptodira are a suborder of Testudines that includes most living tortoises and turtles. Cryptodira differ from Pleurodira in that they lower their necks and pull the heads straight back into the shells, instead of folding their necks sideways along the body under the shells' marginals. They include among their species freshwater turtles, snapping turtles, tortoises, softshell turtles, and sea turtles.
Paracryptodira is an extinct group of reptiles in the clade Testudinata, known from the Jurassic to Paleogene of North America and Europe. Initially treated as a suborder sister to Cryptodira, they were then thought to be a very primitive lineage inside the Cryptodira according to the most common use of the latter taxon. They are now often regarded as late-diverging stem-turtles, lying outside the clade formed by Cryptodira and Pleurodira. The paracryptodires are said to have phylogenic relationships, noted as primary subclades, within the Baenidae and Pleurosternidae. Within each subclade, lies many biodiverse turtles that are continuously being investigated and added to the fossil record. Paracryptodires are divided into three main groups, Compsemydidae, known from the Late Jurassic to Paleocene of North America and Europe, Pleurosternidae, known from the Late Jurassic to Early Cretaceous of North America and Europe, and Baenidae, known from the Early Cretaceous to Eocene of North America. The latter two groups are more closely related to each other than to Compsemys, forming the clade Baenoidea.
Thalassemys is a genus of extinct thalassochelydian turtle from the Late Jurassic of western and central Europe. While the genus was originally named by Rütimeyer in 1859 for a large carapace and other associated fragments from the late Kimmeridgian of the Reuchenette Formation of Switzerland, although the taxon was not validly named until 1873 when Rütimeyer designated the type species T. hugii. Rütimeyer also named T. gresslyi from the Reunchenette Formation in the same paper as T. hugii, but it cannot be differentiated from the type material of T. hugii and is therefore a junior synonym. A large assemblage of shell and postcranial material from the Reunchenette was named as a species of Eurysternum, E. ignoratum, by Bräm in 1965. While originally distinguished based on the presence of fontanelles on the plastron, the feature was later identified on T. hugii and E. ignoratum was designated a junior synonym. Additional material from the Kimmeridge Clay of the United Kingdom has also been referred to T. hugii.
Testudinata is the group of all tetrapods with a true turtle shell. It includes both modern turtles (Testudines) and many of their extinct, shelled relatives (stem-turtles), though excluding Odontochelys and Eorhynchochelys, which are placed in the more inclusive Pantestudines.
The Reuchenette Formation is a Jurassic geologic formation in Switzerland. It is Kimmeridgian in age and predominantly consists of well stratified limestone, with lithology variable both laterally and stratigraphically including wackestones, packstones and grainstones, as well as mudstone. Dinosaur remains are among the fossils that have been recovered from the formation, including the Turiasaurian sauropod Amanzia greppini, alongside a theropod tooth belonging to Ceratosauria indet, originally assigned to Megalosaurus meriani. teleosaurid crocodyliformes are also known, including Sericodon, Proexochokefalos and Machimosaurus. The metriorhynchid Thalatosuchian Torvoneustes. The hybodontid shark Asteracanthus. The thalassochelydian turtle Thalassemys and Solnhofia is known from the formation, as is the platychelyid turtle Platychelys, and the plesiochelyid turtle Plesiochelys.
Eurysternum is an extinct genus of thalassochelydian turtle. Its type species is Eurysternum wagleri, the holotype of which has since been lost and only survives in illustrations.
Pleurosternon is an extinct genus of freshwater pleurosternid turtle from the latest Jurassic to earliest Cretaceous of Europe. Its type species, P. bullockii was described by the paleontologist Richard Owen in 1853. Since then, and throughout the late 19th century, many fossil turtles were incorrectly assigned to this genus, though only two are currently considered valid.
Palaeomedusa testa is an extinct species of thalassochelydian turtle from the Tithonian of the Late Jurassic. It was first described by the German palaeontologist Christian Erich Hermann von Meyer in 1860. It is the only species classified under the genus Palaeomedusa.
Eurysternidae is an extinct family of turtles in the clade Thalassochelydia. It consists of several genera of marine turtles from marine deposits in Europe, including Achelonia, Chelonides, Eurysternum. Hydropelta, Chelonides, Idiochelys, Palaeomedusa, Parachelys. and Solnhofia.
Hydropelta is a genus of Late Jurassic turtle from marine deposits in the Jura Mountains of eastern France.
Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including Eurysternum, Plesiochelys and Thalassemys to the exclusion of Pelomedusa, Testudo and Protostega. While a clade uniting the families Eurysternidae, Plesiochelyidae and Thalassemydidae had been supported by phylogenetic evidence, a name was not given for the clade until 2017, when Jérémy Anquetin and colleagues coined Thalassochelydia.
Solnhofia is a genus of extinct thalassochelydian turtle from the Late Jurassic of Germany. The type species is Solnhofia parsonsi, named by Gaffney in 1975 for a partial skull and jaw from the early Tithonian of the Solnhofen Formation in Bavaria. Additional material including a complete skeleton is known from the late Kimmeridgian of Switzerland and the Kimmeridgian/Tithonian of other deposits within Bavaria, and potentially also unprepared material from the Late Jurassic of France. The genus was referred to the family Eurysternidae by Anquetin and colleagues in 2017, which may represent an artificial grade of early thalassochelydians. In 2020 a new species Solnhofia brachyrhyncha was described from the Kimmeridigan aged Reuchenette Formation of Switzerland.
Aplax is a dubious genus of extinct thalassochelydian turtle from the Late Jurassic of Germany. The type and only species is Aplax oberndorferi, named by Hermann von Meyer in 1843 for a complete juvenile skeleton from the early Tithonian of the Solnhofen Formation in Bavaria. Despite being aware that shell morphology changes during growth, Meyer named Aplax due to his consideration it represented a relative of Dermochelys, where the adults lack distinction of shell regions as in Aplax. However the taxon was later referred to Thalassochelydia by Anquetin and colleagues in 2017, and due to the loss of the original holotype it cannot be identified as a distinct taxon of a juvenile of existing Solnhofen turtles and is therefore a nomen dubium.
Euryaspis is a dubious genus of extinct thalassochelydian turtle from the Late Jurassic of Germany. The type and only species is Euryaspis radians, originally proposed by Wagner in 1859 before being validly described and illustrated in 1861. The only specimen was a partial carapace probably from the Tithonian, although the original locality is unknown. The genus is referred to Thalassochelydia and has been considered a synonym of Eurysternum or Acichelys before, but casts that remain of the lost holotype show that it bears no features that can clarify its validity making it a nomen dubium.
Owadowia is a genus of extinct thalassochelydian turtle from the Late Jurassic of Poland. The type and only species is Owadowia borsukbialynickae, named by Szczygielski and colleagues in 2017 for a partial lower jaw, coracoid, ilium and femur from the early Tithonian Kcynia Formation. The limited material means that it is difficult to compare Owadowia to its relatives, and it may not be a unique taxon. The genus lacks the features diagnostic to its parent clade Thalassochelydia, has similarities to Solnhofia and Portlandemys as well as being a Late Jurassic marine turtle like the remainder of the group.
Pelobatochelys is a genus of extinct thalassochelydian turtle from the Late Jurassic of the United Kingdom. The type and only species is Pelobatochelys blakii, named by Harry Govier Seeley in 1875 for a central carapace fragment along with five additional specimens from the remainder of the shell from the early Kimmeridgian of the Kimmeridge Clay in Dorset, and additional material from the Kimmeridge Clay has also been referred to the taxon. The material was considered to be related to various turtles throughout time, including being reassigned to Tropidemys as T. blakii based on limited evidence. A large shell also from Weymouth, Dorset may represent further material of Pelobatochelys, but could also be from Tropidemys langii which was also present in the deposits.
Sandownidae is a family of extinct marine turtles from the Cretaceous and Paleogene distributed around the Atlantic Ocean and adjacent areas. The family is defined as all taxa closer to the type genus Sandownia than to Pelomedusa, Testudo, Solnhofia, Eurysternum, Plesiochelys, Thalassemys or Protostega, a definition that encompasses the previous concept of the clade while also excluding it from being synonymous with other clades of modern or extinct marine turtles. Sandownidae may be within the larger clade Angolachelonia, defined as inclusive of Angolachelys and Solnhofia, sister to the entirely Late Jurassic marine group Thalassochelydia, although the concepts of the clades may shift with further phylogenetic analysis.
Angolachelonia is a clade of extinct turtles from the Late Jurassic to Paleogene of Eurasia. The group is defined as all taxa derived from the ancestor of the type genus Angolachelys and Solnhofia, a definition that could potentially encompass a clade of entirely marine turtles. Angolachelonia was originally inclusive of only Solnhofia, Angolachelys and Sandownia when originally conceived by Octavio Mateus and colleagues in 2009, but later phylogenetic analyses by Serjocha Evers and Roger Benson in 2018 unites the family Sandownidae, including Angolachelys and Sandownia among other taxa, with the entirely Late Jurassic clade Thalassochelydia, where Solnhofia may be a basal member. While the placement of Solnhofia is weak and the clade that Angolachelonia represents may change with further analysis, the clade of Sandownidae and Thalassochelydia is well-supported, and does not collapse despite the uncertain evolutionary history of the group. Three alternative potential origins of Angolachelonia sensu Evers and Benson are shown below.
