Angolachelonia

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Angolachelonia
Temporal range: Late Jurassic to Paleogene, Oxfordian–Paleocene
Angolachelys mbaxi cropped.jpg
Angolachelys mbaxi
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Pantestudines
Clade: Testudinata
Clade: Mesochelydia
Clade: Perichelydia
Clade: Angolachelonia
Mateus et al., 2009
Subgroups [1]

Angolachelonia is a clade of extinct turtles from the Late Jurassic to Paleogene of Eurasia. The group is defined as all taxa derived from the ancestor of the type genus Angolachelys and Solnhofia , a definition that could potentially encompass a clade of entirely marine turtles. [1] Angolachelonia was originally inclusive of only Solnhofia, Angolachelys and Sandownia when originally conceived by Octavio Mateus and colleagues in 2009, [2] but later phylogenetic analyses by Serjocha Evers and Roger Benson in 2018 unites the family Sandownidae, including Angolachelys and Sandownia among other taxa, with the entirely Late Jurassic clade Thalassochelydia, where Solnhofia may be a basal member. While the placement of Solnhofia is weak and the clade that Angolachelonia represents may change with further analysis, the clade of Sandownidae and Thalassochelydia is well-supported, and does not collapse despite the uncertain evolutionary history of the group. Three alternative potential origins of Angolachelonia sensu Evers and Benson are shown below. [1]

A 2021 study found more evidence for Sandownidae and Thalassochelydia being close to one another, and found that Protostegidae may also be closely linked to the group. [3]

Related Research Articles

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<span class="mw-page-title-main">Cryptodira</span> Suborder of reptiles

The Cryptodira are a suborder of Testudines that includes most living tortoises and turtles. Cryptodira differ from Pleurodira in that they lower their necks and pull the heads straight back into the shells, instead of folding their necks sideways along the body under the shells' marginals. They include among their species freshwater turtles, snapping turtles, tortoises, softshell turtles, and sea turtles.

<span class="mw-page-title-main">Megalosauridae</span> Extinct family of dinosaurs

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Paracryptodira is an extinct group of reptiles in the clade Testudinata, known from the Jurassic to Paleogene of North America and Europe. Initially treated as a suborder sister to Cryptodira, they were then thought to be a very primitive lineage inside the Cryptodira according to the most common use of the latter taxon. They are now often regarded as late-diverging stem-turtles, lying outside the clade formed by Cryptodira and Pleurodira. Paracryptodires are divided into three main groups, Compsemydidae, known from the Late Jurassic to Paleocene of North America and Europe, Pleurosternidae, known from the Late Jurassic to Early Cretaceous of North America and Europe, and Baenidae, known from the Early Cretaceous to Eocene of North America. The latter two groups are more closely related to each other than to Compsemys, forming the clade Baenoidea.

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<span class="mw-page-title-main">Microcleididae</span> Extinct family of reptiles

Microcleididae is an extinct family of basal plesiosauroid plesiosaurs from the Early Jurassic of France, Germany, Portugal and the United Kingdom. Currently, the oldest and the most known microcleidid is Eretmosaurus from the middle Sinemurian of the United Kingdom. Microcleididae was formally named and described by Roger B. J. Benson, Mark Evans and Patrick S. Druckenmiller in 2012.

<span class="mw-page-title-main">Plesiochelyidae</span> Extinct family of turtles

The Plesiochelyidae are an extinct family of turtles in the clade Thalassochelydia originally classified within the Cryptodira suborder, mostly belonging from the Jurassic period. An alternate study placed the clade Thalassochelydia in the Angolachelonia and outside the Testudines.

Hydropelta is a genus of Late Jurassic turtle from marine deposits in the Jura Mountains of eastern France.

<span class="mw-page-title-main">Perichelydia</span> Clade of reptiles

Perichelydia is a clade within Pantestudines known from the Middle Jurassic to Holocene. Alongside crown group Testudines, it also contains Helochelydridae, which is known from the Cretaceous of Europe and North America, Sichuanchelyidae from the Middle Jurassic to Paleocene of Asia and Europe, Meiolaniformes, which is known from the Cretaceous to Holocene of South America, Australia and Oceania, and Spoochelys, known from the Mid-Cretaceous Griman Creek Formation of Australia. Kallokibotion from the Late Cretaceous of Europe is also considered part of this group. Several other groups, including the proposed clade Angolachelonia, Paracryptodira, Macrobaenidae, Sinemydidae and Xinjiangchelyidae, which are sometimes considered members of Cryptodira, have also been found outside crown Testudines in several analyses. These groups are usually considered to be closer to the crown group than the other members of Perichelydia.

<span class="mw-page-title-main">Thalassochelydia</span> Extinct clade of turtles

Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including Eurysternum, Plesiochelys and Thalassemys to the exclusion of Pelomedusa, Testudo and Protostega. While a clade uniting the families Eurysternidae, Plesiochelyidae and Thalassemydidae had been supported by phylogenetic evidence, a name was not given for the clade until 2017, when Jérémy Anquetin and colleagues coined Thalassochelydia.

<i>Solnhofia</i> Extinct genus of turtles

Solnhofia is a genus of extinct thalassochelydian turtle from the Late Jurassic of Germany. The type species is Solnhofia parsonsi, named by Gaffney in 1975 for a partial skull and jaw from the early Tithonian of the Solnhofen Formation in Bavaria. Additional material including a complete skeleton is known from the late Kimmeridgian of Switzerland and the Kimmeridgian/Tithonian of other deposits within Bavaria, and potentially also unprepared material from the Late Jurassic of France. The genus was referred to the family Eurysternidae by Anquetin and colleagues in 2017, which may represent an artificial grade of early thalassochelydians. In 2020 a new species Solnhofia brachyrhyncha was described from the Kimmeridigan aged Reuchenette Formation of Switzerland.

<span class="mw-page-title-main">Sandownidae</span> Extinct family of turtles

Sandownidae is a family of extinct marine turtles from the Cretaceous and Paleogene distributed around the Atlantic Ocean and adjacent areas. The family is defined as all taxa closer to the type genus Sandownia than to Pelomedusa, Testudo, Solnhofia, Eurysternum, Plesiochelys, Thalassemys or Protostega, a definition that encompasses the previous concept of the clade while also excluding it from being synonymous with other clades of modern or extinct marine turtles. Sandownidae may be within the larger clade Angolachelonia, defined as inclusive of Angolachelys and Solnhofia, sister to the entirely Late Jurassic marine group Thalassochelydia, although the concepts of the clades may shift with further phylogenetic analysis.

<span class="mw-page-title-main">Compsemydidae</span> Extinct family of turtles

Compsemydidae is an extinct family of turtles, likely belonging to the clade Paracryptodira. The earliest undisputed member is Tongemys from the Berriasian age of the Early Cretaceous; two Late Jurassic genera have also sometimes been included in the group, but may alternatively be members of the family Pleurosternidae. The genus Compsemys survived the Cretaceous–Paleogene extinction event and lasted until the Thanetian age of the Paleocene.

References

  1. 1 2 3 Evers, S.W.; Benson, R.B.J. (2018). "A new phylogenetic hypothesis of turtles with implications for the timing and number of evolutionary transitions to marine lifestyles in the group". Palaeontology. 62 (2): 93–134. doi:10.1111/pala.12384.
  2. Octávio Mateus; Louis Jacobs; Michael Polcyn; Anne S. Schulp; Diana Vineyard; André Buta Neto; Miguel Telles Antunes (2009). "The oldest African eucryptodiran turtle from the Cretaceous of Angola" (PDF). Acta Palaeontologica Polonica. 54 (4): 581–588. doi: 10.4202/app.2008.0063 . S2CID   55919209.
  3. Joyce, Walter G.; Anquetin, Jérémy; Cadena, Edwin-Alberto; Claude, Julien; Danilov, Igor G.; Evers, Serjoscha W.; Ferreira, Gabriel S.; Gentry, Andrew D.; Georgalis, Georgios L.; Lyson, Tyler R.; Pérez-García, Adán (2021-02-09). "A nomenclature for fossil and living turtles using phylogenetically defined clade names". Swiss Journal of Palaeontology . 140 (1): 5. doi: 10.1186/s13358-020-00211-x . hdl: 11336/155192 . ISSN   1664-2384.