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Tritylodon Temporal range: Early Jurassic | |
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Tritylodon longaevus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Clade: | Cynodontia |
Family: | † Tritylodontidae |
Genus: | † Tritylodon Owen, 1884 |
Type species | |
Tritylodon longaevus Owen, 1884 | |
Species | |
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Tritylodon (from the Greek for "three-cusped tooth") is an extinct genus of tritylodonts, one of the most advanced group of cynodont therapsids. They lived in the Early Jurassic and possibly Late Triassic periods along with dinosaurs. They also shared many characteristics with mammals, and were once considered mammals because of overall skeleton construction. That was changed due to them retaining the vestigial amniote jawbones and a different skull structure. Tritylodonts are now regarded as non-mammalian synapsids. [1]
If a living Tritylodon were to be seen today, it would look a lot like a large rodent. They were about 30 centimetres (12 in) long but there is no certainty about the exact weight. Their method of chewing food, a grinding motion with the bottom teeth sliding against the top teeth, resembled that of rodents as well. The bottom teeth were much like a set of cusps and the top teeth were a set of matching grooves that matched perfectly allowing this motion. There were large incisors at the very front of their mouth separated by a gap from the rest of the teeth. The incisors would stick out and remain slightly visible when the mouth was closed. The legs were directly beneath the body like mammals, unlike the earlier therapsids with sprawling limbs.[ citation needed ]
These animals were burrowers; the structure of the shoulder, front limbs, and large front incisors show this. They used their incisors to help dig and unearth buried plants. The way they ate and the shape of their teeth demonstrate that Tritylodons were probably primarily herbivorous (though some tritylodontids show evidence of more omnivorous diets, and modern analogues like rodents tend to be more omnivorous than their dentitions lead on [2] ). Any of the Tritylodonts including Tritylodon were warm-blooded or endothermic. Like most non-placental mammalimorphs, it had epipubic bones, aiding in its erect gait but preventing the expansion of the abdomen, making it unable to go through prolonged pregnancy and instead give birth to larval young like modern marsupials and monotremes. [3]
The Tritylodons' habitat was limited to the forests of South Africa When the species originated, about 200 million years ago, the African area was drier and hotter. But for most of their existence the climate was tropical and wetter.
The Tritylodon fossils in South Africa are found concentrated mainly in an area about 11,000 km2 (4,250 mi2). They have been found in floodplain deposits of the Lower Jurassic Elliot Formation (upper Karoo Supergroup). [4] In this area there have been so many findings it has been named the Tritylodon Acme Zone. [5] The fossil findings have all been in the Free State of South Africa. [5]
The genus Tritylodon of the Tritylodonts is restricted to the South African forms: Tritylodon longaevus and Tritylodon maximus. [1] It is suggested that T. maximus is either a large T. longeavus or a closely related species. If it is a closely related species it could possibly be ecological succession since the larger T. maximus fossils have been dated in the Sinemurian–Pliensbachian mainly less than 190 million years ago and the T. longaevus in the Hettangian–Sinemurian mainly more than 190 million years ago. With the fossil findings of each species overlapping in Sinemurian stage, the fossils show two differences, T. maximus being larger and having nine upper postcanines (neither species had canine teeth) instead of the seven teeth like T. longeavus. All other structures of the two Tritylodon species were the same. [6] [ self-published source? ]
Below is a cladogram from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: [7]
→ Probainognathia |
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Therapsida is a clade composing of a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, resulting in a more "standing" quadripedal posture, as opposed to the lower sprawling posture of many reptiles and amphibians.
Cynodontia is a clade of eutheriodont therapsids that first appeared in the Late Permian, and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extinct ancestors and close relatives (Mammaliaformes), having evolved from advanced probainognathian cynodonts during the Late Triassic.
Cynognathus is an extinct genus of large-bodied cynodontian therapsids that lived in the Middle Triassic. It is known from a single species, Cynognathus crateronotus. Cynognathus was a predator closely related to mammals and had a southern hemispheric distribution. Fossils have so far been recovered from South Africa, Argentina, Antarctica, and Namibia.
Dinocephalians are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania.
Thrinaxodon is an extinct genus of cynodonts, including the species T. liorhinus which lived in what are now South Africa and Antarctica during the Early Triassic. Thrinaxodon lived just after the Permian–Triassic mass extinction event, its survival during the extinction may have been due to its burrowing habits.
Oligokyphus is an extinct genus of herbivorous tritylodontid cynodont known from the Late Triassic to Early Jurassic of Europe, Asia and North America.
Mammaliaformes is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia, and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes includes Docodonta and Hadrocodium.
The theriodonts are a major group of therapsids which appeared during the Middle Permian and which includes the gorgonopsians and the eutheriodonts, itself including the therocephalians and the cynodonts.
Tritylodontidae is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy and details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous.
Theriognathus is an extinct genus of therocephalian therapsid belonging to the family Whaitsiidae, known from fossils from South Africa, Zambia, and Tanzania. Theriognathus has been dated as existing during the Late Permian. Although Theriognathus means mammal jaw, the lower jaw is actually made up of several bones as seen in modern reptiles, in contrast to mammals. Theriognathus displayed many different reptilian and mammalian characteristics. For example, Theriognathus had canine teeth like mammals, and a secondary palate, multiple bones in the mandible, and a typical reptilian jaw joint, all characteristics of reptiles. It is speculated that Theriognathus was either carnivorous or omnivorous based on its teeth, and was suited to hunting small prey in undergrowth. This synapsid adopted a sleek profile of a mammalian predator, with a narrow snout and around 1 meter long. Theriognathus is represented by 56 specimens in the fossil record.
Scylacops is an extinct genus of Gorgonopsia. It was first named by Broom in 1913, and contains two species, S. bigendens, and S. capensis. Its fossils have been found in South Africa and Zambia. It is believed to be closely related to the Gorgonopsian Sauroctonus progressus. Scylacops was a moderately sized Gorgonopsid.
Morganucodonta is an extinct order of basal Mammaliaformes, a group including crown-group mammals (Mammalia) and their close relatives. Their remains have been found in Southern Africa, Western Europe, North America, India and China. The morganucodontans were probably insectivorous and nocturnal, though like eutriconodonts some species attained large sizes and were carnivorous. Nocturnality is believed to have evolved in the earliest mammals in the Triassic as a specialisation that allowed them to exploit a safer, night-time niche, while most larger predators were likely to have been active during the day.
Cynosaurus is an extinct genus of cynodonts. Remains have been found from the Dicynodon Assemblage Zone in South Africa. Cynosaurus was first described by Richard Owen in 1876 as Cynosuchus suppostus. Cynosaurus has been found in the late Permian period. Cyno- is derived from the Greek word kyon for dog and –sauros in Greek meaning lizard.
Progalesaurus is an extinct genus of galesaurid cynodont from the early Triassic. Progalesaurus is known from a single fossil of the species Progalesaurus lootsbergensis, found in the Lystrosaurus Assemblage Zone of the Balfour Formation. Close relatives of Progalesaurus, other galesaurids, include Galesaurus and Cynosaurus. Galesaurids appeared just before the Permian-Triassic extinction event, and disappeared from the fossil record in the Middle-Triassic.
Dinnebitodon is an extinct genus of tritylodontid mammaliamorphs from the Early Jurassic. It has only been found in the Kayenta Formation in northeastern Arizona. It closely resembles the related genus Kayentatherium from the same formation. It is set apart by differences in the dentition, while resembling in most other respects.
Kayentatherium is an extinct genus of tritylodontid cynodonts that lived during the Early Jurassic. It is one of two tritylodonts from the Kayenta Formation of northern Arizona, United States.
Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. They are thought to have been herbivorous or omnivorous. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil, eastern North America, Europe, China, and Antarctica.
Xenocretosuchus is an extinct genus of tritylodont therapsids from the Aptian Ilek Formation of Siberia, in the Russian Federation. The type species, X. sibiricus, is known only from dental elements, as is X. kolossovi, described from the Batylykh Formation in 2008. Some authors have treated these species as part of the genus Stereognathus, otherwise known from the Middle Jurassic of Britain, but this is rejected by other authors.
Abdalodon is an extinct genus of late Permian cynodonts, known by its only species A. diastematicus.Abdalodon together with the genus Charassognathus, form the clade Charassognathidae. This clade represents the earliest known cynodonts, and is the first known radiation of Permian cynodonts.
Pseudotherium is an extinct genus of prozostrodontian cynodonts from the Late Triassic of Argentina. It contains one species, P. argentinus, which was first described in 2019 from remains found in the La Peña Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin.