Langbergia

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Langbergia
Temporal range: Early Triassic, 251–247  Ma
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Langbergia.jpg
Figure 2 Stereopair and interpretative drawing of NMQR 3255, holotype of Langbergia modisei, in dorsal view. Ept, epipterygoid; L, lacrimal; Mx, maxilla; N, nasal; F, frontal; P, parietal; Pmx, premaxilla; Po, postorbital; Prf, prefrontal; Pro, prootic; Sq, squamosal; tf, trigeminal foramen (for maxillary and mandibular divisions of the trigeminal nerve). Note that bones in the facial region (maxillae, nasals, lacrimals, and prefrontals) are not in their original position. Shading indicates a broken portion (in the jugal) and sediment posteriorly of the postorbital bar. Scale bar = 3 cm.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Cynodontia
Family: Trirachodontidae
Subfamily: Trirachodontinae
Genus: Langbergia
Abdala et al., 2006
Species
  • L. modiseiAbdala et al., 2006 (type)

Langbergia is an extinct genus of trirachodontid cynodont from the Early Triassic of South Africa. The type and only species L. modisei was named in 2006 after the farm where the holotype was found, Langberg 566. [1] Langbergia was found in the Burgersdorp Formation in the Beaufort Group, a part of the Cynognathus Assemblage Zone. The closely related trirachodontids Trirachodon and Cricodon were found in the same area.

Contents

History of discovery

Langbergia was discovered in the Cynognathus Assemblage Zone of the Beaufort Group of South Africa in 2006. It was named after the farm where the holotype was found (Langberg 566).

Description and paleobiology

Trirachodontids are distinguishable due to the shape of their postcanines; the postcanines, distal to the canines, are transversely widened teeth with three main cusps disposed in a transverse row across the center of the tooth. [1] Langbergia are small gomphodontian cynodonts. [2] Cynodonts are a very important component of the Triassic. [3] After the Langbergia fossil was found in 2006, new diagnostic features allowed for a new taxon, Langbergia modisei. Unlike other Trirachodontids, the upper and lower gomphodont post caninines of Langbergia are elliptical in cross section, slightly labiolingually elongated and their long axis is longuomesially orientated. Additionally, both upper and lower lower gomphodont postcanines of Langbergia increase tooth size up until the penultimate tooth and the medial and distal basins are deep and mesiodistally short [4] Trirachodontids Trirachodon and Cricodon have a labial platform of the maxilla that arises lateral to the postcanine series, this is not observed in Langbergia. [5] Langbergia has been recorded having stapes, despite how fragile the structure is. [6]

Dentition of Langbergia modisei Dentition of Langbergia modisei.jpg
Dentition of Langbergia modisei

Langbergia, like most other living in the Permian, went extinct during the end-Permian mass extinction. [7]

Geological information

The Burgersdorp Formation in South Africa is known for having a substantial amount of fossiliferous rocks. [8] The Permo-Triassic strata of the Beaufort Group has a substantial amount of therapsid fossils which are used in the biostratigraphic subdivision. The uppermost division is the Cynognathus Assemblage Zone, this zone covers the late early and early middle Triassic. The Cynognathus Assemblage Zone is then divided into a threefold subdivision. [9] The uppermost subdivision of the Cynognathus Assemblage Zone is named the Lystrosaurus Assemblage Zone. The upper boundary of the Lystrosaurus Assemblage Zone is drawn at the Langbergia and the Garjainia. [10]

Figure 13 Majority-rule consensus tree of the analysis with characters having equal weights plotted against the geological time scale to indicate temporal distribution of the taxa included in the analysis (see Appendix 1). The geological time scale is based on Gradstein & Ogg (2004). Ans, Anisian; Crn, Carnian; E, Early; Ind, Induan; L, Late; Lad, Ladinian; M, Middle; Nor, Norian; Ol, Olenekian. Cladogram Langbergia.jpg
Figure 13 Majority-rule consensus tree of the analysis with characters having equal weights plotted against the geological time scale to indicate temporal distribution of the taxa included in the analysis (see Appendix 1). The geological time scale is based on Gradstein & Ogg (2004). Ans, Anisian; Crn, Carnian; E, Early; Ind, Induan; L, Late; Lad, Ladinian; M, Middle; Nor, Norian; Ol, Olenekian.

Related Research Articles

<i>Cynognathus</i> Extinct genus of cynodonts

Cynognathus is an extinct genus of large-bodied cynodontian therapsids that lived in the Middle Triassic. It is known from a single species, Cynognathus crateronotus. Cynognathus was a predator closely related to mammals and had a southern hemispheric distribution. Fossils have so far been recovered from South Africa, Argentina, Antarctica, and Namibia.

<span class="mw-page-title-main">Cynognathia</span> Clade of cynodonts

Cynognathia is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus Cynognathus and the herbivorous or omnivorous gomphodonts such as traversodontids. Cynognathians can be identified by several synapomorphies including a very deep zygomatic arch that extends above the middle of the orbit.

<span class="mw-page-title-main">Traversodontidae</span> Extinct family of cynodonts

Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil.

<i>Lystrosaurus</i> Assemblage Zone

The Lystrosaurus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the upper Adelaide and lower Tarkastad Subgroups of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. This biozone has outcrops in the south central Eastern Cape and in the southern and northeastern Free State. The Lystrosaurus Assemblage Zone is one of eight biozones found in the Beaufort Group, and is considered to be Early Triassic in age.

<i>Cynognathus</i> Assemblage Zone Biozone which correlates to the Burgersdorp Formation of the Beaufort Group

The Cynognathus Assemblage Zone is a tetrapod biozone utilized in the Karoo Basin of South Africa. It is equivalent to the Burgersdorp Formation, the youngest lithostratigraphic formation in the Beaufort Group, which is part of the fossiliferous and geologically important Karoo Supergroup. The Cynognathus Assemblage Zone is the youngest of the eight biozones found in the Beaufort Group, and is considered to be late Early Triassic (Olenekian) to early Middle Triassic (Anisian) in age. The name of the biozone refers to Cynognathus crateronotus, a large and carnivorous cynodont therapsid which occurs throughout the entire biozone.

<i>Theriognathus</i> Extinct genus of therapsids from late Permian South Africa and Tanzania

Theriognathus is an extinct genus of therocephalian therapsid belonging to the family Whaitsiidae, known from fossils from South Africa, Zambia, and Tanzania. Theriognathus has been dated as existing during the Late Permian. Although Theriognathus means mammal jaw, the lower jaw is actually made up of several bones as seen in modern reptiles, in contrast to mammals. Theriognathus displayed many different reptilian and mammalian characteristics. For example, Theriognathus had canine teeth like mammals, and a secondary palate, multiple bones in the mandible, and a typical reptilian jaw joint, all characteristics of reptiles. It is speculated that Theriognathus was either carnivorous or omnivorous based on its teeth, and was suited to hunting small prey in undergrowth. This synapsid adopted a sleek profile of a mammalian predator, with a narrow snout and around 1 meter long. Theriognathus is represented by 56 specimens in the fossil record.

<i>Progalesaurus</i> Extinct genus of cynodonts

Progalesaurus is an extinct genus of galesaurid cynodont from the early Triassic. Progalesaurus is known from a single fossil of the species Progalesaurus lootsbergensis, found in the Lystrosaurus Assemblage Zone of the Balfour Formation. Close relatives of Progalesaurus, other galesaurids, include Galesaurus and Cynosaurus. Galesaurids appeared just before the Permian-Triassic extinction event, and disappeared from the fossil record in the Middle-Triassic.

Platycraniellus is an extinct genus of carnivorous cynodonts from the Early Triassic. It is known from the Lystrosaurus Assemblage Zone of the Normandien Formation in South Africa. P. elegans is the only species in this genus based on the holotype specimen from the Ditsong National Museum of Natural History in Pretoria, South Africa. Due to limited fossil records for study, Platycraniellus has only been briefly described a handful of times.

<i>Trirachodon</i> Extinct genus of cynodonts

Trirachodon is an extinct genus of cynodonts. Fossils have been found in the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Omingonde Formation of Namibia, dating back to the Early and Middle Triassic.

<i>Lumkuia</i> Extinct genus of cynodonts

Lumkuia is an extinct genus of cynodonts, fossils of which have been found in the Cynognathus Assemblage Zone of the Beaufort Group in the South African Karoo Basin that date back to the early Middle Triassic. It contains a single species, Lumkuia fuzzi, which was named in 2001 on the basis of the holotype specimen BP/1/2669, which can now be found at the Bernard Price Institute in Johannesburg, South Africa. The genus has been placed in its own family, Lumkuiidae. Lumkuia is not as common as other cynodonts from the same locality such as Diademodon and Trirachodon.

<i>Diademodon</i> Extinct genus of cynodonts

Diademodon is an extinct genus of cynodonts. It was about 2 metres (6.6 ft) long.

Neotrirachodon is an extinct genus of therapsids which existed in Russia during the Middle Triassic period. Its type and only species is Neotrirachodon expectatus.

<span class="mw-page-title-main">Trirachodontidae</span> Extinct family of cynodonts

Trirachodontidae is an extinct, possibly paraphyletic family of cynognathian cynodonts from the Triassic of China and southern Africa. Trirachodontids appeared during the Early Triassic soon after the Permian-Triassic extinction event and quickly spread over a wide geographic area in a comparatively brief amount of time from 250 to 237 million years ago.

<span class="mw-page-title-main">Gomphodontia</span> Clade of cynodonts

Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. They are thought to have been herbivorous or omnivorous. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil, eastern North America, Europe, China, and Antarctica.

Titanogomphodon is an extinct genus of diademodontid cynodonts from the Middle Triassic Omingonde Formation of Namibia. It is known from a single partial skull that was described in 1973 from the Omingonde Formation. The type and only species is Titanogomphodon crassus. At about 40 centimetres (16 in), the skull of Titanogomphodon was significantly larger than that of its closest relative, Diademodon. Its teeth are similar to those of another group of cynodonts called Traversodontidae, but the similarities are likely the result of convergent evolution. Aside from its larger size, Titanogomphodon differs from Diademodon in having a bony projection on the postorbital bar behind the eye socket.

<i>Cricodon</i> Extinct genus of cynodonts

Cricodon is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa. A. W. Crompton named Cricodon based on the ring-like arrangement of the cuspules on the crown of a typical postcanine tooth. The epithet of the type species, C. metabolus, indicates the change in structure of certain postcanines resulting from replacement.

<i>Abdalodon</i> Extinct genus of cynodonts

Abdalodon is an extinct genus of late Permian cynodonts, known by its only species A. diastematicus.Abdalodon together with the genus Charassognathus, form the clade Charassognathidae. This clade represents the earliest known cynodonts, and is the first known radiation of Permian cynodonts.

<i>Ufudocyclops</i> Extinct genus of dicynodonts

Ufudocyclops is an extinct genus of stahleckeriid dicynodont from the Middle Triassic of South Africa. It was found in the Burgersdorp Formation, part of the uppermost Cynognathus Assemblage Zone of the Beaufort Group in the Karoo Basin. The type and only known species is U. mukanelai. It was a large, beaked herbivore like other Triassic dicynodonts, lacking tusks, and is mostly characterised by unique features of the skull. It is known from three specimens, two of which were previously referred to the Tanzanian dicynodont Angonisaurus. The separation of Ufudocyclops from Angonisaurus indicates that the Middle Triassic fauna of the Beaufort Group in South Africa was not part of a larger shared fauna with those of the Manda Beds in Tanzania, as was previously supposed, and suggests that they were separated as more localised faunas, possibly by geographic barriers or in time. Ufudocyclops then would have been a unique part of the uppermost Cynognathus Assemblage Zone in South Africa. It is also the oldest known member of the family Stahleckeriidae, and implies that the family was already diversifying in the Middle Triassic alongside other kannemeyeriiforms, not just in the Late Triassic after other families died out.

<i>Etjoia</i> Extinct genus of cynodonts

Etjoia is an extinct genus of traversodontid cynodonts that lived during the Middle Triassic or Late Triassic period in southern Africa. This medium-sized omnivorous cynognathian provides important information on the dental evolution of early diverging gomphodonts and traversodontids.

<i>Impidens</i> Extinct genus of cynodonts

Impidens is an extinct genus of large omnivorous cynodont from the Triassic of South Africa and Antarctica. Its type and only species is Impidens hancoxi. Impidens inhabited high-latitude environments of southern Gondwana during the Middle Triassic, where it was probably the apex predator.

References

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  2. Sidor, Christian A.; Hopson, James A. (2017-11-29). "Cricodon metabolus(Cynodontia: Gomphodontia) from the Triassic Ntawere Formation of northeastern Zambia: patterns of tooth replacement and a systematic review of the Trirachodontidae". Journal of Vertebrate Paleontology. 37 (sup1): 39–64. Bibcode:2017JVPal..37S..39S. doi:10.1080/02724634.2017.1410485. ISSN   0272-4634. S2CID   89932366.
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