Impidens

Impidens Temporal range: Late Anisian–early Ladinian, possibly early Carnian PreꞒ Ꞓ O S D C P T J K Pg N
Life restoration of Impidens hancoxi
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Synapsida
Clade:	Therapsida
Clade:	Cynodontia
Family:	† Trirachodontidae
Genus:	† Impidens Tolchard, Kammerer, Butler, Hendrickx, Benoit, Abdala, & Choiniere, 2021
Type species
Impidens hancoxi Tolchard, Kammerer, Butler, Hendrickx, Benoit, Abdala, & Choiniere, 2021

Impidens ^[a] is an extinct genus of large omnivorous cynodont from the Triassic of South Africa and Antarctica. Its type and only species is Impidens hancoxi. ^[b] Impidens inhabited high-latitude environments of southern Gondwana during the Middle Triassic, where it was probably the apex predator.

History

A specimen of Impidens was collected from Antarctica in 1986. In 1995, it was described as an indeterminate diademodontid possibly belonging to Titanogomphodon . ^[2] The holotype of Impidens hancoxi was found in South Africa in 2014. ^[3] It was described as a new species in 2021, with the Antarctic specimen and a less complete specimen from South Africa being referred to the species. ^[1]

Description

Impidens was one of the largest non-mammalian cynodonts, with a skull well over 400 millimetres (16 in) long, though the herbivorous Scalenodontoides from the Late Triassic was even larger. ^[4]

Classification

Impidens is a member of Trirachodontidae, a family of gomphodont cynodonts. It is closely related to its smaller contemporary Cricodon as well as the earlier Langbergia and Trirachodon . ^[5] However, the phylogeny of Trirachodontidae is not well-understood, and the family may be paraphyletic, with some species more closely related to traversodontids than others.

Paleoecology

Impidens fossils are known from the Cricodon-Ufudocyclops subzone of the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the upper Fremouw Formation in Antarctica. The dating of these strata is controversial; they are conventionally regarded as Middle Triassic in age, probably late Anisian or early Ladinian, but a biostratigraphically correlated fauna in South America has been dated to the beginning of the Late Triassic, in the early Carnian. ^[6] However, another biostratigraphically correlated fauna in China has been dated to the Anisian, suggesting the traditional dating may be correct. ^[7]

Impidens was probably the apex predator of its environment, as the only other large predator known to have coexisted with it was the somewhat smaller cynodont Cynognathus . The known large herbivores of the fauna are the cynodont Diademodon and the dicynodont Ufudocyclops . A smaller trirachodontid, Cricodon , was also present in the environment.

The Cricodon-Ufudocyclops subzone was deposited in a meandering river environment, with a deep channel and high banks. ^[8] Both it and the Fremouw Formation were high-paleolatitude environments, which may explain some of the faunal differences between them and lower-paleolatitude fauna. ^[4]

Footnotes

1. From the isiZulu impi "combat" and the Latin dens "tooth" ^[1]
2. Named in honor of Dr. John Hancox, a pioneering researcher of the strata it was found in. ^[1]

References

1. Tolchard et al. 2021, p. 4.
2. Hammer 1995, pp. 108–109.
3. Tolchard et al. 2021, p. 3.
4. Tolchard et al. 2021, p. 14.
5. Tolchard et al. 2021, p. 12–13.
6. Hancox, Neveling & Rubidge 2020, p. 234.
7. Tolchard et al. 2021, p. 2.
8. Hancox, Neveling & Rubidge 2020, p. 232.

Bibliography

• Hammer, William R. (1995-03-14). "New Therapsids From the Upper Fremouw Formation (Triassic) of Antarctica" . Journal of Vertebrate Paleontology. 15 (1): 105–112. Bibcode:1995JVPal..15..105H. doi:10.1080/02724634.1995.10011210. ISSN   0272-4634.
• Hancox, P.J.; Neveling, J.; Rubidge, B.S. (2020-06-01). "Biostratigraphy of the Cynognathus Assemblage Zone (Beaufort Group, Karoo Supergroup), South Africa" . South African Journal of Geology. 123 (2): 217–238. Bibcode:2020SAJG..123..217H. doi:10.25131/sajg.123.0016. eISSN   1996-8590. ISSN   1012-0750.
• Tolchard, Frederick; Kammerer, Christian F.; Butler, Richard J.; Hendrickx, Christophe; Benoit, Julien; Abdala, Fernando; Choiniere, Jonah N. (2021-07-26). "A new large gomphodont from the Triassic of South Africa and its implications for Gondwanan biostratigraphy" . Journal of Vertebrate Paleontology. 41 (2): e1929265. Bibcode:2021JVPal..41E9265T. doi:10.1080/02724634.2021.1929265. ISSN   0272-4634.