Boreogomphodon Temporal range: Late Triassic | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Clade: | Cynodontia |
Family: | † Traversodontidae |
Subfamily: | † Arctotraversodontinae |
Genus: | † Boreogomphodon Sues and Olsen, 1990 |
Species | |
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Boreogomphodon is an extinct genus of traversodontid cynodonts from the Late Triassic of the eastern United States. Fossils have been found from the Turkey Branch Formation in Virginia and the Pekin Formation of North Carolina.
Boreogomphodon jeffersoni was named in 1990. Its teeth and cranial bones are the most common tetrapod fossils in the Turkey Branch Formation. Remain assigned to B. jeffersoni are also known from the Pekin Formation of North Carolina. [1] Boreogomphodon was distinguished from other traversodontids like the African Luangwa and the South American Traversodon on the basis of its postcanine teeth. Most traversodontids have lower postcanine teeth with two cusps, but Boreogomphodon was the first traversodontid found with three cusps on its lower postcanine teeth. There is a single cusp on the side of the upper postcanine facing the lip, while a flat surface extends outward from it. [2] The traversodontid Arctotraversodon from the Wolfville Formation of Nova Scotia is similar to Boreogomphodon in that it has three cusps on its lower postcanine, but its postcanines are much wider than they are long relative to those of Boreogomphodon. [3]
Boreogomphodon specimens likely represent juvenile individuals because of their small size. The snout is short, while larger, presumably mature traversodonts have longer snouts. Boreogomphodon specimens have only four or five postcanine teeth. Traversodontids acquire more of these teeth as they grow, suggesting that Boreogomphodon individuals were not fully mature. Boreogomphodon also has some sectorial, or cutting, postcanine teeth, which are not present in most larger traversodontids. [2]
Boreogomphodon has several primitive features, including a depression in the upper jaw called the paracanine fossa in front of the canine tooth. This fossa provides space for the lower canine tooth when the jaw is closed. Above the upper canine at the top of the snout there is a small depression on the maxilla. The tip of the snout is covered in irregular pits, a unique feature of Boreogomphodon. Each upper postcanine has large central cusp and a posterior cingulum. [4]
Boreogomphodon is closely related to Arctotraversodon and Nanogomphodon . The three North American traversodontids form a somewhat basal clade within Traversodontidae. Below is a cladogram modified from Sues and Hopson (2010) showing the phylogenetic relations of Boreogomphodon: [4]
Thrinaxodon is an extinct genus of cynodonts, including the species T. liorhinus which lived in what are now South Africa and Antarctica during the Early Triassic. Thrinaxodon lived just after the Permian–Triassic mass extinction event, its survival during the extinction may have been due to its burrowing habits.
Tritylodontidae is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy, and some details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous.
Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil.
Brasilodon is an extinct genus of small, mammal-like cynodonts that lived in what is now Brazil during the Norian age of the Late Triassic epoch, about 225.42 million years ago. While no complete skeletons have been found, the length of Brasilodon has been estimated at 12 centimetres (4.7 in). Its dentition shows that it was most likely an insectivore. The genus is monotypic, containing only the species B. quadrangularis. Brasilodon belongs to the family Brasilodontidae, whose members were some of the closest relatives of mammals, the only cynodonts alive today. Two other brasilodontid genera, Brasilitherium and Minicynodon, are now considered to be junior synonyms of Brasilodon.
Cynosaurus is an extinct genus of cynodonts. Remains have been found from the Dicynodon Assemblage Zone in South Africa. Cynosaurus was first described by Richard Owen in 1876 as Cynosuchus suppostus. Cynosaurus has been found in the late Permian period. Cyno- is derived from the Greek word kyon for dog and –sauros in Greek meaning lizard.
Platycraniellus is an extinct genus of carnivorous cynodonts from the Early Triassic. It is known from the Lystrosaurus Assemblage Zone of the Normandien Formation in South Africa. P. elegans is the only species in this genus based on the holotype specimen from the Ditsong National Museum of Natural History in Pretoria, South Africa. Due to limited fossil records for study, Platycraniellus has only been briefly described a handful of times.
Menadon is an extinct genus of traversodontid cynodonts. The type and only species is Menadon besairiei.
Andescynodon is a genus of traversodontid cynodonts from the Middle Triassic of Argentina. Fossils are known from the Cerro de las Cabras and Cacheutá Formations. Andescynodon is one of the most basal traversodontids. Another traversodontid called Rusconiodon has also been identified from the Cerro de las Cabras Formation but is now considered a junior synonym of Andescynodon.
Pascualgnathus is an extinct genus of traversodontid cynodonts from the Middle Triassic of Argentina. Fossils have been found from the Río Seco de la Quebrada Formation of the Puesto Viejo Group. The type species P. polanskii was named in 1966.
Arctotraversodon is an extinct genus of traversodontid cynodonts from the Late Triassic of Canada. Fossils first described from the Wolfville Formation in Nova Scotia in 1984 represented the first known traversodontid from North America. The type and only species is A. plemmyridon and is represented by teeth and several dentary bones.
Plinthogomphodon is an extinct genus of traversodontid cynodonts from the Late Triassic of the eastern United States. Fossils have been found from the Deep River basin of North Carolina, part of the larger Newark Supergroup. It is known from a single eroded snout. The type and only species is P. herpetairus.
Massetognathinae is an extinct subfamily of cynodonts in the family Traversodontidae. It includes four species from the Middle and Late Triassic: Massetognathus pascuali from Argentina, Massetognathus ochagaviae and Santacruzodon hopsoni from southern Brazil, and Dadadon isaloi from Madagascar. Massetognathines have several distinguishing characteristics, including flattened skulls, small canine teeth, and postcanine teeth with three cusps on their outer edges. Massetgognathinae was defined by Kammerer et al. (2012) as the clade containing all traversodontids more closely related to Massetognathus pascuali than to Gomphodontosuchus brasiliensis, and is the sister taxon of the traversodontid subfamily Gomphodontosuchinae, which was defined by Kammerer et al. (2008) as all traverodontids more closely related to G. brasiliensis than to M. pascuali.
Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. They are thought to have been herbivorous or omnivorous. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil, eastern North America, Europe, China, and Antarctica.
Ruberodon is an extinct genus of traversodontid cynodonts known from the type and only species Ruberodon roychowdhurii from the Late Triassic of India. Ruberodon was named in 2015 on the basis of several isolated lower jaws found in the Tiki Formation. The lower jaw of Ruberodon has three pairs of incisors, one pair of canines, and 9 pairs of postcanine teeth. The first pair of incisors is enlarged and protrudes forward from the tip of the jaw and there is a gap called a diastema between the canines and postcanines. Phylogenetic analysis indicates that among traversodontids, R. roychowdhurii is most closely related to Exaeretodon statisticae, which is also from India.
Cricodon is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa. A. W. Crompton named Cricodon based on the ring-like arrangement of the cuspules on the crown of a typical postcanine tooth. The epithet of the type species, C. metabolus, indicates the change in structure of certain postcanines resulting from replacement.
Abdalodon is an extinct genus of late Permian cynodonts, known by its only species A. diastematicus.Abdalodon together with the genus Charassognathus, form the clade Charassognathidae. This clade represents the earliest known cynodonts, and is the first known radiation of Permian cynodonts.
Etjoia is an extinct genus of traversodontid cynodonts that lived during the Middle Triassic or Late Triassic period in southern Africa. This medium-sized omnivorous cynognathian provides important information on the dental evolution of early diverging gomphodonts and traversodontids.
Bonacynodon is an extinct genus of cynodonts that lived in what is now southern Brazil during the Triassic period. The genus is monotypic, containing only the type species Bonacynodon schultzi. B. schultzi is known from two specimens, consisting of two partial skulls and some badly preserved parts of the postcranium. Both specimens were recovered from the Pinheiros-Chiniquá Sequence, part of the Santa Maria Supersequence of the Paraná Basin. This sequence preserves a faunal association known as the Dinodontosaurus Assemblage Zone, which contains numerous other species of cynodonts, dicynodonts and reptiles. Bonacynodon was a small, likely insectivorous cynodont, whose length has been estimated at around 30 centimetres (12 in). It can be distinguished from other cynodonts by its large, serrated (saw-like) canine teeth. Together with the genus Probainognathus of Argentina, it made up the family Probainognathidae, one of the earliest-diverging lineages of the clade Probainognathia. It was a fairly close relative of mammals, the only group of cynodonts alive today.
Scalenodontoides is an extinct genus of Traversodontidae, a family of herbivorous cynodonts. It lived during the Late Triassic in what is now South Africa. Its type species is Scalenodontoides macrodontes. It was named in 1957 by A. W. Crompton and F. Ellenberger. Arctotraversodon plemmyrodon was originally classified as a species of Scalenodontoides, but was given its own genus in 1992. It is found in the Scalenodontoides Assemblage Zone of the Elliot Formation, which is named for it. It is one of the geologically youngest traversodontids, alongside the putative traversodontid Boreogomphodon. It is closely related to Exaeretodon and Siriusgnathus, but is distinguished by the presence of a shelf-like expansion of its parietal called the nuchal table. Though the largest known complete skull is only 248 millimetres (9.8 in) long, it may have been the largest non-mammalian cynodont, as an incomplete snout would have belonged to a specimen with an estimated skull length of 617 millimetres (24.3 in).