Cynognathus Temporal range: Middle Triassic, | |
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Fossil skull of Cynognathus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Clade: | Cynodontia |
Clade: | † Cynognathia |
Family: | † Cynognathidae Seeley, 1895 |
Genus: | † Cynognathus Seeley, 1895 |
Type species | |
†Cynognathus crateronotus Seeley, 1895 |
Cynognathus is an extinct genus of large-bodied cynodontian therapsids that lived in the Middle Triassic. It is known from a single species, Cynognathus crateronotus. Cynognathus was a predator closely related to mammals and had a southern hemispheric distribution. Fossils have so far been recovered from South Africa, Argentina, Antarctica, and Namibia.
Cynognathus was a heavily built animal, and measured around 1.2 metres (3 ft 11 in) [1] in snout-to-vent body length and up to 2 metres (6 ft 7 in) in total length. [2] It had a particularly large head, up to 40 centimetres (1 ft) in length, with wide jaws and sharp teeth. Its hindlimbs were placed directly beneath the body, but the forelimbs sprawled outwards in a more reptilian fashion. [3] This form of double (erect/sprawling) gait is also found in some primitive mammals alive today. [4]
Possible autapomorphies of C. crateronotus include an extremely elongated postorbital bar and sectorial postcanine teeth with two serrated cusps distal to a recurved apex. [5]
During 1888 and 1889, the British paleontologist Harry Govier Seeley visited southern Africa. In 1889, near Lady Frere, at a location where earlier Alfred Brown had discovered a tooth, Seeley excavated a skull and partial postcranial skeleton of a cynodontian. In 1894, Seeley named the genus Cynognathus with as type species Cynognathus crateronotus. Simultaneously, he named three other species in the genus: Cynognathus berryi, honouring James Berry who had assisted in the excavations, Cynognathus platyceps, the "flat jaw", and Cynognathus leptorhinus, the "slender nose". [6] The generic name Cynognathus is derived from Greek kyon and gnathos, meaning "dog jaw". In 1895, Seeley published a more comprehensive description of these finds. [7]
Fossil material probably belonging to the genus has been given several different names over the years. Generic synonyms include Cynidiognathus, Cynogomphius, Karoomys, Lycaenognathus, Lycochampsa and Lycognathus. Opinions vary as to whether all remains belong to the same species. The genus Karoomys is known only from a tiny juvenile. Species-level synonyms of Cynognathus crateronotus include Cynidiognathus broomi, Cynidiognathus longiceps, Cynidiognathus merenskyi, Cynognathus berryi, Cynognathus minor, Cynognathus platyceps, Cynogomphius berryi, Karoomys browni, Lycaenognathus platyceps, Lycochampsa ferox, Lycognathus ferox, and Nythosaurus browni.
Fossils have been found in the Karoo, the Puesto Viejo Formation, Fremouw Formation, in South Africa/Lesotho, Argentina and Antarctica.
Cynognathus lived between the Anisian and the Ladinian (Middle Triassic). [8]
This genus forms a Cynognathus Assemblage Zone in the Beaufort Group of the Karoo Supergroup. [9] [10] [11]
Seeley in 1894/1895 placed Cynognathus in a separate family Cynognathidae, within the Cynodontia. Cynognathus is presently the only recognized member of the family Cynognathidae. Later a clade Cynognathia was named after the genus, within the Eucynodontia.
Cynognathus crateronotus in a cladogram after Stefanello et al. (2023): [12]
The dentary was equipped with differentiated teeth that show this animal could effectively process its food before swallowing. The presence of a secondary palate in the mouth indicates that Cynognathus would have been able to breathe and swallow simultaneously.
The possible lack of belly ribs, in the stomach region, suggests the presence of an efficient diaphragm: an important muscle for mammalian breathing. Pits and canals on the bone of the snout indicate concentrations of nerves and blood vessels. In mammals, such structures allow hairs (whiskers) to be used as sensory organs.
Cynodontia is a clade of eutheriodont therapsids that first appeared in the Late Permian, and extensively diversified after the Permian–Triassic extinction event. Mammals are cynodonts, as are their extinct ancestors and close relatives (Mammaliaformes), having evolved from advanced probainognathian cynodonts during the Late Triassic.
The theriodonts are a major group of therapsids which appeared during the Middle Permian and which includes the gorgonopsians and the eutheriodonts, itself including the therocephalians and the cynodonts.
Cynognathia is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus Cynognathus and the herbivorous or omnivorous gomphodonts such as traversodontids. Cynognathians can be identified by several synapomorphies including a very deep zygomatic arch that extends above the middle of the orbit.
The Daptocephalus Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important Group of the Karoo Supergroup in South Africa. This biozone has outcrops located in the upper Teekloof Formation west of 24°E, the majority of the Balfour Formation east of 24°E, and the Normandien Formation in the north. It has numerous localities which are spread out from Colesberg in the Northern Cape, Graaff-Reniet to Mthatha in the Eastern Cape, and from Bloemfontein to Harrismith in the Free State. The Daptocephalus Assemblage Zone is one of eight biozones found in the Beaufort Group and is considered Late Permian (Lopingian) in age. Its contact with the overlying Lystrosaurus Assemblage Zone marks the Permian-Triassic boundary.
The Lystrosaurus Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the upper Adelaide and lower Tarkastad Subgroups of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. This biozone has outcrops in the south central Eastern Cape and in the southern and northeastern Free State. The Lystrosaurus Assemblage Zone is one of eight biozones found in the Beaufort Group, and is considered to be Early Triassic in age.
The Cynognathus Assemblage Zone is a tetrapod biozone utilized in the Karoo Basin of South Africa. It is equivalent to the Burgersdorp Formation, the youngest lithostratigraphic formation in the Beaufort Group, which is part of the fossiliferous and geologically important Karoo Supergroup. The Cynognathus Assemblage Zone is the youngest of the eight biozones found in the Beaufort Group, and is considered to be late Early Triassic (Olenekian) to early Middle Triassic (Anisian) in age. The name of the biozone refers to Cynognathus crateronotus, a large and carnivorous cynodont therapsid which occurs throughout the entire biozone.
Cynosaurus is an extinct genus of cynodonts. Remains have been found from the Dicynodon Assemblage Zone in South Africa. Cynosaurus was first described by Richard Owen in 1876 as Cynosuchus suppostus. Cynosaurus has been found in the late Permian period. Cyno- is derived from the Greek word kyon for dog and –sauros in Greek meaning lizard.
Progalesaurus is an extinct genus of galesaurid cynodont from the early Triassic. Progalesaurus is known from a single fossil of the species Progalesaurus lootsbergensis, found in the Lystrosaurus Assemblage Zone of the Balfour Formation. Close relatives of Progalesaurus, other galesaurids, include Galesaurus and Cynosaurus. Galesaurids appeared just before the Permian-Triassic extinction event, and disappeared from the fossil record in the Middle-Triassic.
Platycraniellus is an extinct genus of carnivorous cynodonts from the Early Triassic. It is known from the Lystrosaurus Assemblage Zone of the Normandien Formation in South Africa. P. elegans is the only species in this genus based on the holotype specimen from the Ditsong National Museum of Natural History in Pretoria, South Africa. Due to limited fossil records for study, Platycraniellus has only been briefly described a handful of times.
Trirachodon is an extinct genus of cynodonts. Fossils have been found in the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Omingonde Formation of Namibia, dating back to the Early and Middle Triassic.
Pachygenelus is an extinct genus of tritheledontid cynodonts. Fossils have been found from the Karoo basin in South Africa and date back to the Early Jurassic.
Lumkuia is an extinct genus of cynodont, fossils of which have been found in the Cynognathus Assemblage Zone of the Beaufort Group in the South African Karoo Basin that date back to the early Middle Triassic. It contains a single species, Lumkuia fuzzi, which was named in 2001 on the basis of the holotype specimen BP/1/2669, which can now be found at the Bernard Price Institute in Johannesburg, South Africa. The genus has been placed in its own family, Lumkuiidae. Lumkuia is not as common as other cynodonts from the same locality such as Diademodon and Trirachodon.
Diademodon is an extinct genus of cynodonts. It was about 2 metres (6.6 ft) long.
Langbergia is an extinct genus of trirachodontid cynodont from the Early Triassic of South Africa. The type and only species L. modisei was named in 2006 after the farm where the holotype was found, Langberg 566. Langbergia was found in the Burgersdorp Formation in the Beaufort Group, a part of the Cynognathus Assemblage Zone. The closely related trirachodontids Trirachodon and Cricodon were found in the same area.
Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. They are thought to have been herbivorous or omnivorous. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil, eastern North America, Europe, China, and Antarctica.
Titanogomphodon is an extinct genus of diademodontid cynodonts from the Middle Triassic Omingonde Formation of Namibia. It is known from a single partial skull that was described in 1973 from the Omingonde Formation. The type and only species is Titanogomphodon crassus. At about 40 centimetres (16 in), the skull of Titanogomphodon was significantly larger than that of its closest relative, Diademodon. Its teeth are similar to those of another group of cynodonts called Traversodontidae, but the similarities are likely the result of convergent evolution. Aside from its larger size, Titanogomphodon differs from Diademodon in having a bony projection on the postorbital bar behind the eye socket.
Cricodon is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa. A. W. Crompton named Cricodon based on the ring-like arrangement of the cuspules on the crown of a typical postcanine tooth. The epithet of the type species, C. metabolus, indicates the change in structure of certain postcanines resulting from replacement.
Abdalodon is an extinct genus of late Permian cynodonts, known by its only species A. diastematicus.Abdalodon together with the genus Charassognathus, form the clade Charassognathidae. This clade represents the earliest known cynodonts, and is the first known radiation of Permian cynodonts.
Impidens is an extinct genus of large omnivorous cynodont from the Triassic of South Africa and Antarctica. Its type and only species is Impidens hancoxi. Impidens inhabited high-latitude environments of southern Gondwana during the Middle Triassic, where it was probably the apex predator.