Trirachodontidae

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Trirachodontids
Temporal range: Early - Middle Triassic, 251–237  Ma
Trirachodon.jpg
Life restoration of Trirachodon berryi in a burrow
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Cynodontia
Clade: Neogomphodontia
Family: Trirachodontidae
Crompton, 1955
Genera

Trirachodontidae is an extinct, possibly paraphyletic [1] family of cynognathian cynodonts from the Triassic of China and southern Africa. Trirachodontids appeared during the Early Triassic soon after the Permian-Triassic extinction event and quickly spread over a wide geographic area in a comparatively brief amount of time from 250 to 237 million years ago.

Trirachodontids have wide skulls and short, narrow snouts. Two large holes called temporal fenestrae run along the back of the head and have a uniformly large width. Trirachodontids also have two large canine teeth and smaller cusped postcanines. Most of the features that distinguish trirachodonts from other cynodonts are found in their dentition. [2]

Trirachodontids lived in semi-arid environments with seasonal rainfall. The bone structure of trirachodontids suggests that they grew quickly in seasons with high rainfall and slowly in less favorable seasons. [3] One trirachodontid, Trirachodon , has been found in association with complex burrow systems. These burrows were probably used by many individuals to hide from predators, raise young, or stay warm.

Two subfamilies of trirachodontids are recognized: Trirachodontinae from Africa and Sinognathinae from China. Below is a cladogram from Gao et al. (2010) showing the phylogenetic relationships of trirachodontids: [2]

Cynognathia  

Cynognathus

  Gomphodontia  

Diademodon

 Trirachodontidae 
 Trirachodontinae 

Trirachodon

Langbergia

Cricodon

 Sinognathinae 

Beishanodon

Sinognathus

Traversodontidae

The cladogram above shows a monophyletic Trirachodontidae, but multiple studies have found the family to be paraphyletic with respect to the family Traversodontidae. [1] [4] Below is a cladogram from Hendrickx et al. (2020), who also recovered the members of the subfamily Trirachodontinae as a basal polytomy within the clade Neogomphodontia: [1]

Cynognathia  

Cynognathus

  Gomphodontia  
  Diademodontidae  

Diademodon

Titanogomphodon

  Neogomphodontia  

Langbergia

Trirachodon

  Sinognathinae  

Beishanodon

Cricodon

Sinognathus

Traversodontidae

Trirachodontidae

Related Research Articles

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Probainognathidae is an extinct family of insectivorous cynodonts which lived in what is now South America during the Middle to Late Triassic. The family was established by Alfred Romer in 1973 and includes two genera, Probainognathus from the Chañares Formation of Argentina and Bonacynodon from the Dinodontosaurus Assemblage Zone of Brazil. Probainognathids were closely related to the clade Prozostrodontia, which includes mammals and their close relatives.

<i>Thrinaxodon</i> Extinct genus of cynodonts of Early Triassic South Africa

Thrinaxodon is an extinct genus of cynodonts, most commonly regarded by its species T. liorhinus which lived in what are now South Africa and Antarctica during the Early Triassic. Thrinaxodon lived just after the Permian–Triassic mass extinction event, its survival during the extinction may have been due to its burrowing habits.

<span class="mw-page-title-main">Cynognathia</span> Clade of cynodonts

Cynognathia is one of two major clades of cynodonts, the other being Probainognathia. Cynognathians included the large carnivorous genus Cynognathus and the herbivorous traversodontids. Cynognathians can be identified by several synapomorphies including a very deep zygomatic arch that extends above the middle of the orbit.

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<span class="mw-page-title-main">Traversodontidae</span> Extinct family of cynodonts

Traversodontidae is an extinct family of herbivorous cynodonts. Traversodonts were primarily Gondwanan, with many species known from Africa and South America. Recently, traversodonts have also been found from Europe and eastern North America. Traversodonts first appeared in the Middle Triassic and diversified in the Late Triassic before going extinct at the end of the epoch. The family Traversodontidae was erected by Friedrich von Huene in 1936 for cynodonts first found in São Pedro do Sul in Paleorrota, Brazil.

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<i>Prozostrodon</i> Extinct genus of cynodonts

Prozostrodon is an extinct genus of probainognathian cynodonts that was closely related to the ancestors of mammals. The remains were found in Brazil and are dated to the Carnian age of the Late Triassic. The holotype has a skull length of 6.7 centimetres (2.6 in), indicating that the whole animal may have been the size of a cat. The teeth were typical of advanced cynodonts, and the animal was probably a carnivore hunting reptiles and other small prey.

<i>Brasilodon</i> Extinct genus of mammaliamorphs

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<i>Trirachodon</i> Extinct genus of cynodonts

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<span class="mw-page-title-main">Prozostrodontia</span> Clade of cynodonts

Prozostrodontia is a clade of cynodonts including mammaliaforms and their closest relatives such as Tritheledontidae and Tritylodontidae. It was erected as a node-based taxon by Liu and Olsen (2010) and defined as the least inclusive clade containing Prozostrodon brasiliensis, Tritylodon langaevus, Pachygenelus monus, and Mus musculus. Prozostrodontia is diagnosed by several characters, including:

<span class="mw-page-title-main">Gomphodontia</span> Clade of cynodonts

Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil, eastern North America, Europe, China, and Antarctica.

<i>Cricodon</i> Extinct genus of cynodonts

Cricodon is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa. A. W. Crompton named Cricodon based on the ring-like arrangement of the cuspules on the crown of a typical postcanine tooth. The epithet of the type species, C. metabolus, indicates the change in structure of certain postcanines resulting from replacement.

<i>Abdalodon</i> Extinct genus of cynodonts

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<i>Etjoia</i> Extinct genus of cynodonts

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Arctotraversodontinae is a subfamily of Late Triassic cynodonts belonging to the family Traversodontidae. Members of the subfamily include Arctotraversodon, Boreogomphodon and Plinthogomphodon from North America, and Habayia, Maubeugia, Microscalenodon and Rosieria from Europe.

Impidens is an extinct genus of large omnivorous cynodont from the Triassic of South Africa and Antarctica. Its type and only species is Impidens hancoxi. Impidens inhabited high-latitude environments of southern Gondwana during the Middle Triassic, where it was probably the apex predator.

References

  1. 1 2 3 Hendrickx, C.; Gaetano, L. C.; Choiniere, J. N.; Mocke, H.; Abdala, F. (2020). "A new traversodontid cynodont with a peculiar postcanine dentition from the Middle/Late Triassic of Namibia and dental evolution in basal gomphodonts". Journal of Systematic Palaeontology. 18 (20): 1669–1706. doi:10.1080/14772019.2020.1804470. S2CID   221838726.
  2. 1 2 Gao, K.-Q.; Fox, R.C.; Zhou, C.-F.; Li, D.-Q. (2010). "A new nonmammalian eucynodont (Synapsida: Therapsida) from the Triassic of Northern Gansu Province, China, and its biostratigraphic and biogeographic implications" (PDF). American Museum Novitates (3685): 1–25. doi:10.1206/649.1. hdl:2246/6069. S2CID   85789838.
  3. Botha, J.; Chinsamy, A. (2004). "Growth and life habits of the Triassic cynodont Trirachodon, inferred from bone histology" (PDF). Acta Palaeontologica Polonica. 49 (9): 619–627.
  4. Liu, J.; Olsen, P. (2010). "The Phylogenetic Relationships of Eucynodontia (Amniota: Synapsida)". Journal of Mammalian Evolution. 17 (3): 151. doi:10.1007/s10914-010-9136-8. S2CID   40871206.