Exaeretodon

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Exaeretodon
Temporal range: Ladinian-Norian
~235–205.6  Ma
Cynodont.jpg
Exhibit at Mori Tower
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Cynodontia
Family: Traversodontidae
Subfamily: Gomphodontosuchinae
Genus: Exaeretodon
Cabrera, 1943
Species
  • E. argentinusCabrera, 1943 (type) [originally Belesodon argentinus]
  • ?E. major Huene, 1936 [originally Traversodon ]
  • E. riograndensisAbdala et al., 2002
  • ?E. statisticaeChatterjee, 1982
Synonyms
  • TheropsisCabrera, 1943
  • Ischignathus Bonaparte, 1963

Exaeretodon is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. E. argentinus is from the Carnian-age (Late Triassic) Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina. E. major and E. riograndensis are from the Carnian-age portion of the Santa Maria Formation of the Paraná Basin in southeastern Brazil. [1] E. statisticae is from the Carnian-age Lower Maleri Formation of India.

Contents

Description

Restoration Exaeretodon BW.jpg
Restoration
Skull of Exaeretodon at the Royal Ontario Museum August 1, 2012 - Skull of Exaeretodon on Display at the Royal Ontario Museum (MC2 4483).jpg
Skull of Exaeretodon at the Royal Ontario Museum

This genus was an herbivore up to 1.8 meters (5.9 feet) long, with a specialized grinding action when feeding.

An analysis of the size of the bones of calves collected in Paleorrota concluded that the mother Exaeretodon had one or two calves, for one pregnancy.[ citation needed ]

Taxonomy

Exaeretodon is a gomphodontosuchine cynodont. [2] When he first named the species, Argentine paleontologist José Bonaparte mentioned several features that distinguish it from all other traversodontids. The tooth rows of the upper jaw are more parallel to each other in Ischignathus sudamericanus than they are in Exaeretodon, and they are also inset closer to the inside of the mouth. There are also more postcanine teeth oriented toward middle of the subtemporal fenestrae (two holes in the bottom of the skull) and ascending rama of the dentary (projections of the lower jaw that extend up to the skull). The ascending rama are also wider and taller in the I. sudamericanus specimen. The orbit or eye socket is longer than that of Exaeretodon, as are the palatine bones. [3]

Despite the differences, a 2007 study concluded that the I. sudamericanus skull represents the same species as E. argentinus. Using allometry, paleontologist Jun Liu found I. sudamericanus to be the largest known example of a growth series in E. argentinus. Thus, Ischignathus is now regarded as a junior synonym of Exaeretodon. As the animal grew, the proportions of bones changed. These differing proportions were initially seen as species-distinguishing characters, but are now regarded as natural ontogenic variation. [3]

Proexaeretodon vince was considered a synonym of Exaeretodon argentinus but was accepted as a valid taxon in 2023. [2]

Phylogeny

Exaeretodon in a cladogram after Kammerer et al. (2012): [4]

Cynognathia  

See also

Related Research Articles

<i>Probainognathus</i> Extinct genus of cynodonts

Probainognathus meaning “progressive jaw” is an extinct genus of cynodonts that lived around 235 to 221.5 million years ago, during the Late Triassic in what is now Argentina. Together with the genus Bonacynodon from Brazil, Probainognathus forms the family Probainognathidae. Probainognathus was a relatively small, carnivorous or insectivorous cynodont. Like all cynodonts, it was a relative of mammals, and it possessed several mammal-like features. Like some other cynodonts, Probainognathus had a double jaw joint, which not only included the quadrate and articular bones like in more basal synapsids, but also the squamosal and surangular bones. A joint between the dentary and squamosal bones, as seen in modern mammals, was however absent in Probainognathus.

<i>Chiniquodon</i> Extinct genus of cynodonts

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<i>Hyperodapedon</i> Extinct genus of reptiles

Hyperodapedon is an extinct genus of rhynchosaur reptiles which lived during Late Triassic period. Like other rhynchosaurs, it was an heavily built archosauromorph, distantly related to archosaurs such as crocodilians and dinosaurs. Hyperodapedon in particular was part of the subfamily Hyperodapedontinae, a specialized rhynchosaurian subgroup with broad skulls, beaked snouts, and crushing tooth plates on the roof of the mouth.

<i>Massetognathus</i> Extinct genus of cynodonts

Massetognathus is an extinct genus of plant-eating traversodontid cynodonts. They lived during the Triassic Period about 235 million years ago, and are known from the Chañares Formation in Argentina and the Santa Maria Formation in Brazil.

Tritylodontidae is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy and details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous.

<i>Prozostrodon</i> Extinct genus of cynodonts

Prozostrodon is an extinct genus of probainognathian cynodonts that was closely related to mammals. The remains were found in Brazil and are dated to the Carnian age of the Late Triassic. The holotype has an estimated skull length of 6.7 centimetres (2.6 in), indicating that the whole animal may have been the size of a cat. The teeth were typical of advanced cynodonts, and the animal was probably a carnivore hunting reptiles and other small prey.

<span class="mw-page-title-main">Santa Maria Formation</span> Geologic formation in Brazil

The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age, and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, including the herrerasaurid Staurikosaurus, the basal sauropodomorphs Buriolestes and Saturnalia, and the lagerpetid Ixalerpeton. The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied.

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<span class="mw-page-title-main">Ischigualasto Formation</span> Geological formation in Argentina

The Ischigualasto Formation is a Late Triassic geological formation in the Ischigualasto-Villa Unión Basin of southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation dates to the late Carnian and early Norian stages of the Late Triassic, according to radiometric dating of ash beds.

<i>Proterochampsa</i> Extinct genus of reptiles

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<i>Panphagia</i> Extinct genus of dinosaurs

Panphagia is a genus of sauropodomorph dinosaur described in 2009. It lived around 231 million years ago, during the Carnian age of the Late Triassic period in what is now northwestern Argentina. Fossils of the genus were found in the La Peña Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin. The name Panphagia comes from the Greek words pan, meaning "all", and phagein, meaning "to eat", in reference to its inferred omnivorous diet. Panphagia is one of the earliest known dinosaurs, and is an important find which may mark the transition of diet in early sauropodomorph dinosaurs.

<i>Lumkuia</i> Extinct genus of cynodonts

Lumkuia is an extinct genus of cynodont, fossils of which have been found in the Cynognathus Assemblage Zone of the Beaufort Group in the South African Karoo Basin that date back to the early Middle Triassic. It contains a single species, Lumkuia fuzzi, which was named in 2001 on the basis of the holotype specimen BP/1/2669, which can now be found at the Bernard Price Institute in Johannesburg, South Africa. The genus has been placed in its own family, Lumkuiidae. Lumkuia is not as common as other cynodonts from the same locality such as Diademodon and Trirachodon.

<i>Sanjuansaurus</i> Extinct genus of dinosaurs

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<i>Andescynodon</i> Extinct genus of cynodonts

Andescynodon is a genus of traversodontid cynodonts from the Middle Triassic of Argentina. Fossils are known from the Cerro de las Cabras and Cacheutá Formations. Andescynodon is one of the most basal traversodontids. Another traversodontid called Rusconiodon has also been identified from the Cerro de las Cabras Formation but is now considered a junior synonym of Andescynodon.

Scalenodon is an extinct genus of traversodontid cynodonts from the Middle Triassic of Africa and possibly Russia. The type species S. angustifrons was named in 1946 and several other species were named in the following years. Most of the species from Africa are now thought to belong to different genera than Scalenodon.

<span class="mw-page-title-main">Gomphodontosuchinae</span> Subfamily of traversodontid cynodonts

Gomphodontosuchinae is a subfamily of Triassic traversodontid cynodonts. It includes the genera Gomphodontosuchus, Exaeretodon, Santagnathus, Siriusgnathus, Menadon, Proexaeretodon,Protuberum, Ruberodon and Scalenodontoides.

Mandagomphodon is an extinct genus of traversodontid cynodonts from the Middle Triassic Lifua Member of the Manda Beds of Ruhuhu Valley, Tanzania. The type species Mandagomphodon hirschsoni was named by Crompton in 1972 as a species referable to Scalenodon. Later studies, including a 2003 phylogenetic analysis of traversodontid relationships, did not find the species of Scalenodon from the Manda Formation to form a single clade, meaning that many were not referable to the genus. The study suggested that S. hirschsoni had more in common with other traversodontids like Luangwa. S. attridgei was viewed as a possible synonym of S. charigi, which was also found to be only distantly related to S. angustifrons. Therefore, a new generic name Mandagomphodon was erected for S. hirschsoni by James A. Hopson in 2013.

<i>Dynamosuchus</i> Extinct genus of reptiles

Dynamosuchus is an extinct genus of pseudosuchian archosaurs from the family Ornithosuchidae. It is known from a single species, Dynamosuchus collisensis, which is based on a partial skeleton from the Santa Maria Formation of Brazil. Dynamosuchus is considered a close relative of Venaticosuchus, which is known from the Ischigualasto Formation of Argentina. Ornithosuchids are one of many groups which lived in the Santa Maria and Ischigualasto Formations, which formed at approximately the same time and were ecologically similar. As a large scavenging reptile, Dynamosuchus helps to illuminate the trophic structure of the Santa Maria Formation. It also supports the hypothesis that ornithosuchids had diversified throughout South America by the start of the Carnian, and were not originally endemic to the Ischigualasto-Villa Unión Basin.

The Río Seco de la Quebrada Formation is a Triassic geological formation in Mendoza Province, Argentina. Fossils of cynodonts such as Cynognathus, Diademodon, and Pascualgnathus have been found in this formation, along with dicynodonts such as Vinceria and Acratophorus. Based on biostratigraphy, the Río Seco de la Quebrada Formation is considered to have formed during the Anisian stage, as it shares fauna with the upper subzones of the Cynognathus Assemblage Zone in South Africa. However, radiometric dating controversially argues that it was deposited during the early Carnian stage, 10 million years younger than expected otherwise.

<i>Pseudotherium</i> Extinct genus of cynodonts

Pseudotherium is an extinct genus of prozostrodontian cynodonts from the Late Triassic of Argentina. It contains one species, P. argentinus, which was first described in 2019 from remains found in the La Peña Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin.

References

  1. Langer, Max C.; Ramezani, Jahandar; Da Rosa, Átila A.S. (May 2018). "U-Pb age constraints on dinosaur rise from south Brazil". Gondwana Research. 57: 133–140. Bibcode:2018GondR..57..133L. doi:10.1016/j.gr.2018.01.005.
  2. 1 2 Schmitt, Maurício Rodrigo; Martinelli, Agustín Guillermo; Kaiuca, João Felipe Leal; Schultz, Cesar Leandro & Soares, Marina Bento (2023). "Old fossil findings in the Upper Triassic rocks of southern Brazil improve diversity of traversodontid cynodonts (Therapsida, Cynodontia)". The Anatomical Record. 307 (4): 1474–1514. doi:10.1002/ar.25244. PMID   37246488. S2CID   258960737.
  3. 1 2 Liu, J. (2007). "The taxonomy of the traversodontid cynodonts Exaeretodon and Ischignathus" (PDF). Revista Brasileira de Paleontologia. 10 (2): 133–136. doi: 10.4072/rbp.2007.2.07 .
  4. Kammerer, C. F.; Flynn, J. J.; Ranivoharimanana, L.; Wyss, A. R. (2012). "Ontogeny in the Malagasy Traversodontid Dadadon isaloi and a Reconsideration of its Phylogenetic Relationships". Fieldiana Life and Earth Sciences. 5: 112–125. doi:10.3158/2158-5520-5.1.112. S2CID   84651546.