Gornogomphodon Temporal range: Carnian ~ | |
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Gornogomphodon caffii fossil teeth at the Museo di Scienze Naturali Enrico Caffi, Italy | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Clade: | Cynodontia |
Clade: | † Gomphodontia |
Genus: | † Gornogomphodon Renesto & Lucas, 2009 |
Species: | †G. caffii |
Binomial name | |
†Gornogomphodon caffii Renesto & Lucas, 2009 | |
Gornogomphodon is an extinct genus of cynodonts from the Gorno Formation of Bergamo, Italy. It existed during the Middle Carnian age of the Late Triassic (around 216.5 to 228.0 million years ago). It contains only one species: Gornogomphodon caffi. [1]
Gornogomphodon is known only from a fossil jaw fragment with three transversely elongate teeth preserved. The scant fossil material and the locality they were collected from (the Gorno Formation were once shallow lagoons) lead to initial uncertainty over the nature of the fossil. It was proposed to be the remains of a pycnodont fish but further studies eventually made this hypothesis unlikely. The overall morphology of the teeth suggests they are the upper postcanines of the right maxilla of a gomphodont (herbivorous cynodonts), but they also possess unique characteristics that make their taxonomic assignment difficult. [1]
Gornogomphodon was named after the Gorno Formation (literally 'Gorno gomphodont'). The specific name for its only species Gornogomphodon caffi was named in honor of Enrico Caffi, the first director of the Museo di Scienze Naturali Enrico Caffi. [1]
The Triassic is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.
Chiniquodon is an extinct genus of carnivorous cynodonts, which lived during the Late Triassic (Carnian) in South America and Africa. Chiniquodon was closely related to the genus Aleodon, and close to the ancestry of mammals.
Massetognathus is an extinct genus of plant-eating traversodontid cynodonts. They lived during the Triassic Period about 235 million years ago, and are known from the Chañares Formation in Argentina and the Santa Maria Formation in Brazil.
Exaeretodon is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. E. argentinus is from the Carnian-age Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina. E. major and E. riograndensis are from the Carnian-age portion of the Santa Maria Formation of the Paraná Basin in southeastern Brazil. E. statisticae is from the Carnian-age Lower Maleri Formation of India.
The Chañares Formation is a Carnian-age geologic formation of the Ischigualasto-Villa Unión Basin, located in La Rioja Province, Argentina. It is characterized by drab-colored fine-grained volcaniclastic claystones, siltstones, and sandstones which were deposited in a fluvial to lacustrine environment. The formation is most prominently exposed within Talampaya National Park, a UNESCO World Heritage Site within La Rioja Province.
Trirachodon is an extinct genus of cynodonts. Fossils have been found in the Cynognathus Assemblage Zone of the Beaufort Group in South Africa and the Omingonde Formation of Namibia, dating back to the Early and Middle Triassic.
Lumkuia is an extinct genus of cynodonts, fossils of which have been found in the Cynognathus Assemblage Zone of the Beaufort Group in the South African Karoo Basin that date back to the early Middle Triassic. It contains a single species, Lumkuia fuzzi, which was named in 2001 on the basis of the holotype specimen BP/1/2669, which can now be found at the Bernard Price Institute in Johannesburg, South Africa. The genus has been placed in its own family, Lumkuiidae. Lumkuia is not as common as other cynodonts from the same locality such as Diademodon and Trirachodon.
Diademodon is an extinct genus of cynodonts. It was about 2 metres (6.6 ft) long.
Langbergia is an extinct genus of trirachodontid cynodont from the Early Triassic of South Africa. The type and only species L. modisei was named in 2006 after the farm where the holotype was found, Langberg 566. Langbergia was found in the Burgersdorp Formation in the Beaufort Group, a part of the Cynognathus Assemblage Zone. The closely related trirachodontids Trirachodon and Cricodon were found in the same area.
Nanogomphodon is an extinct genus of cynodonts which existed in Germany during the Middle Triassic period.
Neotrirachodon is an extinct genus of therapsids which existed in Russia during the Middle Triassic period. Its type and only species is Neotrirachodon expectatus.
Andescynodon is a genus of traversodontid cynodonts from the Middle Triassic of Argentina. Fossils are known from the Cerro de las Cabras and Cacheutá Formations. Andescynodon is one of the most basal traversodontids. Another traversodontid called Rusconiodon has also been identified from the Cerro de las Cabras Formation but is now considered a junior synonym of Andescynodon.
Scalenodon is an extinct genus of traversodontid cynodonts from the Middle Triassic of Africa and possibly Russia. The type species S. angustifrons was named in 1946 and several other species were named in the following years. Most of the species from Africa are now thought to belong to different genera than Scalenodon.
Gomphodontia is a clade of cynognathian cynodonts that includes the families Diademodontidae, Trirachodontidae, and Traversodontidae. Gomphodonts are distinguished by wide and closely spaced molar-like postcanine teeth, which are convergent with those of mammals. Other distinguishing characteristics of gomphodonts include deep zygomatic arches, upper postcanines with three or more cusps spanning their widths and lower postcanines with two cusps spanning their widths. They are thought to have been herbivorous or omnivorous. Gomphodonts first appeared in the Early Triassic and became extinct at the end of the Late Triassic. Fossils are known from southern Africa, Argentina and southern Brazil, eastern North America, Europe, China, and Antarctica.
Titanogomphodon is an extinct genus of diademodontid cynodonts from the Middle Triassic Omingonde Formation of Namibia. It is known from a single partial skull that was described in 1973 from the Omingonde Formation. The type and only species is Titanogomphodon crassus. At about 40 centimetres (16 in), the skull of Titanogomphodon was significantly larger than that of its closest relative, Diademodon. Its teeth are similar to those of another group of cynodonts called Traversodontidae, but the similarities are likely the result of convergent evolution. Aside from its larger size, Titanogomphodon differs from Diademodon in having a bony projection on the postorbital bar behind the eye socket.
Cricodon is an extinct genus of trirachodontid cynodonts that lived during the Early Triassic and Middle Triassic periods of Africa. A. W. Crompton named Cricodon based on the ring-like arrangement of the cuspules on the crown of a typical postcanine tooth. The epithet of the type species, C. metabolus, indicates the change in structure of certain postcanines resulting from replacement.
Aleodon is an extinct genus of cynodonts that lived from the Middle to the Late Triassic. Relatively few analyses have been conducted to identify the phylogenetic placement of Aleodon, however those that have place Aleodon as a sister taxon to Chiniquodon. Two species of Aleodon are recognized: A. brachyramphus which was discovered in Tanzania, and A. cromptoni which was discovered most recently in Brazil.
Kraterokheirodon is an extinct genus of enigmatic tetrapod, that was possibly an amniote, from the Late Triassic Chinle Formation of Arizona. The type and only species is K. colberti. Although it is known only from two large teeth, their shape is so unlike those of any other animal that Kraterokheirodon cannot definitively be classified under any known group of tetrapods. Its discovery also indicates that our understanding of Late Triassic tetrapod diversity is still incomplete, with Kraterokheirodon representing an otherwise unknown lineage of large tetrapod in western North America.
Etjoia is an extinct genus of traversodontid cynodonts that lived during the Middle Triassic or Late Triassic period in southern Africa. This medium-sized omnivorous cynognathian provides important information on the dental evolution of early diverging gomphodonts and traversodontids.
Impidens is an extinct genus of large omnivorous cynodont from the Triassic of South Africa and Antarctica. Its type and only species is Impidens hancoxi. Impidens inhabited high-latitude environments of southern Gondwana during the Middle Triassic, where it was probably the apex predator.