Bienotherium Temporal range: | |
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Bienotherium yuannanese | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Clade: | Cynodontia |
Family: | † Tritylodontidae |
Genus: | † Bienotherium Young, 1940 [1] |
Species | |
Synonyms | |
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Bienotherium is an extinct genus of cynodonts from the Early Jurassic of China discovered by Bian Meinian (Mei Nien Bien). [2] Despite its size, it is closely related to Lufengia , and is the largest tritylodont from the Lufeng Formation in China. [3]
Bienotherium had four incisors, no canines, and back molar-like teeth, which it used to chew tough plant material. [4]
Bienotherium is defined as being big and robust compared to other tritylodonts, and also by exposed maxillaries in the skull, an unusually long diastema and thin zygomatic bone. [3]
A chordate is a deuterostomic bilaterial animal belonging to the phylum Chordata. All chordates possess, at some point during their larval or adult stages, five distinctive physical characteristics (synapomorphies) that distinguish them from other taxa. These five synapomorphies are a notochord, a hollow dorsal nerve cord, an endostyle or thyroid, pharyngeal slits, and a post-anal tail.
A fossil is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the fossil record. Though the fossil record is incomplete, numerous studies have demonstrated that there is enough information available to give a good understanding of the pattern of diversification of life on Earth. In addition, the record can predict and fill gaps such as the discovery of Tiktaalik in the arctic of Canada.
Paleontology, also spelled palaeontology or palæontology, is the scientific study of life that existed prior to the start of the Holocene epoch. It includes the study of fossils to classify organisms and study their interactions with each other and their environments. Paleontological observations have been documented as far back as the 5th century BC. The science became established in the 18th century as a result of Georges Cuvier's work on comparative anatomy, and developed rapidly in the 19th century. The term has been used since 1822 formed from Greek παλαιός, ὄν, and λόγος.
Agnatha is a paraphyletic infraphylum of non-gnathostome vertebrates, or jawless fish, in the phylum Chordata, subphylum Vertebrata, consisting of both living (cyclostomes) and extinct. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes.
Gnathostomata are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. Most gnathostomes have retained ancestral traits like true teeth, a stomach, and paired appendages. Other traits are elastin, a horizontal semicircular canal of the inner ear, myelin sheaths of neurons, and an adaptive immune system which has discrete lymphoid organs, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene.
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors.
Repenomamus is a genus of opossum- to badger-sized gobiconodontid mammal containing two species, Repenomamus robustus and Repenomamus giganticus. Both species are known from fossils found in China that date to the early Cretaceous period, about 125-123.2 million years ago. R. robustus is one of several Mesozoic mammals for which there is good evidence that it fed on vertebrates, including dinosaurs. Though it is not entirely clear whether or not these animals primarily hunted live dinosaurs or scavenged dead ones, evidence for the former is present in fossilized remains showcasing the results of what was most likely a predation attempt by R. robustus directed at a specimen of the dinosaur Psittacosaurus lujiatunensis. R. giganticus is among the largest mammals known from the Mesozoic era, only surpassed by Patagomaia.
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland, the Calcaire de Caen of France, the Daohugou Beds in China, the Itat Formation in Russia, the Tiouraren Formation of Niger, and the Isalo III Formation of western Madagascar.
Spencer George Lucas is an American paleontologist and stratigrapher, and curator of paleontology at the New Mexico Museum of Natural History and Science. His main areas of study are late Paleozoic, Mesozoic and early Cenozoic vertebrate fossils, stratigraphy, and continental deposits, particularly in the American Southwest. His research has taken him on field trips to northern Mexico, Costa Rica, Nicaragua, Jamaica, Kazakhstan, and Georgia, and he conducted extensive field and museum research in China in the 1980s and 1990s. He has written more than 500 scientific contributions, three books, and has co-edited 14 books.
Oligokyphus is an extinct genus of herbivorous tritylodontid cynodont known from the Late Triassic to Early Jurassic of Europe, Asia and North America.
Tritylodon is an extinct genus of tritylodonts, one of the most advanced group of cynodont therapsids. They lived in the Early Jurassic and possibly Late Triassic periods along with dinosaurs. They also shared many characteristics with mammals, and were once considered mammals because of overall skeleton construction. That was changed due to them retaining the vestigial amniote jawbones and a different skull structure. Tritylodonts are now regarded as non-mammalian synapsids.
Zhucheng is a county-level city in the southeast of Shandong province, People's Republic of China. It is under the administration of Weifang city and had at the 2010 census a population of 1,086,222 even though its built-up area is much smaller.
Peter Dodson is an American paleontologist who has published many papers and written and collaborated on books about dinosaurs. An authority on Ceratopsians, he has also authored several papers and textbooks on hadrosaurs and sauropods, and is a co-editor of The Dinosauria, widely considered the definitive scholarly reference on dinosaurs. Dodson described Avaceratops in 1986; Suuwassea in 2004, and many others, while his students have named Paralititan and Auroraceratops. He has conducted field research in Canada, the United States, India, Madagascar, Egypt, Argentina, and China. A professor of vertebrate paleontology and of veterinary anatomy at the University of Pennsylvania, Dodson has also taught courses in geology, history, history and sociology of science, and religious studies. Dodson is also a research associate at the Academy of Natural Sciences. In 2001, two former students named an ancient frog species, Nezpercius dodsoni, after him. Dodson has also been skeptical to the theory of a dinosaurian origin of birds, but more recently has come down on the side of this theory.
The history of life on Earth traces the processes by which living and extinct organisms evolved, from the earliest emergence of life to the present day. Earth formed about 4.5 billion years ago and evidence suggests that life emerged prior to 3.7 Ga. The similarities among all known present-day species indicate that they have diverged through the process of evolution from a common ancestor.
The evolution of mammalian auditory ossicles was an evolutionary process that resulted in the formation of the bones of the mammalian middle ear. These bones, or ossicles, are a defining characteristic of all mammals. The event is well-documented and important as a demonstration of transitional forms and exaptation, the re-purposing of existing structures during evolution.
Edward B. 'Ted' Daeschler is an American vertebrate paleontologist and Associate Curator and Chair of Vertebrate Biology at the Academy of Natural Sciences in Philadelphia. He is a specialist in fish paleontology, especially in the Late Devonian, and in the development of the first limbed vertebrates. He is the discoverer of the transitional fossil tetrapod Hynerpeton bassetti, and a Devonian fish-like specimen of Sauripterus taylori with fingerlike appendages, and was also part of a team of researchers that discovered the transitional fossil Tiktaalik.
John Albert Long is an Australian paleontologist who is currently Strategic Professor in Palaeontology at Flinders University in Adelaide, South Australia. He was previously the Vice President of Research and Collections at the Natural History Museum of Los Angeles County. He is also an author of popular science books. His main area of research is on the fossil fish of the Late Devonian Gogo Formation from northern Western Australia. It has yielded many important insights into fish evolution, such as Gogonasus and Materpiscis, the later specimen being crucial to our understanding of the origins of vertebrate reproduction.
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Richard Hall Tedford was Curator Emeritus in the Department of Vertebrate Paleontology of the American Museum of Natural History in New York City, having been named as curator in 1969.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
Chinese Fossil Vertebrates, p. 133