Kollikodon

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Kollikodon
Temporal range: Cenomanian, 99–96  Ma
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Kollikodon.fossil.jpg
Opalised jaw of Kollikodon (backlit)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Monotremata
Family: Kollikodontidae
Genus: Kollikodon
Flannery, Archer, Rich & Jones, 1995 [1]
Species:
K. ritchiei
Binomial name
Kollikodon ritchiei
Flannery, Archer, Rich & Jones, 1995 [1]
Synonyms
  • Hotcrossbunodon(informal)

Kollikodon is an extinct species of mammal, considered to be an early monotreme. [2] It is known only from an opalised dentary fragment, with one premolar and two molars in situ , as well as a referred maxillary fragment containing the last premolar and all four molars. The fossils were found in the Griman Creek Formation at Lightning Ridge, New South Wales, Australia. Kollikodon lived in the Late Cretaceous period, during the Cenomanian age (99–96 million years ago). Several other monotremes are known from the Griman Creek Formation, including Dharragarra , Opalios , Parvopalus , Steropodon , and Stirtodon . [3]

Contents

Etymology

Kollix is an ancient Greek word (κολλίξ) for a bread roll. The strange teeth of Kollikodon, when seen from above, resemble hot cross buns, traditionally toasted and eaten on Good Friday. Originally, Michael Archer wanted to name it "Hotcrossbunodon", but met disapproval from his associates. [4]

Description

Like Steropodon, Kollikodon was a relatively large mammal for the Mesozoic. The molars have a length of around 5.5 mm and a width of between about 4 and 6 mm. [5] Based upon these data, the potential body length could be up to a metre. [6] Assuming the accuracy of such a guess, Kollikodon would be a contender for the largest Mesozoic mammal known, along with other possible giants such as Repenomamus , Schowalteria , and Bubodens .

Aside from its size, it is difficult to say what Kollikodon looked like. It is certain that its teeth were specialised to crush food, being perhaps a shellfish-eater or herbivore. The description of the upper jaw showed that it was strongly specialised, with molars being subdivided into numerous rounded cuspules, some of which exhibit pits, possibly the result of crushing hard items. [7]

In museums

Both Kollikodon and Steropodon can be found at the Australian Museum in Sydney, along with Eric, the opalised pliosaur.

Related Research Articles

<i>Monotrematum</i> Extinct genus of monotremes

Monotrematum sudamericanum is an extinct monotreme species from the Paleocene (Peligran) Salamanca Formation in Patagonia, Argentina. It is one of only two monotremes found outside Oceania.

<span class="mw-page-title-main">Ornithorhynchidae</span> Family of monotremes

The Ornithorhynchidae are one of the two extant families in the order Monotremata, and contain the platypus and its extinct relatives. The other family is the Tachyglossidae, or echidnas.

<span class="mw-page-title-main">Prototheria</span> Subclass of mammalia

Prototheria is an obsolete subclass of mammals which includes the living Monotremata and to which a variety of extinct groups, including Morganucodonta, Docodonta, Triconodonta and Multituberculata, have also been assigned. It is today no longer considered a valid grouping, but rather a paraphyletic evolutionary grade of basal mammals and mammaliaform cynodonts.

<span class="mw-page-title-main">Theria</span> Subclass of mammals in the clade Theriiformes

Theria is a subclass of mammals amongst the Theriiformes. Theria includes the eutherians and the metatherians but excludes the egg-laying monotremes and various extinct mammals evolving prior to the common ancestor of placentals and marsupials.

<i>Steropodon</i> Extinct genus of monotremes

Steropodon is a genus of prehistoric platypus-like monotreme, or egg-laying mammal. It contains a single species, Steropodon galmani, that lived about 100.2–96.6 million years ago during the Cretaceous period, from early to middle Cenomanian. It is one of the oldest monotremes discovered, and is one of the oldest Australian mammal discoveries. Several other monotremes are known from the Griman Creek Formation, including Dharragarra, Kollikodon, Opalios, Parvopalus, and Stirtodon.

Teinolophos is a prehistoric species of monotreme, or egg-laying mammal, from the Teinolophidae. It is known from four specimens, each consisting of a partial lower jawbone collected from the Wonthaggi Formation at Flat Rocks, Victoria, Australia. It lived during the late Barremian age of the Lower Cretaceous.

Kollikodontidae is an extinct family of early monotreme mammals. Defining characteristics of kollikodontids include very specialized bunodont molars, which may indicate a diet of hard-shelled animals like mollusks in some species, and a somewhat narrow mandibular canal. The known genera have estimated body masses between 3.5–8 kilograms (7.7–17.6 lb).

<span class="mw-page-title-main">Tribosphenida</span> Infralegion of mammals

Tribosphenida is a group (infralegion) of mammals that includes the ancestor of Hypomylos, Aegialodontia and Theria. It belongs to the group Zatheria. The current definition of Tribosphenida is more or less synonymous with Boreosphenida.

<span class="mw-page-title-main">Australosphenida</span> Subclass of mammals

The Australosphenida are a clade of mammals, containing mammals with tribosphenic molars, known from the Jurassic to Mid-Cretaceous of Gondwana. Although they have often been suggested to have acquired tribosphenic molars independently from those of Tribosphenida, this has been disputed. Fossils of australosphenidans have been found from the Jurassic of Madagascar and Argentina, and Cretaceous of Australia and Argentina. Monotremes have also been considered a part of this group in its original definition and in many subsequent studies, but its relationship with the relationship with other members has been disputed by some scholars.

<i>Megalibgwilia</i> Extinct genus of monotremes

Megalibgwilia is a genus of echidna known only from Australian fossils that incorporates the oldest-known echidna species. The genus ranged from the Miocene until the late Pleistocene, becoming extinct about 50,000 years ago. Megalibgwilia species were more widespread in warmer and moist climates. Their extinction can be attributed to increasing aridification in Southern Australia.

The Griman Creek Formation is a geological formation in northern New South Wales and southern Queensland, Australia whose strata date back to the Albian-Cenomanian stages of the mid-Cretaceous. It is most notable being a major source of opal, found near the town of Lightning Ridge, New South Wales. Alongside the opal opalised fossils are also found, including those of dinosaurs and primitive monotremes.

<span class="mw-page-title-main">Monotreme</span> Order of egg-laying mammals

Monotremes are mammals of the order Monotremata. They are the only known group of living mammals that lay eggs, rather than bearing live young. The extant monotreme species are the platypus and the four species of echidnas. Monotremes are typified by structural differences in their brains, jaws, digestive tract, reproductive tract, and other body parts, compared to the more common mammalian types. Although they are different from almost all mammals in that they lay eggs, like all mammals, the female monotremes nurse their young with milk.

<span class="mw-page-title-main">Yinotheria</span> Subclass of mammals

Yinotheria is a proposed basal subclass clade of crown mammals uniting the Shuotheriidae, an extinct group of mammals from the Jurassic of Eurasia, with Australosphenida, a group of mammals known from the Jurassic to Cretaceous of Gondwana, which possibly include living monotremes. Today, there are only five surviving species of monotremes which live in Australia and New Guinea, consisting of the platypus and four species of echidna. Fossils of yinotheres have been found in Britain, China, Russia, Madagascar and Argentina. Contrary to other known crown mammals, they retained postdentary bones as shown by the presence of a postdentary trough. The extant members (monotremes) developed the mammalian middle ear independently.

Teinolophidae is an extinct family of small, carnivorous mammals that were the earliest known monotremes and were endemic to what would become Australia. Two genera are known: Teinolophos, and possibly also Stirtodon.

Opalios is an extinct genus of monotreme mammal from the Late Cretaceous (Cenomanian) Griman Creek Formation of Australia. The genus contains a single species, O. splendens, known from a fragmentary left dentary.

Dharragarra is an extinct genus of monotreme mammal from the Late Cretaceous (Cenomanian) Griman Creek Formation of Australia. The genus contains a single species, D. aurora, known from a partial left mandibular ramus. Dharragarra was likely more closely related to the living platypus than many other monotremes of the Cretaceous.

Parvopalus is an extinct genus of monotreme mammal from the Late Cretaceous (Cenomanian) Griman Creek Formation of Australia. The genus contains a single species, P. clytiei, known from a partial left dentary.

<span class="mw-page-title-main">Ornithorhynchoidea</span> Superfamily of monotreme mammals

Ornithorhynchoidea is a superfamily of mammals containing the only living monotremes, the platypus and the echidnas, as well as their closest fossil relatives, to the exclusion of more primitive fossil monotremes of uncertain affinity.

Sundrius is an extinct genus of probable monotreme mammal from the Late Cretaceous (Albian) Eumeralla Formation of Australia. The genus contains a single species, S. ziegleri, known from a broken molar.

Stirtodon is an extinct genus of monotreme mammal from the Late Cretaceous (Cenomanian) Griman Creek Formation of Australia. The genus contains a single species, S. elizabethae, known from a large isolated premolar. Stirtodon may be the largest toothed monotreme discovered. Several other monotremes are known from the Griman Creek Formation, including Dharragarra, Kollikodon, Opalios, Parvopalus, and Steropodon.

References

  1. 1 2 Flannery, Timothy F.; Archer, Michael; Rich, Thomas H. & Jones, Robert (1995). "A new family of monotremes from the Cretaceous of Australia". Nature. 377 (6548): 418–420. doi:10.1038/377418a0. S2CID   4301982.
  2. Flannery, Timothy F.; Rich, Thomas H.; Vickers-Rich, Patricia; Ziegler, Tim; Veatch, E. Grace; Helgen, Kristofer M. (2022-01-02). "A review of monotreme (Monotremata) evolution". Alcheringa: An Australasian Journal of Palaeontology. 46 (1): 3–20. doi: 10.1080/03115518.2022.2025900 . ISSN   0311-5518.
  3. Flannery, Timothy F.; McCurry, Matthew R.; Rich, Thomas H.; Vickers-Rich, Patricia; Smith, Elizabeth T.; Helgen, Kristofer M. (2024-05-26). "A diverse assemblage of monotremes (Monotremata) from the Cenomanian Lightning Ridge fauna of New South Wales, Australia". Alcheringa: An Australasian Journal of Palaeontology : 1–19. doi: 10.1080/03115518.2024.2348753 . ISSN   0311-5518.
  4. Long, John A.; Archer, Michael; Flannery, Timothy & Hand, Suzanne (2002). Prehistoric mammals of Australia and New Guinea: One hundred million years of evolution. Baltimore, Maryland: Johns Hopkins University Press. p. 39. ISBN   978-0801872235.
  5. Clemens, William A.; Wilson, Gregory P. & Molnar, Ralph E. (2003). "An enigmatic (Synapsid?) tooth from the Early Cretaceous of New South Wales, Australia" (PDF). Journal of Vertebrate Paleontology. 23 (1): 232–237. doi:10.1671/0272-4634(2003)23[232:AESTFT]2.0.CO;2. S2CID   131689322.
  6. Weil, Anne (2005). "Mammalian palaeobiology: Living large in the Cretaceous". Nature. 433 (7022) (published 2005-01-12): 116–117. doi:10.1038/433116b. PMID   15650725. S2CID   52869101.
  7. Pian, Rebecca; Archer, Michael; Hand, Suzanne J.; Beck, Robin M.D. & Cody, Andrew (2016). "The upper dentition and relationships of the enigmatic Australian Cretaceous mammal Kollikodon ritchiei" (PDF). Memoirs of Museum Victoria. 74: 97–105. doi: 10.24199/j.mmv.2016.74.10 . ISSN   1447-2546.[ permanent dead link ]