Tribosphenida is a group (infralegion) of mammals that includes the ancestor of Hypomylos, Aegialodontia and Theria. It belongs to the group Zatheria. The current definition of Tribosphenida is more or less synonymous with Boreosphenida.
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland, the Calcaire de Caen of France, the Daohugou Beds in China, the Itat Formation in Russia, the Tiouraren Formation of Niger, and the Isalo III Formation of western Madagascar.
Echinodon is a genus of heterodontosaurid dinosaur that lived during the earliest Cretaceous of southern England and possibly western France in the Berriasian epoch. The first specimens were jaw bones named Echinodon becklesii by Sir Richard Owen in 1861, and since their original description only additional teeth have been discovered. The specific name honours collector Samuel Beckles who discovered the material of Echinodon and many other taxa from across England, while the genus name translates as "prickly tooth" in reference to the dental anatomy of the taxon.
Docodonta is an order of extinct Mesozoic mammaliaforms. They were among the most common mammaliaforms of their time, persisting from the Middle Jurassic to the Early Cretaceous across the continent of Laurasia. They are distinguished from other early mammaliaforms by their relatively complex molar teeth. Docodont teeth have been described as "pseudotribosphenic": a cusp on the inner half of the upper molar grinds into a basin on the front half of the lower molar, like a mortar-and-pestle. This is a case of convergent evolution with the tribosphenic teeth of therian mammals. There is much uncertainty for how docodont teeth developed from their simpler ancestors. Their closest relatives may have been certain Triassic "symmetrodonts", namely Woutersia, Delsatia, and Tikitherium.
The family Peramuridae is a family of mammals that lived in the Late Jurassic and Early Cretaceous. They are considered to be advanced cladotherians closely related to therian mammals as part of Zatheria.
Dryolestida is an extinct order of mammals, known from the Jurassic and Cretaceous. They are considered basal members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they developed a fully mammalian jaw and also had the three middle ear bones. Most members of the group, as with most Mesozoic mammals, are only known from fragmentary tooth and jaw remains.
Morganucodonta is an extinct order of basal Mammaliaformes, a group including crown-group mammals (Mammalia) and their close relatives. Their remains have been found in Southern Africa, Western Europe, North America, India and China. The morganucodontans were probably insectivorous and nocturnal, though like eutriconodonts some species attained large sizes and were carnivorous. Nocturnality is believed to have evolved in the earliest mammals in the Triassic as a specialisation that allowed them to exploit a safer, night-time niche, while most larger predators were likely to have been active during the day.
Dryolestidae is an extinct family of Mesozoic mammals, known from the Middle Jurassic to the Early Cretaceous of the North Hemisphere. The oldest known member, Anthracolestes, is known from the Middle Jurassic Itat Formation of Western Siberia, but most other representatives are known from the Late Jurassic of North America and the Late Jurassic and Early Cretaceous of Europe. Most members are only known from isolated teeth and jaw fragments. Like many other groups of early mammals, they are thought to have been insectivores. They are generally classified in Cladotheria, meaning that they are considered to be more closely related to marsupials and placentals than to monotremes. They are placed as part of the broader Dryolestida, which also includes the Paurodontidae, and also sometimes the South American-Antarctic Meridiolestida, which are often considered unrelated cladotherians. Dryolestidae taxon is not based on a phylogenetic definition, but instead on the possession of unequal roots for the molars of the lower jaw. Additionally, the clade is distinguished by hypsodonty in lower molars, and uneven labio-lingual height for the alveolar borders of the dentary.
Shuotherium is a fossil mammaliaform known from Middle-Late Jurassic of the Forest Marble Formation of England, and the Shaximiao Formation of Sichuan, China.
Cladotheria is a clade of mammals. It contains modern therian mammals and several extinct groups, such as the "dryolestoids", amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to Mus musculus than to the "symmetrodont" Spalacotherium tricuspidens.
Duriavenator is a genus of theropod dinosaur that lived in what is now England during the Middle Jurassic, about 168 million years ago. In 1882, upper and lower jaw bones of a dinosaur were collected near Sherborne in Dorset, and Richard Owen considered the fossils to belong to the species Megalosaurus bucklandii, the first named non-bird dinosaur. By 1964, the specimen was recognised as belonging to a different species, and in 1974 it was described as a new species of Megalosaurus, M. hesperis; the specific name means 'the West' or 'western'. Later researchers questioned whether the species belonged to Megalosaurus, in which many fragmentary theropods from around the world had historically been placed. After examining the taxonomic issues surrounding Megalosaurus, Roger B. J. Benson moved M. hesperis to its own genus in 2008, Duriavenator; this name means "Dorset hunter".
Brancatherulum is an extinct genus of Late Jurassic mammal from the Tendaguru Formation of Lindi Region of Tanzania. It is based on a single toothless dentary 21 mm in length. It is currently considered either a stem-zatherian or dryolestidan.
Donodon is an extinct genus of mammal from the Ksar Metlili Formation of Talssint, Morocco, which has been dated to the Late Jurassic to Early Cretaceous epochs. The type species D. perscriptoris was described in 1991 by the palaeontologist Denise Sigogneau-Russell. A second species, D. minor, was named in 2022. Donodon was a member of Cladotheria, a group that includes therian mammals and some of their closest relatives. It differed from dryolestids in having upper molars that were not compressed mesiodistally. Some studies have suggested that it was closely related to various South American cladotherians in the clade Meridiolestida, with specific similarities to Mesungulatum, a herbivorous mesungulatid, being noted. On the other hand, a 2022 phylogenetic analysis found it to be only distantly related to meridiolestidans, and instead closer to crown group therians.
The Kilmaluag Formation is a Middle Jurassic geologic formation in Scotland. It was formerly known as the Ostracod Limestone for preserving an abundance of fossil freshwater/low salinity ostracods. Gastropods, bivalves, trace fossil burrows, and vertebrate fossil remains have also been recorded from the formation. Vertebrate fossils include fish, crocodylomorphs, mammals, small reptiles, amphibians, theropod and sauropod dinosaurs and pterosaurs.
Amphitherium is an extinct genus of stem cladotherian mammal that lived during the Middle Jurassic of England. It was one of the first Mesozoic mammals ever described. A recent phylogenetic study found it to be the sister taxon of Palaeoxonodon. It is found in the Forest Marble Formation and the Taynton Limestone Formation.
Borealestes is a genus of docodontan from the Middle Jurassic of Britain, first discovered on the Isle of Skye near the village of Elgol. It was the earliest mammaliaform from the Mesozoic found and named in Scotland. A second species and was later found in other Middle Jurassic sites in England, but is now shown to be a different genus. A new species, B. cuillinensis was named in 2021, also from Skye.
Wareolestes rex is a mammaliaform from the Middle Jurassic (Bathonian) rocks of England and Scotland. It was originally known from isolated teeth from England, before a more complete jaw with teeth was found in the Kilmaluag Formation of Skye, Scotland.
Palaeoxonodon is an extinct genus of cladotherian mammal from the Middle Jurassic of England and Scotland.
Peramus is an extinct genus of cladotherian mammal. It lived in the Late Jurassic and Early Cretaceous of Europe and North Africa.
Kouriogenys is a genus of extinct mammal from the Early Cretaceous of southern England. The type and only species was originally described as Spalacotherium minus by Richard Owen in 1871 for a dentary with teeth from the Berriasian Lulworth Formation, although it was given its own genus in 2012 by Brian Davis. The genus name is taken from the Ancient Greek "youthful" and "jaw" in reference to the replacement method of the premolars. Kouriogenys is closely related to coexisting genera Peramus and Peramuroides, and along with other genera these make up the family Peramuridae, a group of extinct zatherians.
