Stereognathus | |
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The postcanine tooth of Stereognathus ooliticus, from Owen 1857. | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Clade: | Cynodontia |
Family: | † Tritylodontidae |
Genus: | † Stereognathus Charlesworth, 1854 |
Type species | |
Stereognathus ooliticus Charlesworth, 1854 | |
Potential species | |
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Synonyms | |
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Stereognathus is an extinct genus of tritylodontid cynodonts from the Middle Jurassic of the United Kingdom. [1] There is a single named species: S. ooliticus, named after the Great Oolite deposits of England. A second species, S. hebridicus, was named after the Hebrides in Scotland, where it was found; it was synonymized with S. ooliticus in 2017. [2]
S. ooliticus was the first tritylodontid species ever found, being described by Charlesworth in 1855 [3] and later by Sir Richard Owen. [4] In 1972, S. hebridicus was named from several postcanine teeth recovered in Bathonian age deposits on the Isle of Skye in Scotland by palaeontologists R. J. G. Savage and Michael Waldman. [5] Despite being the first tritylodontid genus found and named, Stereognathus remains poorly represented, being known mainly from isolated molar teeth. There is, however, one holotype fragment of maxilla with three damaged molars, and a second fragment of maxilla with four sets of molar roots.
In 2017, Elsa Panciroli and colleagues found no points of variance between the anatomy of S. ooliticus and S. hebridicus, accounting for changes with growth, which makes S. hebridicus a junior synonym of S. ooliticus. [2] Also in 2017, Alexander Averianov and colleagues considered species of Polistodon , Xenocretosuchus , and Montirictus to be within the genus Stereognathus; they also retained S. hebridicus as a separate species. [6] However, this would mean Stereognathus persisted almost unchanged for over 60 million years across the whole of Eurasia, which would be very unusual. The issue remains unresolved.
Oligokyphus is an extinct genus of herbivorous tritylodontid cynodont known from the Late Triassic to Early Jurassic of Europe, Asia and North America.
Castorocauda is an extinct, semi-aquatic, superficially otter-like genus of docodont mammaliaforms with one species, C. lutrasimilis. It is part of the Yanliao Biota, found in the Daohugou Beds of Inner Mongolia, China dating to the Middle to Late Jurassic. It was part of an explosive Middle Jurassic radiation of Mammaliaformes moving into diverse habitats and niches. Its discovery in 2006, along with the discovery of other unusual mammaliaforms, disproves the previous hypothesis of Mammaliaformes remaining evolutionarily stagnant until the extinction of the non-avian dinosaurs.
Docodonta is an order of extinct Mesozoic mammaliaforms. They were among the most common mammaliaforms of their time, persisting from the Middle Jurassic to the Early Cretaceous across the continent of Laurasia. They are distinguished from other early mammaliaforms by their relatively complex molar teeth. Docodont teeth have been described as "pseudotribosphenic": a cusp on the inner half of the upper molar grinds into a basin on the front half of the lower molar, like a mortar-and-pestle. This is a case of convergent evolution with the tribosphenic teeth of therian mammals. There is much uncertainty for how docodont teeth developed from their simpler ancestors. Their closest relatives may have been certain Triassic "symmetrodonts", namely Woutersia, Delsatia.
Mammaliaformes is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia, and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes includes Docodonta and Hadrocodium.
Tritylodontidae is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy and details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous.
Gobiconodon is an extinct genus of carnivorous mammals belonging to the family Gobiconodontidae. Undisputed records of Gobiconodon are restricted to the Early Cretaceous of Asia and North America, but isolated teeth attributed to the genus have also been described from formations in England and Morocco dating as far back as the Middle Jurassic. Species of Gobiconodon varied considerably in size, with G. ostromi, one of the larger species, being around the size of a modern Virginia opossum. Like other gobiconodontids, it possessed several speciations towards carnivory, such as shearing molariform teeth, large canine-like incisors and powerful jaw and forelimb musculature, indicating that it probably fed on vertebrate prey. Unusually among predatory mammals and other eutriconodonts, the lower canines were vestigial, with the first lower incisor pair having become massive and canine-like. Like the larger Repenomamus there might be some evidence of scavenging.
The Itat Formation is a geologic formation in western Siberia. It was deposited in the Bajocian to Bathonian ages of the Middle Jurassic. Dinosaur remains are among the fossils that have been recovered from the formation, including the proceratosaurid Kileskus, as well as fish, amphibians, mammals and many other vertebrates. The formation is noted for bearing significant coal reserves, with large open pit coal mines extracting lignite from the unit currently in operation.
Paritatodon is an extinct mammaliaform which existed in Kyrgyzstan and England during the Jurassic period. It was originally the holotype specimen of Shuotherium kermacki, but Martin and Averianov (2010) argued that it resembled the genus Itatodon (Docodonta) and so renamed it Paritatodon.
Several mammals are known from the Mesozoic of Madagascar. The Bathonian Ambondro, known from a piece of jaw with three teeth, is the earliest known mammal with molars showing the modern, tribosphenic pattern that is characteristic of marsupial and placental mammals. Interpretations of its affinities have differed; one proposal places it in a group known as Australosphenida with other Mesozoic tribosphenic mammals from the southern continents (Gondwana) as well as the monotremes, while others favor closer affinities with northern (Laurasian) tribosphenic mammals or specifically with placentals. At least five species are known from the Maastrichtian, including a yet undescribed species known from a nearly complete skeleton that may represent a completely new group of mammals. The gondwanathere Lavanify, known from two teeth, is most closely related to other gondwanatheres found in India and Argentina. Two other teeth may represent another gondwanathere or a different kind of mammal. One molar fragment is one of the few known remains of a multituberculate mammal from Gondwana and another has been interpreted as either a marsupial or a placental.
Polistodon is an extinct genus of tritylodonts from the Bajocian or Callovian of China. It contains one species, P. chuannanensis, which is known from a single skull from the Xiashaximiao Formation. According to one study, it may be a species of the genus Stereognathus.
Xenocretosuchus is an extinct genus of tritylodont therapsids from the Aptian Ilek Formation of Siberia, in the Russian Federation. The type species, X. sibiricus, is known only from dental elements, as is X. kolossovi, described from the Batylykh Formation in 2008. Some authors have treated these species as part of the genus Stereognathus, otherwise known from the Middle Jurassic of Britain, but this is rejected by other authors.
The Ilek Formation is a Lower Cretaceous geologic formation in Western Siberia. Many different fossils have been recovered from the formation. It overlies the Late Jurassic Tyazhin Formation and underlies the Albian Kiya Formation.
The Kilmaluag Formation is a Middle Jurassic geologic formation in Scotland. It was formerly known as the Ostracod Limestone for preserving an abundance of fossil freshwater/low salinity ostracods. Gastropods, bivalves, trace fossil burrows, and vertebrate fossil remains have also been recorded from the formation. Vertebrate fossils include fish, crocodylomorphs, mammals, small reptiles, amphibians, theropod and sauropod dinosaurs and pterosaurs.
Robert Joseph Gay Savage was a British palaeontologist known as Britain's leading expert on fossil mammals. He worked at the University of Bristol for nearly 40 years and studied fossils around the world, especially in North and East Africa. He produced the 1986 popular science book Mammal Evolution: An Illustrated Guide and co-edited several technical books in the Fossil Vertebrates of Africa series with fellow palaeontologist Louis Leakey.
Jugulator is an extinct genus of mammals from the Cretaceous of North America. It contains one species, Jugulator amplissimus. A eutriconodont, it is known from the Cedar Mountain Formation, and is both a large sized and possibly ecologically specialised taxon, showcasing the diversity of mammals in the Mesozoic.
Montirictus is an extinct genus of tritylodonts known from the Early Cretaceous Kuwajima Formation of Japan. It was among the latest surviving tritylodontids, and is closely related to the earlier Xenocretosuchus from mainland Asia, and the Jurassic Stereognathus from the UK. It may be a species of the genus Stereognathus, but resolution of its affinities conditions upon the discovery of additional material.
Borealestes is a genus of docodontan from the Middle Jurassic of Britain, first discovered on the Isle of Skye near the village of Elgol. It was the earliest mammaliaform from the Mesozoic found and named in Scotland. A second species and was later found in other Middle Jurassic sites in England, but is now shown to be a different genus. A new species, B. cuillinensis was named in 2021, also from Skye.
Palaeoxonodon is an extinct genus of cladotherian mammal from the Middle Jurassic of England and Scotland.
Dobunnodon its an extinct genus of docodont from the Middle Jurassic (Bathonian) Forest Marble Formation of England, first discovered in Oxfordshire near the village of Kirtlington. The type species, D. mussettae, was originally named as a species of Borealestes in 2003.
Michael Waldman is a British palaeontologist known for his work on fossil fish, mammals, and reptiles. He also discovered the globally important fossil site of Cladach a'Ghlinne, near Elgol on the Isle of Skye, Scotland. This site exposes the Kilmaluag Formation and provides a valuable record of Middle Jurassic ecosystems. During the 1970s he visited the site several times with fellow palaeontologist Robert Savage. The fossil turtle Eileanchelys waldmani was named after Michael in recognition of his notable contribution to palaeontology.