Redfieldiiformes Temporal range: | |
---|---|
Dictyopyge macrurus from the Late Triassic of Virginia | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Subclass: | Neopterygii (?) |
Order: | † Redfieldiiformes Berg, 1940 |
Type genus | |
† Redfieldius Hay, 1899 | |
Genera | |
See text |
Redfieldiiformes is an extinct order of ray-finned fish (actinopterygians) which lived from the Early Triassic to Early Jurassic. Redfieldiiforms were fairly typical Triassic fish in overall anatomy. They had a fusiform (streamlined, tuna-like) body shape with thick, ganoine-covered scales. The dorsal and anal fins were large, positioned opposite from each other, and shifted back, close to the tail. The caudal fin was hemiheterocercal, with the vertebral column and body scales extending into an upper lobe equal in size and shape to the lower lobe. They also had several characteristic skeletal traits, such as a hatchet-shaped preopercle, a series of fulcra (thin spiny scales) fringing the fins, a reduced number of branchiostegal rays (typically just one), and a snout ornamented with tubercles. [1] [2] [3]
The maxilla has small teeth and is strongly connected to the preopercle; this would have allowed a deep gape to assist in ram feeding. [1] The function of the snout tubercles is uncertain; some authors have suggested that they attached to a fleshy upper lip, [2] while others argue that they could have held sensory organs akin to the tubercles of siluriforms (catfish). [3]
Redfieldiiforms were fairly primitive 'subholostean' fish (more primitive than holosteans such as gars or the bowfin), with uncertain relations to neopterygians. [1] Some studies draw comparisons to 'perleidiforms' or ptycholepids, while others consider redfieldiiforms to be early neopterygians related to pholidopleuriforms. [4] Redfieldiiforms were exclusively freshwater fish which became prominent in southern Gondwana (Argentina, Australia, southern Africa) during the Middle Triassic. [3] By the Late Triassic they had become a major component of freshwater ecosystems in western Laurasia (North America and Morocco). However, they were much rarer further east (South Korea, [3] possibly Switzerland [5] ). The last member of the order, Redfieldius , lived in the Early Jurassic of eastern North America. [1] [3]
The following families and genera are known:
The enigmatic marine fish Pacorichthys from the Ladinian of Europe somewhat resembles a redfieldiiform, although it was marine unlike the freshwater nature of other redfieldiiforms. [5]
The Amiiformes order of fish has only two extant species, the bowfins: Amia calva and Amia ocellicauda, the latter recognized as a separate species in 2022. These Amiiformes are found in the freshwater systems of North America, in the United States and parts of southern Canada. They live in freshwater streams, rivers, and swamps. The order first appeared in the Triassic, and the extinct members include both marine and freshwater species, many of which are morphologically disparate from bowfins, such as the caturids.
Birgeria is a genus of carnivorous marine ray-finned fish from the Triassic period. Birgeria had a global distribution, with fossil known from Madagascar, Spitsbergen, Germany, Switzerland, Italy, Slovenia, China, Russia, Canada and Nevada, United States. The oldest fossils are from Griesbachian aged beds of the Wordie Creek Formation of East Greenland. Birgeria existed throughout the entire Triassic period, from the very beginning just after the Permian-Triassic mass extinction, up to the very end with its extinction during the Triassic-Jurassic mass extinction.
Pachycormiformes is an extinct order of marine ray-finned fish known from the Early Jurassic to the end of the Cretaceous. It only includes a single family, Pachycormidae. They were characterized by having serrated pectoral fins, reduced pelvic fins and a bony rostrum. Pachycormiformes are morphologically diverse, containing both tuna and swordfish-like carnivorous forms, as well as edentulous suspension-feeding forms.
Daedalichthys is an extinct genus of prehistoric freshwater ray-finned fish that lived during the Early Triassic epoch. It contains a single species, D. formosa from the Olenekian-aged Cynognathus Assemblage Zone of South Africa. It was previously classified in Dictyopyge.
Brookvalia is an extinct genus of prehistoric freshwater ray-finned fish that lived during the Middle Triassic epoch.
Cosmoptychius is an extinct genus of freshwater and estuarine ray-finned fish that lived during the Mississippian epoch. It contains a single species, C. striatus from the Visean of Scotland. It has a wide stratigraphic range in the Wardie Shale, with many specimens known.
Ctenognathichthys is an extinct genus of prehistoric marine ray-finned fish that lived during the Middle Triassic epoch of Europe, in the former Tethys Ocean.
Ptychoceratodus is an extinct genus of lungfish living from Early Triassic to Middle Jurassic. It was established by Otto Jaekel for one species, transferred from Ceratodus genus. Type species is P. serratus from the Middle Triassic of Switzerland and Germany. Ptychoceratodus had two pairs of massive dental plates, bearing 4-6 acute ridges. Its skull roof was composed from massive, plate-like bones. In the central part of skull roof was localized an unossified fenestra. Most of the Ptychoceratodus findings are isolated dental plates, some associated with jaws. Other parts of skull or postcranial skeleton are relatively rarely found as fossils. The anatomy of skull is the best recognized in P. serratus, whereas less complete cranial material is available also for P. concinuus, P. phillipsi, and P. rectangulus. Although Ptychoceratodus is known exclusively from the Triassic and Jurassic, there were also Cretaceous specimens referred to this genus. However, they are more often regarded as representants of Metaceratodus. Ptychoceratodus is the only member of the family Ptychoceratodontidae. The first named species is P. phillipsi by Louis Agassiz in 1837 as a species of Ceratodus and later moved to Ptychoceratodus genus. Occurrences of Ptychoceratodus come mainly from Europe. However, occurrences from other continents suggest it was dispersed globally during the Triassic. After 2010, the new fossil material behind the Europe was reported from South America, India, and Greenland
Acentrophorus is an extinct genus of prehistoric freshwater and marine ray-finned fish from the Roadian to the Wuchiapingian of England, Germany (Kupferschiefer), Italy and Russia. There may also be a Triassic occurrence in Australia.
Cyranorhis is an extinct genus of prehistoric marine ray-finned fish that lived during the Serpukhovian age of the Carboniferous period. One species is known, C. bergeraci in the Bear Gulch Limestone what is now Montana, United States. It is named after French novelist Cyrano de Bergerac.
Cycloptychius is an extinct genus of prehistoric freshwater & marine ray-finned fish that existed throughout much of the Carboniferous period in Eurasia, and possibly into the Early Permian in South Africa. It was a member of the Rhadinichthyidae, a family of basal ray-finned fish that was formerly placed in the now-paraphyletic order Palaeonisciformes.
Lonchidion is a genus of extinct hybodont in the family Lonchidiidae. The genus first appears in the fossil record during the Middle Triassic (Ladinian) and was among the last surviving hybodont genera, with its youngest known fossils dating to the very end of the Cretaceous (Maastrichtian).
The Colorado City Formation is a Late Triassic geologic formation in the Dockum Group of Texas, United States. It has previously been known as the Iatan Member, Colorado City Member or 'Pre-Tecovas Horizon'.
Dorsetichthys is an extinct genus of stem-teleost ray-finned fish from the Early Jurassic period of Europe.
The Zhuganpo Formation is a Triassic geologic unit found in southern China. It has historically been known as the Zhuganpo Member of the Falang Formation. A diverse fossil assemblage known as the Xingyi biota or Xingyi Fauna can be found in the upper part of the Zhuganpo Formation. Fossils of the Xingyi biota include articulated skeletons of marine reptiles, abundant fish, and a plentiful assortment of invertebrates indicating a Ladinian to Carnian age for the sediments of the formation.
Ginglymodi is a clade of ray-finned fish containing modern-day gars (Lepisosteidae) and their extinct relatives in the order Lepisosteiformes, the extinct orders Semionotiformes and Kyphosichthyiformes, and various other extinct taxa. Ginglymodi is one of the two major subgroups of the infraclass Holostei, the other one being Halecomorphi, which contains the bowfin and eyespot bowfin and their fossil relatives.
Dapediidae is an extinct family of neopterygian ray-finned fish that lived from the Middle Triassic to Late Jurassic. It is the only family of the order Dapediiformes. Its members were historically placed within the ginglymodian family Semionotidae, but were moved to their own family in 1966.
Australopachycormus is an extinct genus of marine pachycormiform fish. It is only known from the type species, A. hurleyi, from the Early Cretaceous (Albian) Toolebuc Formation of Queensland, Australia. Like the related Protosphyraena, it possessed an elongated rostrum. Recent studies have recovered it as a junior synonym of Protosphyraena because of their lack of chronological separation and because of the fragmentary pectoral fin of Australopachycormus which is indistinguishable to consider it as a valid taxon.
Redfieldius is an extinct genus of freshwater ray-finned fish that inhabited eastern North America during the Early Jurassic period. It contains a single species, R. gracilis, known from the Hettangian to the Sinemurian of the northeastern United States. It is the type genus and was the last surviving member of the order Redfieldiiformes, which was widespread and diverse throughout the preceding Triassic period. It is notable for representing possibly the first fossil bony fish collected from North America, with a specimen from Middletown collected in 1816 by Benjamin Silliman.
Tanaocrossus is an extinct genus of primitive freshwater ray-finned fish that inhabited southwestern and eastern North America during the Late Triassic period. It contains a single known species, T. kalliokoskii, known from the United States, with indeterminate species also known.