Redfieldiiformes Temporal range: | |
---|---|
Dictyopyge macrurus from the Late Triassic of Virginia | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Subclass: | Neopterygii (?) |
Order: | † Redfieldiiformes Berg, 1940 |
Genera | |
See text |
Redfieldiiformes is an extinct order of ray-finned fish (actinopterygians) which lived from the Middle Triassic to Early Jurassic. Redfieldiiforms were fairly typical Triassic fish in overall anatomy. They had a fusiform (streamlined, tuna-like) body shape with thick, ganoine-covered scales. The dorsal and anal fins were large, positioned opposite from each other, and shifted back, close to the tail. The caudal fin was hemiheterocercal, with the vertebral column and body scales extending into an upper lobe which was equal in size and shape to the lower lobe. They also had several characteristic skeletal traits, such as a hatchet-shaped preopercle, a series of fulcra (thin spiny scales) fringing the fins, a reduced number of branchiostegal rays (typically just one), and a snout ornamented with tubercles. [1] [2] [3]
The maxilla has small teeth and is strongly connected to the preopercle; this would have allowed a deep gape to assist in ram feeding. [1] The function of the snout tubercles is uncertain; some authors have suggested that they attached to a fleshy upper lip, [2] while others argue that they could have held sensory organs akin to the tubercles of siluriforms (catfish). [3]
Redfieldiiforms were fairly primitive 'subholostean' fish (more primitive than holosteans such as gars or the bowfin), with uncertain relations to neopterygians. [1] Some studies draw comparisons to 'perleidiforms' or ptycholepids, while others consider redfieldiiforms to be early neopterygians related to pholidopleuriforms. [4] Redfieldiiforms were exclusively freshwater fish which became prominent in southern Gondwana (Argentina, Australia, southern Africa) during the Middle Triassic. [3] By the Late Triassic they had become a major component of freshwater ecosystems in western Laurasia (North America and Morocco), though they were much rarer further east (South Korea, [3] possibly Switzerland [5] ). The last member of the order, Redfieldius , lived in the Early Jurassic of eastern North America. [1] [3]
Actinopterygii, members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webs of skin supported by radially extended bony spines, as opposed to the bulkier, fleshy lobed fins of the sister class Sarcopterygii. Resembling folding fans, the actinopterygian fins can change shape and wetted area easily, providing superior thrust-to-weight ratios per movement compared to sarcopterygian and chondrichthyian fins. The fin rays attach directly to the proximal or basal skeletal elements, the radials, which represent the articulation between these fins and the internal skeleton.
The Amiiformes order of fish has only two extant species, the bowfins: Amia calva and Amia ocellicauda, the latter recognized as a separate species in 2022. These Amiiformes are found in the freshwater systems of North America, in the United States and parts of southern Canada. They live in freshwater streams, rivers, and swamps. The order first appeared in the Triassic, and the extinct members include both marine and freshwater species, many of which are morphologically disparate from bowfins, such as the caturids.
Chondrostei is a group of non-neopterygian ray-finned fish. While the term originally referred to the paraphyletic grouping of all non-neopterygian ray-finned fish, it was redefined by Patterson in 1982 to be a clade comprising the Acipenseriformes and their extinct relatives.
Neopterygii is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant fishes, and over half of all living vertebrate species. While living holosteans include only freshwater taxa, teleosts are diverse in both freshwater and marine environments. Many new species of teleosts are scientifically described each year.
Dapedium is an extinct genus of primitive neopterygian ray-finned fish. The first-described finding was an example of D. politum, found in the Lower Lias of Lyme Regis, on the Jurassic Coast of England. Dapedium lived in the late Triassic and Jurassic periods.
Saurichthys is an extinct genus of predatory ray-finned fish from the Triassic Period. It is the type genus of the family Saurichthyidae, and the most speciose and longest lasting genus in the family. This family also includes the Permian Eosaurichthys (China) and the Jurassic Saurorhynchus from Europe and North America, though it may be more appropriate to treat these as subgenera of Saurichthys, due to the genus Saurichthys otherwise being paraphyletic.
Birgeria is a genus of carnivorous marine ray-finned fish from the Triassic period. Birgeria had a global distribution. Fossils were found in Madagascar, Spitsbergen, Germany, Switzerland, Italy, Slovenia, China, Russia, Canada and Nevada, United States. The oldest fossils are from Griesbachian aged beds of the Wordie Creek Formation of East Greenland.
Dicellopyge is an extinct genus of freshwater ray-finned fish that lived during the Anisian age of the Middle Triassic epoch in what is now South Africa. It was originally named "Dicellopygae" by James Brough but the name was later corrected to Dicellopyge by Peter Hutchinson.
Pycnodontiformes is an extinct order of primarily marine bony fish. The group first appeared during the Late Triassic and disappeared during the Eocene. The group has been found in rock formations in Africa, Asia, Europe, North and South America. They were small to middle-sized fish, generally with laterally-compressed deep bodies, some with almost circular outlines, adapted for manuverability in reef-like environments, though the group was morphologically diverse. Most, but not all members of the groups had jaws with round and flattened teeth, well adapted to crush food items (durophagy), such as echinoderms, crustaceans and molluscs. Some pyncodontiformes developed piranha like teeth used for eating flesh. Most species inhabited shallow marine reef environments, while a handful of species lived in freshwater or brackish conditions. While rare during the Triassic and Early-Middle Jurassic, Pycnodontiformes became abundant and diverse during the Late Jurassic, exhibiting a high but relatively static diversity during the Early Cretaceous. At the beginning of the Late Cretaceous they reached their apex of morphological and species diversity, after which they began to gradually decline, with a more sudden decline at the end of the Cretaceous due to the collapse of reef ecosystems, finally becoming extinct during the Eocene. They are considered to belong to the Neopterygii, but their relationship to other members of that group is uncertain.
Ctenognathichthys is an extinct genus of prehistoric bony fish that lived during the Anisian and Ladinian ages of the Middle Triassic epoch in what is now southern/southeastern Switzerland and northern Italy.
Saurichthyiformes is an extinct order of ray-finned fish which existed in Asia, Africa, Australia, Europe and North America, during the late Permian to early Middle Jurassic. Saurichthyiiformes comprise two families, Saurichthyidae and Yelangichthyidae. Yelangichthyidae is monotypic, containing only the genus Yelangichthys. The gar or needlefish-like Saurichthyidae is primarily known from the genus Saurichthys. Additionally, the subgenera SaurorhynchusCostasaurichthys, Eosaurichthys, Lepidosaurichthys, and Sinosaurichthys are frequently used to group species, and are sometimes considered separate genera. Species are known from both marine end freshwater deposits. They had their highest diversity during the Early and Middle Triassic. Their phylogenetic position is uncertain, while they have often been considered members of Chondrostei, and thus related to living sturgeons and paddlefish, phylogenetic analysis of well-preserved remains has considered this relationship equivocal. They may actually belong to the stem-group of Actinopterygii, and thus not closely related to any living group of ray-finned fish.
Kyphosichthys is an extinct genus of basal actinopterygian bony fish known from the lower Middle Triassic (Anisian) marine deposits in Luoping, eastern Yunnan Province, southwestern China. The species is the first known fossil record of highly deep-bodied ginglymodians.
The evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared. Most jawless fish are now extinct; but the extant lampreys may approximate ancient pre-jawed fish. Lampreys belong to the Cyclostomata, which includes the extant hagfish, and this group may have split early on from other agnathans.
This list of fossil fishes described in 2016 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes and other fishes of every kind that have been described during the year 2016, as well as other significant discoveries and events related to paleontology of fishes that occurred in the year 2016. The list only includes taxa at the level of genus or species.
Sam Giles is a palaeobiologist at the University of Birmingham. Her research combines modern imaging with fossils to understand the evolution of life, in particular that of early fish, and in 2015 "rewrote" the vertebrate family tree. She was a 2017 L'Oréal-UNESCO Rising Star and won the 2019 Geological Society of London Lyell Fund.
The Intertemporal bone is a paired cranial bone present in certain sarcopterygians and extinct amphibian-grade tetrapods. It lies in the rear part of the skull, behind the eyes.
Ginglymodi is a clade of ray-finned fish containing modern-day gars (Lepisosteidae) and their extinct relatives, including the family Lepidotidae and the orders Semionotiformes and Kyphosichthyiformes, and various other extinct taxa. Ginglymodi is one of the two major subgroups of the infraclass Holostei, the other one being Halecomorphi, which contains the bowfin and its fossil relatives.
Dapediidae is an extinct family of neopterygian ray-finned fish that lived from the Late Triassic to Late Jurassic. It is the only family of the order Dapediiformes. Its members were historically placed within the ginglymodian family Semionotidae, but were moved to their own family in 1966.
Coccolepididae is an extinct family of ray-finned fish, known from the Early Jurassic to Early Cretaceous, most of which were originally referred to the type genus Coccolepis. They had a widespread distribution, being found in North and South America, Australia, Asia and Europe. They are mostly known from freshwater environments, though several species have been found in marine environments. They are morphologically conservative, and have poorly ossified endo and exoskeletons, which usually results in poor preservation. This makes it difficult to distinguish species. They are generally small fish, with the largest known specimens reaching a length of 210 mm. Historically, they have been classified as members of “Palaeonisciformes”, a paraphyletic grouping of non-neopterygian fish, due to their plesiomorphic conservative morphology closely resembling those of many other groups of primitive fish. They have been suggested to be relatives of the Acipenseriformes within the Chondrostei.
Turseodus is an extinct genus of ray-finned fish found in Late Triassic freshwater sediments of the United States. Two species have been described, T. acutus from the Lockatong Formation of Pennsylvania, and T. dolorensis from the Chinle Formation of Colorado.