Grallator | |
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Typical footprint form | |
Trace fossil classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Ichnofamily: | † Grallatoridae |
Ichnogenus: | † Grallator Hitchcock, 1858 |
Type ichnospecies | |
†Grallator cursorius Hitchcock, 1858 | |
Ichnospecies | |
Synonyms | |
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Grallator ["GRA-luh-tor"] is an ichnogenus (form taxon based on footprints) which covers a common type of small, three-toed print made by a variety of bipedal theropod dinosaurs. Grallator-type footprints have been found in formations dating from the Early Triassic through to the early Cretaceous periods. They are found in the United States, Canada, Europe, India, Australia, Brazil (Sousa and Santa Maria Formations) and China, [1] but are most abundant on the east coast of North America, especially the Triassic and Early Jurassic formations of the northern part of the Newark Supergroup. [2] [3] The name Grallator translates into "stilt walker", although the actual length and form of the trackmaking legs varied by species, usually unidentified. The related term "Grallae" is an ancient name for the presumed group of long-legged wading birds, such as storks and herons. These footprints were given this name by their discoverer, Edward Hitchcock, in 1858. [2]
Grallator footprints are characteristically three-toed (tridactyl) and range from 10 to 20 centimeters (or 4 to 8 inches) long. Though the tracks show only three toes, the trackmakers likely had between four and five toes on their feet. While it is usually impossible to match these prints with the exact dinosaur species that left them, it is sometimes possible to narrow down potential trackmakers by comparing the proportions in individual Grallator ichnospecies with known dinosaurs of the same formation. For example, Grallator tracks identified from the Yixian Formation may have been left by Caudipteryx . [3]
Source: [4]
Fossil tracks can be informative about theropod pathologies but apparently pathological traits may be due to unusual behaviors. Sandstone stratum dating to the Norian in southern Wales preserves tracks of an individual with a deformed digit III attributed to the ichnogenus Anchisauripus . The distal end of the digit was consistently flexed. However, this apparent pathology could be caused by the animal rotating the tip of that digit when lifting the foot. [6]
Grallator-type footprints have been found in formations dating from the Early Triassic through to the early Cretaceous periods. They are found in the United States, Canada, Europe, Australia, Brazil (Sousa and Santa Maria Formations) and China, [1] but are most abundant on the east coast of North America, especially the Triassic and Early Jurassic formations of the northern part of the Newark Supergroup. [2] [3]
The most famous, and archetypal tracks that conform to the Grallator type are those found on the East Coast of North America, specifically from the Late Triassic to Early Jurassic Newark Supergroup. These footprints were likely made by an unidentified, primitive dinosaur similar to Coelophysis . [2] The Newark Supergroup footprints show digits II, III and IV, but no trace of the shorter digits I and V which would likely have been present in a dinosaur of this stage. The outer two digits would have been stubby and ineffective, not touching the ground during walking or running. [2] Despite losing most of their effectiveness, dinosaur evolution had not yet removed these digits to fully streamline the foot. This is known because rare specimens are found with traces of these outer digits. Digits II, III and IV have 3, 4 and 5 phalanges respectively, giving Grallator a ?-3-4-5-? digital formula.
Although the Newark Supergroup Grallator tracks were made by a bipedal saurischian dinosaur, they can easily be mistaken for those of the late Triassic ichnogenus Atreipus . [7] The trackmaker of Atreipus prints was a quadrupedal ornithischian. The reason for this similarity is a lack of divergence in the foot evolution of the two distinct groups of dinosaurs: ornithischians and saurischians.
In January 2021, while walking with her father Richard Wilder, a four-year-old girl called Lily Wilder found a 215- to 220-million year-old dinosaur footprint at Bendricks Bay in the Vale of Glamorgan, Wales. [8] [9] [10]
Experts believe that the footprint was most likely left by a dinosaur (grallator) that stood about 75 centimeters (29.5 inches) tall and 2.5 meters (about 8 feet) long and walked on its two hind feet. [11] [12]
The scientists called the girl's discovery "the finest impression of a 215 million-year-old dinosaur print found in Britain in a decade". [8] Karl-James Langford of Archaeology Cymru considered the find to be "internationally important". [8] [13] Cindy Howells, a palaeontologist at Amgueddfa Cymru – National Museum Wales, described it as "one of the best-preserved examples from anywhere in the UK" and said that it "will really aid palaeontologists to get a better idea about how these early dinosaurs walked". [10]
Gwyneddosaurus is a possibly invalid genus of extinct aquatic tanystropheid reptile. The type species, G. erici was described in 1945 by Wilhelm Bock, who identified it as a coelurosaurian dinosaur related to Podokesaurus. Its remains were found in the Upper Triassic Lockatong Formation of Montgomery County, eastern Pennsylvania, and the holotype includes skull fragments, several vertebra, ribs, gastralia, partial shoulder and hip bones, and several forelimb and hindlimb elements found in soft shale, while the paratype includes a femur and a tibia. The type specimen is ANSP 15072 and it was discovered by Bock's four-year-old son while the paratype is only listed as ?(ASNP coll.). It was not a large animal; the type skeleton was estimated by Bock as 18 centimetres (7.1 in) long, and its thigh bone was only 23 millimeters long (0.91 in).
Eubrontes is the name of fossilised dinosaur footprints dating from the Late Triassic and Early Jurassic. They have been identified from France, Poland, Slovakia, Czech Republic, Italy, Spain, Sweden, Australia (Queensland), US, India, China and Brazil (South).
Otozoum is an extinct ichnogenus of sauropodomorph dinosaur from the Late Triassic-Middle Jurassic sandstones. Footprints were made by heavy, bipedal or, sometimes, quadrupedal animals with a short stride that walked on four toes directed forward. These footprints are relatively large, over 20 cm in pes length. Otozoum differs from Plateosaurus by having a notable homopody.
New Chum is a suburb in the City of Ipswich, Queensland, Australia. In the 2021 census, New Chum had "no people or a very low population".
The Blackstone Formation is a geologic formation of the Ipswich Coal Measures Group in southeastern Queensland, Australia, dating to the Carnian to Norian stages of the Late Triassic. The shales, siltstones, coal and tuffs were deposited in a lacustrine environment. The Blackstone Formation contains the Denmark Hill Insect Bed.
Stegomosuchus is an extinct genus of small protosuchian crocodyliform. It is known from a single incomplete specimen discovered in the late 19th century in Lower Jurassic rocks of south-central Massachusetts, United States. It was originally thought to be a species of Stegomus, an aetosaur, but was eventually shown to be related to Protosuchus and thus closer to the ancestry of crocodilians. Stegomosuchus is also regarded as a candidate for the maker of at least some of the tracks named Batrachopus in the Connecticut River Valley.
Komlosaurus is an ichnogenus of theropod dinosaur from the Early Jurassic of Baranya, Hungary. The type species, Komlosaurus carbonis, was described by Kordos in 1983. The type remains come from the Mecsek Coal Formation, from the Middle Hettangian to the Early Sinemurian, and comprise several footprints.
Sphingopus is an ichnogenus of dinosauromorph footprints found in sediments dating to 250 and 228 Ma. The exact species which created the Sphingopus tracks have not been identified.
Jialingpus is an ichnogenus of dinosaur, likely a theropod. Its footprints have been found in the Feitianshan Formation, a low-energy lake formation. Holotype is SCFP-24, which was found in Late Jurassic (Oxfordian)-aged Yuechi tracksite at Huanglong, China. These footprints were found within the vicinity of those of the smaller theropod Minisauripus, meaning that Jialingpus likely hunted Minisauripus.
Anticheiropus is an ichnogenus of dinosaur footprint belonging to a saurischian. It has only been discovered in Massachusetts. Two ichnospecies are known : A. hamatus and A. pilulatus, both discovered around 1863 and named by Edward Hitchcock in 1865.
Paleontology in Virginia refers to paleontological research occurring within or conducted by people from the U.S. state of Virginia. The geologic column in Virginia spans from the Cambrian to the Quaternary. During the early part of the Paleozoic, Virginia was covered by a warm shallow sea. This sea would come to be inhabited by creatures like brachiopods, bryozoans, corals, and nautiloids. The state was briefly out of the sea during the Ordovician, but by the Silurian it was once again submerged. During this second period of inundation the state was home to brachiopods, trilobites and entire reef systems. During the mid-to-late Carboniferous the state gradually became a swampy environment.
Tanytrachelos is an extinct genus of tanystropheid archosauromorph reptile from the Late Triassic of the eastern United States. It contains a single species, Tanytrachelos ahynis, which is known from several hundred fossil specimens preserved in the Solite Quarry in Cascade, Virginia. Abundant fossils of Tanytrachelos are found in a series of lakebed sediments that were deposited over the course of about 350 thousand years in a lake which existed approximately 230 million years ago. Some fossils are very well-preserved and include the remains of soft tissues. Tanytrachelos is the most likely trackmaker of the ichnogenus Gwyneddichnium.
The Tomanová Formation is a Late Triassic geologic formation in Poland and Slovakia. Fossil theropod tracks have been reported from the formation.
The 20th century in ichnology refers to advances made between the years 1900 and 1999 in the scientific study of trace fossils, the preserved record of the behavior and physiological processes of ancient life forms, especially fossil footprints. Significant fossil trackway discoveries began almost immediately after the start of the 20th century with the 1900 discovery at Ipolytarnoc, Hungary of a wide variety of bird and mammal footprints left behind during the early Miocene. Not long after, fossil Iguanodon footprints were discovered in Sussex, England, a discovery that probably served as the inspiration for Sir Arthur Conan Doyle's The Lost World.
The 19th century in ichnology refers to advances made between the years 1800 and 1899 in the scientific study of trace fossils, the preserved record of the behavior and physiological processes of ancient life forms, especially fossil footprints. The 19th century was notably the first century in which fossil footprints received scholarly attention. British paleontologist William Buckland performed the first true scientific research on the subject during the early 1830s.
This article records new taxa of trace fossils of every kind that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to trace fossil paleontology that are scheduled to occur in the year 2019.
Gwyneddichnium is an ichnogenus from the Late Triassic of North America and Europe. It represents a form of reptile footprints and trackways, likely produced by small tanystropheids such as Tanytrachelos. Gwyneddichnium includes a single species, Gwyneddichnium major. Two other proposed species, G. elongatum and G. minore, are indistinguishable from G. major apart from their smaller size and minor taphonomic discrepancies. As a result, they are considered junior synonyms of G. major.
Bellatoripes is an ichnogenus of footprint produced by a large theropod dinosaur so far known only from the Late Cretaceous of Alberta and British Columbia in Canada. The tracks are large and three-toed, and based on their size are believed to have been made by tyrannosaurids, such as Albertosaurus and Daspletosaurus. Fossils of Bellatoripes are notable for preserving trackways of multiple individual tyrannosaurids all travelling in the same direction at similar speeds, suggesting the prints may have been made by a group, or pack, of tyrannosaurids moving together. Such inferences of behaviour cannot be made with fossil bones alone, so the record of Bellatoripes tracks together is important for understanding how large predatory theropods such as tyrannosaurids may have lived.
Wakinyantanka is an ichnogenus of footprint produced by a large theropod dinosaur from the Late Cretaceous Hell Creek Formation of South Dakota. Wakinyantanka tracks are large with three long, slender toes with occasional impressions of a short hallux and narrow metatarsals. Wakinyantanka was the first dinosaur track to be discovered in the Hell Creek Formation, which remain rare in the preservational conditions of the rocks. The potential trackmakers may be a large oviraptorosaur or a small tyrannosaurid.
Rhynchosauroides is an ichnogenus, a form taxon based on footprints. The organism producing the footprints was likely a lepidosaur and may have been a sphenodont, an ancestor of the modern tuatara. The footprint consists of five digits, of which the fifth is shortened and the first highly shortened.