Riojasaurids Temporal range: Late Triassic, | |
---|---|
Riojasaurus skull cast | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Massopoda |
Clade: | † Riojasauridae Yates, 2007 |
Genera | |
Riojasauridae is an extinct family of sauropodomorph dinosaurs from the Late Triassic Period (late Carnian to Norian Ages). It contains the genera Riojasaurus and Eucnemesaurus . The Riojasauridae is considered a stem taxon, and is defined as "the most inclusive clade containing Riojasaurus incertus but not Plateosaurus engelhardti , Massospondylus carinatus , or Anchisaurus polyzelus ". [1] Geologic formations containing riojasaurid fossils include the Lower Elliot Formation of Orange Free State, South Africa (where fossils of Eucnemesaurus have been found), and the Los Colorados Formation, in La Rioja Province, Argentina (where fossils of Riojasaurus have been recovered). [1]
The Riojasauridae are considered to be sauropodomorphs, but not sauropods themselves. This means that they were generally much smaller than the Sauropods of the Jurassic and Cretaceous Periods, and members of this family may not have been obligate quadrupeds, the way more derived Sauropods were. The relationships of this family within other sauropodomorphs has been considered by studies in 2007 [1] and 2020 [2] with slightly different results.
The cladogram below shows basal sauropodomorph relationships simplified after Yates, 2007. In this analysis, Riojasauridae was recovered as the earliest-diverging lineage of the clade Massopoda, and represents a sister group to the node formed by Massospondylidae, the genus Jingshanosaurus and the Anchisauria. [1]
Plateosauria |
| ||||||||||||||||||||||||||||||||||||||||||||||||
The following cladogram is from a phylogenetic analysis by Peyre de Fabrègues et al., 2020. Here, Riojasauridae was placed in a more derived position, lying closer to Sauropoda than to Massospondylidae, and appearing as the out group to the clade formed by the genus Seitaad and the Sauropodiformes. [2] The Massospondylidae and the taxa considered to be related to the family by Yates, are shown to be represent more primitive lineages. [2]
Massopoda |
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sauropodomorpha is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The prosauropods, which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic until their decline and extinction at the end of the Cretaceous.
Melanorosaurus is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. A herbivore from South Africa, it had a large body and sturdy limbs, suggesting it moved about on all fours. Its limb bones were massive and heavy like the limb bones of true sauropods.
Massospondylus was a genus of sauropodomorph dinosaur from the Early Jurassic. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. Fossils have since been found at other locations in South Africa, Lesotho, and Zimbabwe. Material from Arizona's Kayenta Formation, India, and Argentina has been assigned to the genus at various times, but the Arizonan and Argentinian material are now assigned to other genera.
Isanosaurus is an extinct genus of sauropod dinosaur from Thailand. It was originally dated to approximately 210 million years ago during the Late Triassic, which would make it one of the oldest known sauropods. Its age was later considered uncertain, and may be Early Jurassic or even as young as Late Jurassic. The only species is Isanosaurus attavipachi. Though important for the understanding of sauropod origin and early evolution, Isanosaurus is poorly known. Exact relationships to other early sauropods remain unresolved.
Antetonitrus is a genus of sauropod dinosaur found in the Early Jurassic Elliot Formation of South Africa. The only species is Antetonitrus ingenipes. As one of the oldest known sauropods, it is crucial for the understanding of the origin and early evolution of this group. It was a quadrupedal herbivore, like all of its later relatives, but shows primitive adaptations to use the forelimbs for grasping, instead of purely for weight support.
Blikanasaurus is a genus of sauropodomorph dinosaur from the late Triassic of South Africa. The generic name Blikanasaurus is derived from Greek, meaning "lizard from Blikana". The species name cromptoni is taken from the surname of A.W. “Fuzz” Crompton, an American paleontologist who led numerous field expeditions in Elliot Formation outcrop localities in South Africa. Blikanasaurus is only known from partial hindlimb bones that were recovered from the lower Elliot Formation (LEF) in the Eastern Cape.
Coloradisaurus is a genus of massospondylid sauropodomorph dinosaur. It lived during the Late Triassic period in what is now La Rioja Province, Argentina. It is known from two specimens collected from the Los Colorados Formation of the Ischigualasto-Villa Unión Basin.
Eucnemesaurus is a basal sauropodomorph dinosaur genus usually considered to be a synonym of Euskelosaurus. Recent study by Yates (2006), however, indicates that it is valid and the same animal as putative "giant herrerasaurid" Aliwalia.
Gryponyx is an extinct genus of massopod sauropodomorph known from southern Free State, central South Africa.
Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and Antarctica during the Late Triassic to the Early Jurassic periods. Several dinosaurs have been classified as massospondylids over the years. The largest cladistic analysis of early sauropodomorphs, which was presented by Apaldetti and colleagues in November 2011, found Adeopapposaurus, Coloradisaurus, Glacialisaurus, Massospondylus, Leyesaurus and Lufengosaurus to be massospondylids. This result supports many previous analyses that tested fewer taxa. However, this analysis found the two recently described North American massopods, Sarahsaurus and Seitaad, and the South African Ignavusaurus to nest outside Massospondylidae, as opposed to some provisional proposals. Earlier in 2011, Pradhania, a sauropodomorph from India, was tested for the first time in a large cladistic analysis and was found to be a relatively basal massospondylid. Mussaurus and Xixiposaurus may also be included within Massospondylidae.
Anchisauria is an extinct clade of sauropodomorph dinosaurs that lived from the Late Triassic to the Late Cretaceous. The name Anchisauria was first used Haekel and defined by Galton and Upchurch in the second edition of The Dinosauria. It is a node-based taxon containing the most recent common ancestor of Anchisaurus polyzelus and Melanorosaurus readi, and all its descendants. Galton and Upchurch assigned a family of dinosaurs to the Anchisauria: the Melanorosauridae. The more common prosauropods Plateosaurus and Massospondylus were placed in the sister clade Plateosauria.
Glacialisaurus is a genus of sauropodomorph dinosaur. It lived during the Pliensbachian stage of the Early Jurassic period around 186 to 182 million years ago in what is now the central region of the Transantarctic Mountains of Antarctica. It is known from two specimens; the holotype, a partial tarsus (ankle) and metatarsus, and a partial left femur. The fossils were collected by a team led by paleontologist William R. Hammer during a 1990–91 field expedition to the Hanson Formation of Antarctica. They were described in 2007, and made the basis of the new genus and species Glacialisaurus hammeri. The genus name translates as “icy” or "frozen lizard”, and the species name honors Hammer.
Plateosauria is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to the Late Cretaceous. The name Plateosauria was first coined by Gustav Tornier in 1913. The name afterwards fell out of use until the 1980s.
Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, defined as all animals more closely related to Saltasaurus loricatus than to Plateosaurus engelhardti.
Aardonyx is a genus of basal sauropodomorph dinosaur. It is known from the type species Aardonyx celestae found from the Early Jurassic Elliot Formation of South Africa. A. celestae was named after Celeste Yates, who prepared much of the first known fossil material of the species. It has arm features that are intermediate between basal sauropodomorphs and more derived sauropods.
Leonerasaurus is a basal genus of sauropodomorph dinosaur. Currently, there is only one species known, named L. taquetrensis by Diego Pol, Alberto Garrido and Ignacio A. Cerda in 2011. The fossil, an incomplete subadult individual, was found in the Las Leoneras Formation in Argentina. This formation is probably Early Jurassic in age. Leonerasaurus was a small non-sauropod sauropodomorph, showing an unusual combination of basal and derived characters. This indicates that the evolution of early sauropodomorphs witnessed a great degree of convergent evolution.
Meroktenos is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period of what is now Lesotho.
Xingxiulong is a genus of bipedal sauropodiform from the Early Jurassic of China. It contains a single species, X. chengi, described by Wang et al. in 2017 from three specimens, two adults and an immature individual, that collectively constitute a mostly complete skeleton. Adults of the genus measured 4–5 metres (13–16 ft) long and 1–1.5 metres tall. Phylogenetic analysis suggests that Xingxiulong is most closely related to its contemporary Jingshanosaurus, although an alternative position outside of both the Sauropodiformes and Massospondylidae is also plausible.
Ledumahadi is a genus of lessemsaurid sauropodomorph dinosaur from the Early Jurassic Elliot Formation in Free State Province, South Africa. The type and only species is L. mafube, known from a singular incomplete postcranial specimen. A quadruped, it was one of the first giant sauropodomorphs, reaching a weight of around 12 tonnes, despite not having evolved columnar limbs like its later huge relatives.
Irisosaurus is an extinct genus of sauropodiform sauropodomorph dinosaur, from the Fengjiahe Formation of China. The type species, Irisosaurus yimenensis was formally described in 2020. It was the sister taxon to Mussaurus.