Musankwa

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Musankwa
Temporal range: Triassic, Norian
Musankwa holotype (NHMZ 2521).png
Holotype femur (A) and tibia with astragalus (B)
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Massopoda
Genus: Musankwa
Species:
M. sanyatiensis
Binomial name
Musankwa sanyatiensis
Barrett et al., 2024

Musankwa is an extinct genus of massopodan sauropodomorph dinosaur from the Late Triassic (Norian) Pebbly Arkose Formation of Zimbabwe. The genus contains a single species, M. sanyatiensis, known from a partial leg. Musankwa represents the fourth dinosaur genus to be named from Zimbabwe.

Contents

Discovery and naming

Geographical, stratigraphic, and sedimentological details of the Musankwa discovery, including the holotype in situ (E) Musankwa (geography and sedimentology).png
Geographical, stratigraphic, and sedimentological details of the Musankwa discovery, including the holotype in situ (E)
Skeletal diagram of the holotype Musankwa Skeletal.svg
Skeletal diagram of the holotype

The Musankwa holotype specimen, NHMZ 2521, was discovered in March 2018 by Paul Barrett in sediments of the Pebbly Arkose Formation of the Upper Karoo Group (Spurwing East Palaeosol site) on the shoreline of Spurwing Island in Lake Kariba, Zimbabwe. The specimen consists of an articulated partial right leg, including the femur, tibia, and astragalus, in addition to associated unidentifiable bone fragments. A fragment of the distal end of the fibula was also found in the field but lost during collection and preparation before its description. [1]

The Musankwa fossil material was mentioned in a 2020 review of the Upper Karoo Group's geology and paleontology, where it was preliminarily identified as belonging to a non-dinosaurian archosauromorph. [2] The material later was then alluded to in a 2023 conference abstract as belonging to a Riojasaurus -like sauropodomorph, before its formal description the following year. [3]

In 2024, Barrett et al. described Musankwa sanyatiensis as a new genus and species of basal sauropodomorph based on these fossil remains. The generic name, Musankwa, honors the house boat of the same name that served as the paleontologists' laboratory and home while conducting the fieldwork during which the holotype was found. "Musankwa" is a Tonga word meaning "boy close to marriage". The specific name, sanyatiensis, references the Sanyati River which flows into Lake Kariba near the type locality. [1]

Musankwa is the fourth genus of dinosaurs to be named from Zimbabwe, [1] following "Syntarsus" rhodesiensis in 1969, [4] Vulcanodon in 1972, [5] and Mbiresaurus in 2022. [6]

Description

Based on more complete relatives, Musankwa was likely an obligate biped. As such, the body mass of the holotype individual was calculated at about 386 kilograms (851 lb). [1]

Classification

In their phylogenetic analyses, Barrett et al. (2024) recovered Musankwa as the basalmost member of the sauropodomorph clade Massopoda, with the Riojasauridae found to be the next diverging clade. Their results are displayed in the cladogram below: [1]

Massopoda

Related Research Articles

<span class="mw-page-title-main">Sauropodomorpha</span> Extinct clade of dinosaurs

Sauropodomorpha is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The prosauropods, which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic until their decline and extinction at the end of the Cretaceous.

<i>Melanorosaurus</i> Sauropodomorph dinosaur genus from the Late Triassic period

Melanorosaurus is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. A omnivore from South Africa, it had a large body and sturdy limbs, suggesting it moved about on all fours. Its limb bones were massive and heavy like the limb bones of true sauropods.

<i>Massospondylus</i> Sauropodomorph dinosaur genus from Early Jurassic South Africa and Botswana

Massospondylus was a genus of sauropodomorph dinosaur from the Early Jurassic. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been named. Fossils have since been found at other locations in South Africa, Lesotho, and Zimbabwe. Material from Arizona's Kayenta Formation, India, and Argentina has been assigned to the genus at various times, but the Arizonan and Argentinian material are now assigned to other genera.

Basutodon was a genus of suchian archosaur from the late Carnian-early Norian-age Upper Triassic Lower Elliot Formation of Lesotho. The type species is B. ferox.

<i>Hortalotarsus</i> Extinct genus of reptiles

Hortalotarsus is a dubious genus of extinct sauropodomorph from the Early Jurassic (Sinemurian-aged) Clarens Formation of Eagle's Crag, South Africa.

<span class="mw-page-title-main">Massospondylidae</span> Extinct family of dinosaurs

Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and Antarctica during the Late Triassic to the Early Jurassic periods. Several dinosaurs have been classified as massospondylids over the years. The largest cladistic analysis of early sauropodomorphs, which was presented by Apaldetti and colleagues in November 2011, found Adeopapposaurus, Coloradisaurus, Glacialisaurus, Massospondylus, Leyesaurus and Lufengosaurus to be massospondylids. This result supports many previous analyses that tested fewer taxa. However, this analysis found the two recently described North American massopods, Sarahsaurus and Seitaad, and the South African Ignavusaurus to nest outside Massospondylidae, as opposed to some provisional proposals. Earlier in 2011, Pradhania, a sauropodomorph from India, was tested for the first time in a large cladistic analysis and was found to be a relatively basal massospondylid. Mussaurus and Xixiposaurus may also be included within Massospondylidae.

<span class="mw-page-title-main">Forest Sandstone</span> Geologic formation in Southern Africa

The Forest Sandstone is a geological formation in southern Africa, dating to roughly between 200 and 190 million years ago and covering the Hettangian to Sinemurian stages of the Jurassic Period in the Mesozoic Era. As its name suggests, it consists mainly of sandstone.

<span class="mw-page-title-main">Massopoda</span> Extinct clade of dinosaurs

Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, defined as all animals more closely related to Saltasaurus loricatus than to Plateosaurus engelhardti.

<span class="mw-page-title-main">Elliot Formation</span> Lithostratigraphic layer of the Stormberg Group in South Africa

The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, southern Free State, and in the eastern KwaZulu-Natal provinces of South Africa. Outcrops and exposures are also found in several localities in Lesotho such as Qacha's Neck, Hill Top, Quthing, and near the capital, Maseru. The Elliot Formation is further divided into the lower (LEF) and upper (UEF) Elliot formations to differentiate significant sedimentological differences between these layers. The LEF is dominantly Late Triassic (Norian-Hettangian) in age while the UEF is mainly Early Jurassic (Sinemurian-Pliensbachian) and is tentatively regarded to preserve a continental record of the Triassic-Jurassic boundary in southern Africa. This geological formation is named after the town of Elliot in the Eastern Cape, and its stratotype locality is located on the Barkly Pass, 9 km north of the town.

<span class="mw-page-title-main">Batoka Formation</span> Geological formation in Southern Africa

The Batoka Formation is a geological formation in the Zambezi valley in Botswana, Zambia and Zimbabwe. It is predominantly a volcanic unit comprising mainly basalts. It was formerly thought to contain sand stones containing the dinosaur Vulcanodon, however this was shown to be in error resulting from interpreting folding of the rocks as separate layers, with the sandstone layers actually being from the underlying Forest Sandstone.

Notochampsa is an extinct genus of protosuchian crocodyliform. Fossils have been found from the lower Clarens Formation of the Karoo Supergroup in South Africa, dating back to the Pliensbachian stage of the Early Jurassic. Notochampsa comes from a period of relative fossil scarcity, and is the youngest known occurrence of a crocodylomorph from the Karoo Basin of South Africa.

<span class="mw-page-title-main">Pebbly Arkose Formation</span>

The Pebbly Arkose Formation is a Late Triassic geologic formation found in southern Africa.

<span class="mw-page-title-main">Stormberg Group</span> Triassic/Jurassic geological group in the Karoo Supergroup in South Africa

The Stormberg Group is one of the four geological groups that comprises the Karoo Supergroup in South Africa. It is the uppermost geological group representing the final phase of preserved sedimentation of the Karoo Basin. The Stormberg Group rocks are considered to range between Lower Triassic (Olenekian) to Lower Jurassic (Pliensbachian) in age. These estimates are based on means of geological dating including stratigraphic position, lithostratigraphic and biostratigraphic correlations, and palynological analyses.

<span class="mw-page-title-main">Angwa Sandstone</span>

The Angwa Sandstone is a geological formation of the mid-Triassic Cabora Bassa Basin and Mana Pools Basin of southern Africa, consisting mainly of sandstone.

<i>Megapnosaurus</i> Extinct genus of dinosaur

Megapnosaurus is an extinct genus of coelophysid theropod dinosaur that lived approximately 188 million years ago during the early part of the Jurassic Period in what is now Africa. The species was a small to medium-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to 2.2 m (7.2 ft) long and weigh up to 13 kg (29 lb). It was originally given the genus name Syntarsus, but that name was later determined to be preoccupied by a beetle. The species was subsequently given a new genus name, Megapnosaurus, by Ivie, Ślipiński & Węgrzynowicz in 2001. Some studies have classified it as a species within the genus Coelophysis, but this interpretation has been challenged by more subsequent studies and the genus Megapnosaurus is now considered valid.

<i>Pentasaurus</i> Genus of dicynodont therapsid from the late Triassic of South Africa

Pentasaurus is an extinct genus of dicynodont of the family Stahleckeriidae, closely related to the well known Placerias. It was found in the Lower Elliot Formation of South Africa, dated to the Norian of the Late Triassic period. The genus contains the type and only species, Pentasaurus goggai. Pentasaurus is named after the ichnogenus Pentasauropus, fossil footprints that were originally described from the lower Elliot Formation in 1970 decades before the body fossils of Pentasaurus itself were recognised. Pentasauropus footprints were likely made by dicynodonts, and in South Africa Pentasaurus itself was the likely trackmaker. The name reflects the fact that a large dicynodont was predicted to have existed in the lower Elliot Formation before any body fossils were recognised, and so Pentasaurus was named after its probable footprints. This is a reversal of the more typical occurrence where fossil footprints are named after their presumed trackmakers. The name of the species honours its collector Alfred Brown, nicknamed "Gogga", which means "bug" in Afrikaans.

<span class="mw-page-title-main">Lessemsauridae</span> Extinct clade of dinosaurs

Lessemsauridae is a clade of early sauropodiform dinosaurs that lived in the Triassic and Jurassic of Argentina, South Africa and possibly Lesotho. A phylogenetic analysis performed by Apaldetti and colleagues in 2018 recovered a new clade of sauropodiforms uniting Lessemsaurus, Antetonitrus, and Ingentia which they named Lessemsauridae. It is a node-based taxon, defined as all descendants of the most recent common ancestor of Lessemsaurus sauropoides and Antetonitrus ingenipes. Depending on the definition of Sauropoda, Lessemsauridae is either one of the most basal sauropod taxa, or a sister taxon of Sauropoda. An additional member of the clade was named later in 2018, Ledumahadi. A 2021 study by Pol and colleagues also assigned the genera Kholumolumo and Meroktenos to the group.

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Mbiresaurus is an extinct genus of basal sauropodomorph dinosaur from the Late Triassic (Carnian) Pebbly Arkose Formation of Zimbabwe. The genus contains a single species, Mbiresaurus raathi, known from a nearly complete skeleton. Mbiresaurus represents one of Africa’s earliest known definitive dinosaurs.

References

  1. 1 2 3 4 5 Barrett, Paul M.; Chapelle, Kimberley E.J.; Sciscio, Lara; Broderick, Timothy J.; Zondo, Michel; Munyikwa, Darlington; Choiniere, Jonah N. "A new Late Triassic sauropodomorph dinosaur from the Mid-Zambezi Basin, Zimbabwe". Acta Palaeontologica Polonica . 69 (2): 227–241. doi: 10.4202/app.01100.2023 .
  2. Sciscio, Lara; Viglietti, Pia A.; Barrett, Paul M.; Broderick, Timothy J.; Munyikwa, Darlington; Chapelle, Kimberley E. J.; Dollman, Kathleen N.; Edwards, Steve F.; Zondo, Michel; Choiniere, Jonah N. (2020-10-26). "Sedimentology and palaeontology of the Upper Karoo Group in the Mid-Zambezi Basin, Zimbabwe: new localities and their implications for interbasinal correlation". Geological Magazine . 158 (6): 1035–1058. Bibcode:2021GeoM..158.1035S. doi:10.1017/S0016756820001089. ISSN   0016-7568.
  3. Barrett, Paul M.; Sciscio, Lara; Zondo, Michel; Broderick, Timothy J.; Munyikwa, Darlington; Viglietti, Pia A.; Edwards, Steve F.; Chapelle, Kimberley E. J.; Dollman, Kathleen N.; Choiniere, Jonah N. (2023). Faunal change across the Triassic–Jurassic boundary: New insights from the Mid-Zambezi Basin of Zimbabwe. 14th Symposium on Mesozoic Terrestrial Ecosystems and Biota. Vol. 306. pp. 30–32. doi: 10.1002/ar.25219 .
  4. Raath, Michael A. (1969). "A new Coelurosaurian dinosaur from the Forest Sandstone of Rhodesia". Arnoldia . 4 (28): 1–25.
  5. Raath, Michael A. (1972). "Fossil vertebrate studies in Rhodesia: a new dinosaur (Reptilia, Saurischia) from near the Triassic-Jurassic boundary". Arnoldia . 5: 1–2, 4.
  6. Griffin, Christopher T.; Wynd, Brenen M.; Munyikwa, Darlington; Broderick, Tim J.; Zondo, Michel; Tolan, Stephen; Langer, Max C.; Nesbitt, Sterling J.; Taruvinga, Hazel R. (2022-08-31). "Africa's oldest dinosaurs reveal early suppression of dinosaur distribution". Nature. 609 (7926): 313–319. Bibcode:2022Natur.609..313G. doi:10.1038/s41586-022-05133-x. ISSN   0028-0836. PMID   36045297. S2CID   251977824.