Musankwa Temporal range: Triassic, | |
---|---|
Holotype femur (A) and tibia with astragalus (B) | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Massopoda |
Genus: | † Musankwa |
Species: | †M. sanyatiensis |
Binomial name | |
†Musankwa sanyatiensis Barrett et al., 2024 | |
Musankwa is an extinct genus of massopodan sauropodomorph dinosaur from the Late Triassic (Norian) Pebbly Arkose Formation of Zimbabwe. The genus contains a single species, M. sanyatiensis, known from a partial leg. Musankwa represents the fourth dinosaur genus to be named from Zimbabwe.
The Musankwa holotype specimen, NHMZ 2521, was discovered in March 2018 by Paul Barrett in sediments of the Pebbly Arkose Formation of the Upper Karoo Group (Spurwing East Palaeosol site) on the shoreline of Spurwing Island in Lake Kariba, Zimbabwe. The specimen consists of an articulated partial right leg, including the femur, tibia, and astragalus, in addition to associated unidentifiable bone fragments. A fragment of the distal end of the fibula was also found in the field but lost during collection and preparation before its description. [1]
The Musankwa fossil material was mentioned in a 2020 review of the Upper Karoo Group's geology and paleontology, where it was preliminarily identified as belonging to a non-dinosaurian archosauromorph. [2] The material later was then alluded to in a 2023 conference abstract as belonging to a Riojasaurus -like sauropodomorph, before its formal description the following year. [3]
In 2024, Barrett et al. described Musankwa sanyatiensis as a new genus and species of basal sauropodomorph based on these fossil remains. The generic name, Musankwa, honors the house boat of the same name that served as the paleontologists' laboratory and home while conducting the fieldwork during which the holotype was found. "Musankwa" is a Tonga word meaning "boy close to marriage". The specific name, sanyatiensis, references the Sanyati River which flows into Lake Kariba near the type locality. [1]
Musankwa is the fourth genus of dinosaurs to be named from Zimbabwe, [1] following "Syntarsus" rhodesiensis in 1969, [4] Vulcanodon in 1972, [5] and Mbiresaurus in 2022. [6]
Based on more complete relatives, Musankwa was likely an obligate biped. As such, the body mass of the holotype individual was calculated at about 386 kilograms (851 lb). [1]
In their phylogenetic analyses, Barrett et al. (2024) recovered Musankwa as the basalmost member of the sauropodomorph clade Massopoda, with the Riojasauridae found to be the next diverging clade. Their results are displayed in the cladogram below: [1]
Melanorosaurus is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. An omnivore from South Africa, it had a large body and sturdy limbs, suggesting it moved quadrupedally. Its limb bones were massive and heavy like the limb bones of true sauropods.
Vulcanodon is an extinct genus of sauropod dinosaur from the Early Jurassic Uppermost Forest Sandstone of southern Africa. The only known species is V. karibaensis. Discovered in 1969 in Zimbabwe, it was regarded as the earliest-known sauropod for decades, and is still one of the most primitive sauropods that has been discovered. As a quadrupedal, ground-dwelling herbivore, Vulcanodon already showed the typical sauropod body plan with column-like legs and a long neck and tail. It was smaller than most other sauropods, measuring approximately eleven metres (36 ft) in length. Vulcanodon is known from a fragmentary skeleton including much of the pelvic girdle, hindlimbs, forearms, and tail, but lacking the trunk and neck vertebrae as well as the skull.
Basutodon was a genus of suchian archosaur from the late Carnian-early Norian-age Upper Triassic Lower Elliot Formation of Lesotho. The type species is B. ferox.
Hortalotarsus is a dubious genus of extinct sauropodomorph from the Early Jurassic (Sinemurian-aged) Clarens Formation of Eagle's Crag, South Africa.
Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and Antarctica during the Late Triassic to the Early Jurassic periods. Several dinosaurs have been classified as massospondylids over the years. The largest cladistic analysis of early sauropodomorphs, which was presented by Apaldetti and colleagues in November 2011, found Adeopapposaurus, Coloradisaurus, Glacialisaurus, Massospondylus, Leyesaurus and Lufengosaurus to be massospondylids. This result supports many previous analyses that tested fewer taxa. However, this analysis found the two recently described North American massopods, Sarahsaurus and Seitaad, and the South African Ignavusaurus to nest outside Massospondylidae, as opposed to some provisional proposals. Earlier in 2011, Pradhania, a sauropodomorph from India, was tested for the first time in a large cladistic analysis and was found to be a relatively basal massospondylid. Mussaurus and Xixiposaurus may also be included within Massospondylidae.
The Forest Sandstone is a geological formation in southern Africa, dating to roughly between 200 and 190 million years ago and covering the Hettangian to Sinemurian stages of the Jurassic Period in the Mesozoic Era. As its name suggests, it consists mainly of sandstone.
Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, defined as all animals more closely related to Saltasaurus loricatus than to Plateosaurus engelhardti. Sauropodiformes is a more exclusive stem-based clade within Massopoda, defined as "the most inclusive clade containing Saltasaurus but not Massospondylus".
Ferganoceratodus is a genus of prehistoric lungfish known from the Mesozoic of Asia and Africa. Based on morphological evidence, it has either been recovered as a basal member of the Ceratodontiformes or to be the sister group of the Neoceratodontidae.
The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, southern Free State, and in the eastern KwaZulu-Natal provinces of South Africa. Outcrops and exposures are also found in several localities in Lesotho such as Qacha's Neck, Hill Top, Quthing, and near the capital, Maseru. The Elliot Formation is further divided into the lower (LEF) and upper (UEF) Elliot formations to differentiate significant sedimentological differences between these layers. The LEF is dominantly Late Triassic (Norian-Hettangian) in age while the UEF is mainly Early Jurassic (Sinemurian-Pliensbachian) and is tentatively regarded to preserve a continental record of the Triassic-Jurassic boundary in southern Africa. This geological formation is named after the town of Elliot in the Eastern Cape, and its stratotype locality is located on the Barkly Pass, 9 km north of the town.
The Batoka Formation is a geological formation in the Zambezi valley in Botswana, Zambia and Zimbabwe. It is predominantly a volcanic unit comprising mainly basalts. It was formerly thought to contain sand stones containing the dinosaur Vulcanodon, however this was shown to be in error resulting from interpreting folding of the rocks as separate layers, with the sandstone layers actually being from the underlying Forest Sandstone.
Notochampsa is an extinct genus of protosuchian crocodyliform. Fossils have been found from the lower Clarens Formation of the Karoo Supergroup in South Africa, dating back to the Pliensbachian stage of the Early Jurassic. Notochampsa comes from a period of relative fossil scarcity, and is the youngest known occurrence of a crocodylomorph from the Karoo Basin of South Africa.
The Pebbly Arkose Formation is a Late Triassic geologic formation found in southern Africa.
The Stormberg Group is one of the four geological groups that comprises the Karoo Supergroup in South Africa. It is the uppermost geological group representing the final phase of preserved sedimentation of the Karoo Basin. The Stormberg Group rocks are considered to range between Lower Triassic (Olenekian) to Lower Jurassic (Pliensbachian) in age. These estimates are based on means of geological dating including stratigraphic position, lithostratigraphic and biostratigraphic correlations, and palynological analyses.
The Angwa Sandstone is a geological formation of the mid-Triassic Cabora Bassa Basin and Mana Pools Basin of southern Africa, consisting mainly of sandstone.
Pentasaurus is an extinct genus of dicynodont of the family Stahleckeriidae, closely related to the well known Placerias. It was found in the Lower Elliot Formation of South Africa, dated to the Norian of the Late Triassic period. The genus contains the type and only species, Pentasaurus goggai. Pentasaurus is named after the ichnogenus Pentasauropus, fossil footprints that were originally described from the lower Elliot Formation in 1970 decades before the body fossils of Pentasaurus itself were recognised. Pentasauropus footprints were likely made by dicynodonts, and in South Africa Pentasaurus itself was the likely trackmaker. The name reflects the fact that a large dicynodont was predicted to have existed in the lower Elliot Formation before any body fossils were recognised, and so Pentasaurus was named after its probable footprints. This is a reversal of the more typical occurrence where fossil footprints are named after their presumed trackmakers. The name of the species honours its collector Alfred Brown, nicknamed "Gogga", which means "bug" in Afrikaans.
Lessemsauridae is a clade of early sauropodiform dinosaurs that lived in the Triassic and Jurassic of Argentina, South Africa and possibly Lesotho. A phylogenetic analysis performed by Apaldetti and colleagues in 2018 recovered a new clade of sauropodiforms uniting Lessemsaurus, Antetonitrus, and Ingentia which they named Lessemsauridae. It is a node-based taxon, defined as all descendants of the most recent common ancestor of Lessemsaurus sauropoides and Antetonitrus ingenipes. Depending on the definition of Sauropoda, Lessemsauridae is either one of the most basal sauropod taxa, or a sister taxon of Sauropoda. An additional member of the clade was named later in 2018, Ledumahadi. A 2021 study by Pol and colleagues also assigned the genera Kholumolumo and Meroktenos to the group.
Ngwevu is a genus of massospondylid sauropodomorph dinosaur from the Lower Jurassic of South Africa. The genus contains one species, Ngwevu intloko.
Scalenodontoides is an extinct genus of Traversodontidae, a family of herbivorous cynodonts. It lived during the Late Triassic in what is now South Africa. Its type species is Scalenodontoides macrodontes. It was named in 1957 by A. W. Crompton and F. Ellenberger. Arctotraversodon plemmyrodon was originally classified as a species of Scalenodontoides, but was given its own genus in 1992. It is found in the Scalenodontoides Assemblage Zone of the Elliot Formation, which is named for it. It is one of the geologically youngest traversodontids, alongside the putative traversodontid Boreogomphodon. It is closely related to Exaeretodon and Siriusgnathus, but is distinguished by the presence of a shelf-like expansion of its parietal called the nuchal table. Though the largest known complete skull is only 248 millimetres (9.8 in) long, it may have been the largest non-mammalian cynodont, as an incomplete snout would have belonged to a specimen with an estimated skull length of 617 millimetres (24.3 in).
Mbiresaurus is an extinct genus of basal sauropodomorph dinosaur from the Late Triassic (Carnian) Pebbly Arkose Formation of Zimbabwe. The genus contains a single species, Mbiresaurus raathi, known from a nearly complete skeleton. Mbiresaurus represents one of Africa’s earliest known definitive dinosaurs.