Hamititan Temporal range: Early Cretaceous, | |
---|---|
Skeletal diagram showing known material | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Macronaria |
Clade: | † Titanosauria |
Genus: | † Hamititan Wang et al., 2021 |
Type species | |
†Hamititan xinjiangensis Wang et al., 2021 |
Hamititan (meaning "Hami giant") is a genus of titanosaur sauropod dinosaur from the Shengjinkou Formation of Xinjiang, China. It contains one species, the type species, Hamititan xinjiangensis. [1]
In 2006, a Konservat-Lagerstätte was reported from the Shengjinkou Formation in the Hami region of Xinjiang, China. This consisted of lake sediments allowing for exceptional preservation of fossils. The same year, Qiu Zhanxiang and Wang Banyue started official excavations.
Among the excavated fossils were seven caudal vertebrae with three chevrons preserved. These were established as the holotype (HM V22) of the new sauropod taxon, Hamititan. Four sacral elements, specimen IVPP V27875, were not referred. A theropod tooth was found near the sixth caudal vertebra of the holotype. It is likely that this theropod preyed on the holotype shortly after it died. The generic name refers to the city of Hami, where the holotype was found, while the specific name, xinjiangensis, refers to the Chinese province, Xinjiang. [1]
The phylogenetic analysis of Wang et al. places Hamititan as a derived non-lithostrotian titanosaur in a polytomy with Epachthosaurus , Kaijutitan , Notocolossus , rinconsaurs, lognkosaurs, and lithostrotians. If it is scored with the contemporary Silutitan as a single taxon, the combined taxon will be sister to Euhelopus . A cladogram depicting the former hypothesis is shown below: [1]
Other animals from the holotype locality include the pterosaur Hamipterus and contemporary euhelopodid Silutitan , which is described in the same paper. Together, Hamititan, the aforementioned fauna and an unnamed theropod represent the known vertebrate taxa of the area. [1]
Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan, estimated at 37 m (121 ft) long with a weight of 69 tonnes, and the comparably-sized Argentinosaurus and Puertasaurus from the same region.
Magyarosaurus is a genus of dwarf sauropod dinosaur from late Cretaceous Period in Romania. It is one of the smallest-known adult sauropods, measuring only 6 m (20 ft) in length and 750–1,000 kg (1,650–2,200 lb) in body mass. The type and only certain species is Magyarosaurus dacus. It has been found to be a close relative of Rapetosaurus in the family Saltasauridae in the sauropod clade Titanosauria in a 2005 study.
Hypselosaurus is a dubious genus of titanosaurian sauropod that lived in southern France during the Late Cretaceous, approximately 70 million years ago in the early Maastrichtian. Hypselosaurus was first described in 1846, but was not formally named until 1869, when Phillip Matheron named it under the binomial Hypselosaurus priscus. The holotype specimen includes a partial hindlimb and a pair of caudal vertebrae, and two eggshell fragments were found alongside these bones. Because of the proximity of these eggshells to the fossil remains, many later authors, including Matheron and Paul Gervais, have assigned several eggs from the same region of France all to Hypselosaurus, although the variation and differences between these eggs suggest that they do not all belong to the same taxon. Hypselosaurus has been found in the same formation as the dromaeosaurids Variraptor and Pyroraptor, the ornithopod Rhabdodon, and the ankylosaurian Rhodanosaurus, as well as indeterminate bones from other groups.
Epachthosaurus was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America.
Baurutitan is a genus of sauropod dinosaur that lived during the Late Cretaceous in what is now Brazil. The type species, Baurutitan britoi, was described in 2005 by Kellner and colleagues, although the fossil remains had already been discovered in 1957. Baurutitan is classified as a lithostrotian titanosaur, and is distinguished from related genera based on its distinctive caudal vertebrae. This South American dinosaur was found in the Serra da Galga Formation near Uberaba, in the Brazilian state of Minas Gerais.
Pellegrinisaurus is a genus of titanosaurian sauropod dinosaur that lived in South America during the Late Cretaceous period. The holotype was found in the Allen Formation, Argentina.
Phuwiangosaurus is a genus of titanosaur dinosaur from the Early Cretaceous (Valanginian-Hauterivian) Sao Khua Formation of Thailand. The type species, P. sirindhornae, was described by Martin, Buffetaut, and Suteethorn in a 1993 press release and was formally named in 1994. The species was named to honor Princess Maha Chakri Sirindhorn of Thailand, who was interested in the geology and palaeontology of Thailand, while the genus was named after the Phu Wiang area, where the fossil was discovered. The precise affinities of Phuwiangosaurus among titanosauriform sauropods are uncertain, though several studies have classified it as a euhelopodid.
Sonidosaurus is a genus of sauropod dinosaur from the Late Cretaceous. It was a titanosaur which lived in what is now Inner Mongolia. The type species, Sonidosaurus saihangaobiensis, was described by Xu, Zhang, Tan, Zhao, and Tan in 2006. It was a small titanosaur, about 9 meters (30 ft) long. It was first discovered in the Saihangaobi, Iren Dabasu (Erlian) Formation, in 2001 in a quarry which would later yield the remains of Gigantoraptor.
Euhelopodidae is a family of sauropod dinosaurs of disputed membership and affinities, which contains Euhelopus and its close relatives. Most proposed euhelopodids are from East Asia.
Wintonotitan is a genus of titanosauriform dinosaur from Cenomanian -age Winton Formation of Australia. It is known from partial postcranial remains.
Xianshanosaurus is a genus of sauropod dinosaur from the Early Cretaceous (Aptian-Albian) of the Ruyang Basin in Henan Province, China. Its type and only species is Xianshanosaurus shijiagouensis. It was described in 2009 by a team of paleontologists led by Lü Junchang. Xianshanosaurus may be a titanosaur, and Daxiatitan may be its closest relative, but its evolutionary relationships remain controversial.
Brasilotitan is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Adamantina Formation of Brazil. The type species is Brasilotitan nemophagus. Brasilotitan was a small titanosaur with a squared-off snout, and may be closely related to another Brazilian titanosaur, Uberabatitan.
Hamipterus is an extinct genus of pteranodontoid pterosaurs from the Early Cretaceous Shengjinkou Formation of northwestern China. It is known from a single species, the type species, H. tianshanensis.
Savannasaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Winton Formation of Queensland, Australia. It contains one species, Savannasaurus elliottorum, named in 2016 by Stephen Poropat and colleagues. The holotype and only known specimen, originally nicknamed "Wade", is the most complete specimen of an Australian sauropod, and is held at the Australian Age of Dinosaurs museum. Dinosaurs known from contemporary rocks include its close relative Diamantinasaurus and the theropod Australovenator; associated teeth suggest that Australovenator may have fed on the holotype specimen.
Dzharatitanis is a genus of sauropod from the Bissekty Formation in Uzbekistan, dating to the Turonian age of the Late Cretaceous. The genus contains a single species, Dzharatitanis kingi, named after geologist Christopher King, who contributed to the Cretaceous geology of Asia. It is currently one of two known sauropods from the Bissekty Formation, alongside an indeterminate titanosaur. In its original publication it was considered to be a member of Rebbachisauridae, but later papers considered it to be a titanosaur.
Silutitan is a genus of euhelopodid sauropod dinosaur from the Shengjinkou Formation of Xinjiang, China. It contains only the type species, Silutitan sinensis.
The Shengjinkou Formation is an Early Cretaceous (Aptian)-aged Konservat-Lagerstätte composed of "interbedded red green and yellow variegated mudstones and siltstones" that is part of the larger Tugulu Group of China. Dinosaur and pterosaur remains have been recovered from the formation.
Elemgasem is an extinct genus of brachyrostran abelisaurid from the Late Cretaceous Portezuelo Formation of Patagonia, Argentina. The genus contains a single species, Elemgasem nubilis. The cladistic position of Elemgasem within Brachyrostra is uncertain, given that phylogenetic analyses recover it as either a sister taxon to Furileusauria or in several positions within this clade.
Ruixinia is an extinct genus of somphospondylan titanosauriform dinosaur from the Early Cretaceous (Barremian) Yixian Formation of China. The genus contains a single species, Ruixinia zhangi. The Ruixinia holotype is a partial articulated skeleton with the most complete series of caudal vertebrae known from any Asian titanosauriform.