| Brohisaurus Temporal range: Late Jurassic, | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Clade: | Dinosauria |
| Genus: | † Brohisaurus Malkani, 2003 |
| Species: | †B. kirthari |
| Binomial name | |
| †Brohisaurus kirthari Malkani, 2003 | |
Brohisaurus is a problematic genus of dubious dinosaurs known from the Late Jurassic (Kimmeridgian-aged) Sembar Formation of Pakistan. The type species is B. kirthari.
The Brohisaurus holotype, MSM-86-K to 94-K, 96-K to 98-K, 101-K to 105-K, largely consisting of indeterminate fragments of some ribs, vertebrae, and limb bones, was discovered in the Sembar Formation of Pakistan. In 2003 Muhammad Sadiq Malkani named and described Brohisaurus kirthari as a new genus and species of titanosaurian sauropods. [1]
The genus name means "Brohi lizard" and refers to the Brohi people who live in the area where it was found. The species name refers to the Kirthar Mountains. [1]
If Brohisaurus is a sauropod, as originally identified, it would have been a large-bodied, long-necked herbivore. Its femur was 12 centimetres (4.7 in) in diameter. [1] The femur of the 15–20 metres (49–66 ft)-long titanosauriform Phuwiangosaurus , by contrast, is 20 centimetres (7.9 in) in diameter. [2]
Brohisaurus was originally described as a titanosaur. [1] Malkani suggested it was similar to the early African titanosaur Malawisaurus , arguing that this provided evidence for a biogeographic link between the Indian subcontinent and Africa. However, the phylogenetic position of Brohisaurus is far from clear. None of the proposed traits uniting it to Titanosauria are definitive synapomorphies of that clade, and none of the fossil material is reliably diagnostic, rendering it a dubious taxon. In 2013, Mannion et al. identified two possible synapomorphies of the Titanosauriformes in the type material: pneumatic cavities in its thoracic ribs and femora with elliptical cross sections. [3]
In a 2025 paper published in Scientific Research Publishing, a known predatory publisher, Malkani revisited Brohisaurus and proposed that it could be allied with the Ankylosauridae based on the presence of several osteoderms apparently referable to this taxon. [4]
Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan, estimated at 37 m (121 ft) long with a weight of 69 tonnes, and the comparably-sized Argentinosaurus and Puertasaurus from the same region.
The Brachiosauridae are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the leaves of tall trees that other sauropods would have been unable to reach. In addition, they possessed thick spoon-shaped teeth which helped them to consume tough plants more efficiently than other sauropods. They have also been characterized by a few unique traits or synapomorphies; dorsal vertebrae with 'rod-like' transverse processes and an ischium with an abbreviated pubic peduncle.
Antarctosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. The type species, Antarctosaurus wichmannianus, and a second species, Antarctosaurus giganteus, were described by prolific German paleontologist Friedrich von Huene in 1929. Three additional species of Antarctosaurus have been named since then but later studies have considered them dubious or unlikely to pertain to the genus.
Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located. Fossil evidence suggests that macronarian dinosaurs lived from the Middle Jurassic (Bathonian) through the Late Cretaceous (Maastrichtian). Macronarians have been found globally, including discoveries in Argentina, the United States, Portugal, China, and Tanzania. Like other sauropods, they are known to have inhabited primarily terrestrial areas, and little evidence exists to suggest that they spent much time in coastal environments. Macronarians are diagnosed through their distinct characters on their skulls, as well as appendicular and vertebral characters. Macronaria is composed of several subclades and families notably including Camarasauridae and Titanosauriformes, among several others. Titanosauriforms are particularly well known for being some of the largest terrestrial animals to ever exist.
Andesaurus is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Earth, it would have had a small head on the end of a long neck and an equally long tail.
Huabeisaurus was a genus of dinosaur from the Late Cretaceous. It was a sauropod which lived in what is present-day northern China. The type species, Huabeisaurus allocotus, was first described by Pang Qiqing and Cheng Zhengwu in 2000. Huabeisaurus is known from numerous remains found in the 1990s, which include teeth, partial limbs and vertebrae. Due to its relative completeness, Huabeisaurus represents a significant taxon for understanding sauropod evolution in Asia. Huabeisaurus comes from Kangdailiang and Houyu, Zhaojiagou Town, Tianzhen County, Shanxi province, China. The holotype was found in the unnamed upper member of the Huiquanpu Formation, which is Late Cretaceous (?Cenomanian–?Campanian) in age based on ostracods, charophytes, and fission-track dating.
Tangvayosaurus is a genus of sauropod dinosaur from the Aptian-Albian age Lower Cretaceous Gres superieurs Formation of Savannakhet Province, Laos. It was a basal somphospondylan, about 15 m long, and is known from the remains of two or three individuals.
Phuwiangosaurus is a genus of titanosaur dinosaur from the Early Cretaceous (Valanginian-Hauterivian) Sao Khua Formation of Thailand. The type species, P. sirindhornae, was described by Martin, Buffetaut, and Suteethorn in a 1993 press release and was formally named in 1994. The species was named to honor Princess Maha Chakri Sirindhorn of Thailand, who was interested in the geology and palaeontology of Thailand, while the genus was named after the Phu Wiang area, where the fossil was discovered. The precise affinities of Phuwiangosaurus among titanosauriform sauropods are uncertain, though several studies have classified it as a euhelopodid.
Huanghetitan, is a genus of sauropod dinosaur from the early Cretaceous Period. It was a basal titanosauriform which lived in what is now Gansu, China.
Fusuisaurus is a genus of sauropod dinosaur from the Early Cretaceous of China. Fragmentary postcranial remains of this animal have been discovered in 2001 in the Napai Formation of Guangxi, China and consist of the left ilium, left pubis, anterior caudals, most of the dorsal ribs and distal end of the left femur. This sauropod has been described as a basal titanosauriform.
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch et al. in 2004 as the most recent common ancestor of Malawisaurus and Saltasaurus and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek lithostros, meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch et al. include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous.
Malarguesaurus is a genus of titanosauriform sauropod dinosaur from the Late Cretaceous of Mendoza Province, Argentina. Its fossils, consisting of tail vertebrae, chevrons, ribs, and limb bones, were found in the upper Turonian-lower Coniacian Portezuelo Formation of the Neuquén Group. The type species, described by González Riga et al. in 2008, is M. florenciae.
Ruyangosaurus is a genus of titanosauriform sauropod dinosaur recovered from the Early Cretaceous Haoling Formation of China. The type species is R. giganteus, described in 2009 by Lü Junchang et al.
Xianshanosaurus is a genus of sauropod dinosaur from the Early Cretaceous (Aptian-Albian) of the Ruyang Basin in Henan Province, China. Its type and only species is Xianshanosaurus shijiagouensis. It was described in 2009 by a team of paleontologists led by Lü Junchang. Xianshanosaurus may be a titanosaur, and Daxiatitan may be its closest relative, but its evolutionary relationships remain controversial.
Mierasaurus is an extinct genus of sauropod dinosaur from the Early Cretaceous of Utah, United States. The taxon was first described and named in 2017 by Rafael Royo-Torres and colleagues, from a mostly complete skeleton including a disarticulated partial skull and mandible, teeth, multiple vertebrae from along the length of the body, both scapulae, radius and ulna bones, a left manus, a complete pelvis, both femora and the entire left hindlimb. Additionally, they referred a lower jaw and femur from juvenile individuals, which were found nearby, to the genus. Collectively, Mierasaurus is among the most completely known North American sauropods. The genus name honours Bernardo de Miera y Pacheco, the first European scientist to enter what is now Utah. The type species for Mierasaurus is Mierasaurus bobyoungi, named after Robert Glen Young, a paleontologist who researched the Early Cretaceous of Utah.
Sibirotitan is a genus of somphospondyl sauropod from the Ilek Formation of Russia. The type and only species is S. astrosacralis.
Wamweracaudia is a large herbivorous sauropod dinosaur from the Late Jurassic Tendaguru Formation of Tanzania, Africa, 155-145 million years ago.
Australotitan is an extinct genus of possibly titanosaurian somphospondylan dinosaurs from the Late Cretaceous Winton Formation (Cenomanian–Turonian) of southern-central Queensland, Australia. The genus contains a single species, A. cooperensis, known from multiple partial skeletons. The genus Australotitan may be synonymous with Diamantinasaurus, a contemporary relative.
Ruixinia is an extinct genus of somphospondylan titanosauriform dinosaur from the Early Cretaceous (Barremian) Yixian Formation of China. The genus contains a single species, Ruixinia zhangi. The Ruixinia holotype is a partial articulated skeleton with the most complete series of caudal vertebrae known from any Asian titanosauriform.
