Kaatedocus Temporal range: Late Jurassic, | |
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Mounted skeleton cast, Museum voor Natuurwetenschappen in Brussels, Belgium | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Superfamily: | † Diplodocoidea |
Clade: | † Flagellicaudata |
Genus: | † Kaatedocus Tschopp & Mateus, 2012 |
Type species | |
†Kaatedocus siberi Tschopp & Mateus, 2012 |
Kaatedocus is a genus of flagellicaudatan sauropod known from the middle Late Jurassic (Kimmeridgian stage) of northern Wyoming, United States. It is known from well-preserved skull and cervical vertebrae which were collected in the lower part of the Morrison Formation. The type and only species is Kaatedocus siberi, described in 2012 by Emanuel Tschopp and Octávio Mateus. [1]
In 1934, a team of the American Museum of Natural History (AMNH) headed by Barnum Brown, financed by the Sinclair Oil Corporation, uncovered about three thousand sauropod bones on the land of rancher Barker Howe near Shell, in Big Horn County. Plans for further excavations in 1935 had to be cancelled after Howe, convinced by the large publicity surrounding the find that the remains were very valuable, demanded higher payment. The bones would not be described and most of them were lost in a fire at the AMNH during the 1940s; others were thrown away in the 1960s after having rotted because of being stowed in a chicken run at Shell. Only about 10% of the fossils survived, among them a skull. They were generally interpreted as belonging to Barosaurus . In 2015, based on specimen-level phylogenetic analysis, one of these specimens (AMNH FARB 7530) was reinterpreted to belong to Kaatedocus. [2]
In 1989 the site was reopened by Hans-Jakob Siber, the founder of the Swiss Aathal Dinosaur Museum. His team immediately adjacent to the old Howe Quarry discovered another 450 bones that became part of the collection of the Swiss museum.
The finds included an exceptionally complete neck, specimen SMA 004 In Switzerland, this became the subject of several lines of scientific investigation. In 2005 Daniela Schwarz studied the pneumatisation of the vertebrae by tomography scanning them with neutrons and X-rays. [3] In 2010 Andreas Christian used the well-preserved vertebrae to support his hypothesis that sauropod necks were held in a rather upright position, [4] which was confirmed by Armin Schmitt studying the vestibular system of Kaatedocus. In 2012 Tschopp used a scan to create a replica of the neck by means of a 3D-printer. [5]
During the intense study of the fossils it became clear that they did not represent Barosaurus but a species new to science. In 2012 this was named Kaatedocus siberi, by the Swiss palaeontologist Emanuel Tschopp, who as a boy had visited the excavations, and his Portuguese colleague Octávio Mateus. The generic name, which means "small beam", combines a reference to the related form Diplodocus with a Crow Indian diminutive suffix ~kaate. The specific name honours Siber. [1]
An additional specimen consisting of a braincase from the Aathal Dinosaur Museum collection (SMA D16-3) was referred to Kaatedocus in 2013, [6] and its referral was corroborated in 2015 based on an exhaustive phylogenetic analysis. [2]
Kaatedocus was relatively small compared to most diplodocids. The type individual was estimated to have been approximately 14 metres (46 ft) long. The combined length of the skull and neck is approximately 3.8 metres (12 ft). The neck was composed of at least 14 vertebrae, comparable to the typical number in diplodocids. [1]
Tschopp, Mateus, and Benson (2015) found Kaatedocus to be an advanced member of the Diplodocinae, closely related to Barosaurus , as shown below. [2]
Diplodocidae |
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On the other hand, Whitlock and Wilson (2020) find Kaatedocus to be the most basal dicraeosaurid, finding such a position more likely than being a diplodocid. The position of Kaatedocus is supported by many cranial features, which are only found within Diplodocoidea. The cladogram of the analysis is shown below: [7]
To support their analysis, Whitlock and Wilson evaluated the characters that Kaatedocus supposedly shared with Diplodocidae: "antorbital fenestra with concave dorsal margin; a 'hooked' prefrontal; box-like basal tubera; and the elongate coel on posterior cervical neural spines." They found the first character to be a more general character diagnostic of Flagellicaudata, because it is currently unknown if dicraeosaurids possessed this feature. They believed that the second character, the hooked prefrontal, was incorrectly interpreted, and that the prefrontal is actually linear, similar to Dicraeosaurus. The presence of a box-like basal tubera was excluded as a character from their analysis because the original intent of the character was to distinguish Diplodocus and Apatosaurus, but in its current form, all other taxa exhibit intermediate morphologies. Lastly, the fourth character was also dismissed because it was unclear in its current definition and varied along the spinal column of all diplodocid taxa. Additionally, they independently test their results by conducting a supplemental analysis utilizing a modified dataset of Mannion et al. (2019) [8] which also recovered Kaatedocus within Dicraeosauridae. [7]
Apatosaurus is a genus of herbivorous sauropod dinosaur that lived in North America during the Late Jurassic period. Othniel Charles Marsh described and named the first-known species, A. ajax, in 1877, and a second species, A. louisae, was discovered and named by William H. Holland in 1916. Apatosaurus lived about 152 to 151 million years ago (mya), during the late Kimmeridgian to early Tithonian age, and are now known from fossils in the Morrison Formation of modern-day Colorado, Oklahoma, New Mexico, Wyoming, and Utah in the United States. Apatosaurus had an average length of 21–23 m (69–75 ft), and an average mass of 16.4–22.4 t. A few specimens indicate a maximum length of 11–30% greater than average and a mass of approximately 33 t.
Supersaurus is a genus of diplodocid sauropod dinosaur that lived in North America during the Late Jurassic period. The type species, S. vivianae, was first discovered by Vivian Jones of Delta, Colorado, in the middle Morrison Formation of Colorado in 1972. The fossil remains came from the Brushy Basin Member of the formation, dating between 153 to 145 million years ago. It was a very large sauropod, with the WDC and BYU specimens reaching 33–35 meters (108–115 ft) in length and approximately 35–40 metric tons in body mass. A potential second species, S. lourinhanensis (Dinheirosaurus), is known from Portugal and has been dated to a similar time.
Camarasaurus was a genus of quadrupedal, herbivorous dinosaurs and is the most common North American sauropod fossil. Its fossil remains have been found in the Morrison Formation, dating to the Late Jurassic epoch, between 155 and 145 million years ago.
Diplodocids, or members of the family Diplodocidae, are a group of sauropod dinosaurs. The family includes some of the longest creatures ever to walk the Earth, including Diplodocus and Supersaurus, some of which may have reached lengths of up to 42 metres (138 ft).
Barosaurus was a giant, long-tailed, long-necked, plant-eating sauropod dinosaur closely related to the more familiar Diplodocus. Remains have been found in the Morrison Formation from the Upper Jurassic Period of Utah and South Dakota. It is present in stratigraphic zones 2–5.
Amargasaurus is a genus of sauropod dinosaur from the Early Cretaceous epoch of what is now Argentina. The only known skeleton was discovered in 1984 and is virtually complete, including a fragmentary skull, making Amargasaurus one of the best-known sauropods of its epoch. Amargasaurus was first described in 1991 and contains a single known species, Amargasaurus cazaui. It was a large animal, but small for a sauropod, reaching 9 to 13 meters in length. Most distinctively, it sported two parallel rows of tall spines down its neck and back, taller than in any other known sauropod. In life, these spines could have stuck out of the body as solitary structures that supported a keratinous sheath. An alternate hypothesis, now more favored, postulates that they could have formed a scaffold supporting a skin sail. They might have been used for display, combat, or defense.
Brontosaurus is a genus of herbivorous sauropod dinosaur that lived in present-day United States during the Late Jurassic period. It was described by American paleontologist Othniel Charles Marsh in 1879, the type species being dubbed B. excelsus, based on a partial skeleton lacking a skull found in Como Bluff, Wyoming. In subsequent years, two more species of Brontosaurus were named: B. parvus in 1902 and B. yahnahpin in 1994. Brontosaurus lived about 156 to 146 million years ago (mya) during the Kimmeridgian and Tithonian ages in the Morrison Formation of what is now Utah and Wyoming. For decades, the animal was thought to have been a taxonomic synonym of its close relative Apatosaurus, but a 2015 study by Emmanuel Tschopp and colleagues found it to be distinct. It has seen widespread representation in popular culture, being the archetypal "long-necked" dinosaur in general media.
Suuwassea is a genus of dicraeosaurid sauropod dinosaur found in the Upper Jurassic strata of the Morrison Formation, located in southern Carbon County, Montana, United States. The fossil remains were recovered in a series of expeditions during a period spanning the years 1999 and 2000 and were described by J.D. Harris and Peter Dodson in 2004. They consist of a disarticulated but associated partial skeleton, including partial vertebral series and limb bones.
Diplodocoidea is a superfamily of sauropod dinosaurs, which included some of the longest animals of all time, including slender giants like Supersaurus, Diplodocus, Apatosaurus, and Amphicoelias. Most had very long necks and long, whip-like tails; however, one family are the only known sauropods to have re-evolved a short neck, presumably an adaptation for feeding low to the ground. This adaptation was taken to the extreme in the highly specialized sauropod Brachytrachelopan. A study of snout shape and dental microwear in diplodocoids showed that the square snouts, large proportion of pits, and fine subparallel scratches in Apatosaurus, Diplodocus, Nigersaurus, and Rebbachisaurus suggest ground-height nonselective browsing; the narrow snouts of Dicraeosaurus, Suuwassea, and Tornieria and the coarse scratches and gouges on the teeth of Dicraeosaurus suggest mid-height selective browsing in those taxa. This taxon is also noteworthy because diplodocoid sauropods had the highest tooth replacement rates of any vertebrates, as exemplified by Nigersaurus, which had new teeth erupting every 30 days.
Dicraeosauridae is a family of diplodocoid sauropods who are the sister group to Diplodocidae. Dicraeosaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as Amargasaurus, Suuwassea, Dicraeosaurus, and Brachytrachelopan. Specimens of this family have been found in North America, Asia, Africa, and South America. In 2023, a dicraeosaurid fossil was discovered in India for the first time. Their temporal range is from the Early or Middle Jurassic to the Early Cretaceous. Few dicraeosaurids survived into the Cretaceous, the youngest of which was Amargasaurus.
Camarasaurus lewisi is a species of sauropod dinosaur from the Upper Jurassic of the United States. It was named by James A. Jensen in 1988. C. lewisi was originally placed in its own genus, Cathetosaurus, but in 1996 it was reclassified as a species of Camarasaurus; most researchers since have considered it to be one of the four valid species of Camarasaurus. Two unpublished studies have since argued that the genus Cathetosaurus should be reinstated, whereas two other studies have argued that C. lewisi may be a junior synonym of another species of Camarasaurus.
Tornieria is a genus of diplodocid sauropod dinosaur from the Late Jurassic in Lindi Region of Tanzania. It has a convoluted taxonomic history.
Brachytrachelopan is a short-necked sauropod dinosaur from the Late Jurassic of Argentina. The holotype and only known specimen was collected from an erosional exposure of fluvial sandstone within the Cañadón Calcáreo Formation on a hill approximately 25 kilometres (16 mi) north-northeast of Cerro Cóndor, Chubut Province, in west-central Argentina, South America. Though very incomplete, the skeletal elements recovered were found in articulation and include eight cervical, twelve dorsal, and three sacral vertebrae, as well as proximal portions of the posterior cervical ribs and all the dorsal ribs, the distal end of the left femur, the proximal end of the left tibia, and the right ilium. Much of the specimen was probably lost to erosion many years before its discovery. The type species is Brachytrachelopan mesai. The specific name honours Daniel Mesa, a local shepherd who discovered the specimen while searching for lost sheep. The genus name translates as "short-necked Pan", Pan being the god of the shepherds.
Dinheirosaurus is a genus of diplodocid sauropod dinosaur that is known from fossils uncovered in modern-day Portugal. It may represent a species of Supersaurus. The only species is Dinheirosaurus lourinhanensis, first described by José Bonaparte and Octávio Mateus in 1999 for vertebrae and some other material from the Lourinhã Formation. Although the precise age of the formation is not known, it can be dated around the early Tithonian of the Late Jurassic.
Apatosaurinae is a subfamily of diplodocid sauropods, an extinct group of large, quadrupedal dinosaurs, the other subfamily in Diplodocidae being Diplodocinae. Apatosaurines are distinguished by their more robust, stocky builds and shorter necks proportionally to the rest of their bodies. Several fairly complete specimens are known, giving a comprehensive view of apatosaurine anatomy.
Diplodocus was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a Neo-Latin term derived from Greek διπλός (diplos) "double" and δοκός (dokos) "beam", in reference to the double-beamed chevron bones located in the underside of the tail, which were then considered unique.
Diplodocinae is an extinct subfamily of diplodocid sauropods that existed from the Late Jurassic to Early Cretaceous of North America, Europe, Africa and South America, about 161.2 to 136.4 million years ago. Genera within the subfamily include Tornieria, Supersaurus, Leinkupal, Galeamopus, Diplodocus, and Barosaurus.
Leinkupal is a genus of diplodocine sauropod known from the Early Cretaceous of the Bajada Colorada Formation, southeastern Neuquén Basin in the Neuquén Province of Argentina. It contains a single species, Leinkupal laticauda.
Galeamopus is a genus of herbivorous diplodocid sauropod dinosaurs. It contains two known species: Galeamopus hayi, known from the Late Jurassic lower Morrison Formation of Wyoming, United States, and Galeamopus pabsti, known from the Late Jurassic fossils from Wyoming and Colorado. The type species is known from one of the most well preserved diplodocid fossils, a nearly complete skeleton with associated skull.
Bajadasaurus is a genus of sauropod dinosaur from the Early Cretaceous epoch of northern Patagonia, Argentina. It was first described in 2019 based on a single specimen found in 2010 that includes a largely complete skull and parts of the neck. The only species is Bajadasaurus pronuspinax. The genus is classified as a member of the Dicraeosauridae, a group of relatively small and short-necked sauropods.