Rinconsaurus

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Rinconsaurus
Temporal range: 86–83  Ma
Rinconsaurus test 2.jpg
Life restoration comparing the animal's size with a human
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Sauropoda
Clade: Macronaria
Clade: Titanosauria
Clade: Rinconsauria
Genus: Rinconsaurus
Calvo & González Riga, 2003
Type species
Rinconsaurus caudamirus
Calvo & González Riga, 2003

Rinconsaurus is a genus of titanosaur sauropod dinosaur from the Late Cretaceous in what is now Argentina. The type species, Rinconsaurus caudamirus, was described by Calvo and Riga in 2003, and is based on three partial skeletons.

Contents

Description

Like all sauropods, Rinconsaurus was a large long-necked quadrupedal animal, with a long, whip-like tail and four pillar-like legs. Rinconsaurus was an unusually slender sauropod. Although fossil discoveries are incomplete, and no complete necks or heads have been found, fully grown Rinconsaurus are estimated to have been 11 meters (36 ft) long and approximately 2.5 meters (8 ft) high at the shoulder. [1] The body mass of Rinconsaurus has been estimated to be between 3.21 and 5.39 tonnes. [2]

Discovery and species

Fossils of Rinconsaurus were discovered in 1997 by Gabriel Benítezin strata belonging to the Bajo de la Carpa Formation near Rincón de los Sauces, in the Neuquen province of Argentina. The fossils, which consisted of three individuals (two adults and one juvenile), were excavated by Jorge Calvo of the Paleontology Museum of the National University of Comahue. [1]

The type, and only known species, Rinconsaurus caudamirus, was described by Calvo and Bernardo J. González Riga of the Paleovertebrate Laboratory in Mendoza, Argentina, in 2003. The authors noted derived traits unique to the genus, including distinctive tail vertebrae with bony processes. The fossil remains included vertebrae, limb bones, scapula, hip bones (pubis, ilium, and ischium) and several ribs. Two teeth are also known. These teeth resemble those of another titanosaurid, Malawisaurus dixeyi . [1] Calvo and González Riga initially interpreted two elements as belonging to the skull of Rinconsaurus, a prefrontal and part of the mandible, but these were subsequently identified as the ilium of the crocodyliform Pehuenchesuchus [3] and part of a rib of Rinconsaurus [4] respectively.

The generic name refers to Rincón de los Sauces, where the fossils were discovered, while the specific name, caudamirus, means "amazing in the tail" in Latin, in reference to the unusual shape of the tail vertebrae, especially a strange sequence of procoelous, amphicoelous, opisthocoelous, biconvex and ultimately again procoelous vertebrae shown by one individual. [1]

Classification

Rinconsaurus is a titanosaurid sauropod. Within the titanosaurs, Calvo and Riga regard this dinosaur as closely related to Aeolosaurus based on several derived traits. [1] In 2007, Casal et al. assigned Rinconsaurus, Gondwanatitan , and Aeolosaurus to the Aeolosauridae, a proposed stem-based clade of titanosaurs. Rinconsaurus would form a smaller clade Rinconsauria with Muyelensaurus . [5] It remains to be seen, however, if this classification will receive widespread acceptance.

Paleobiology

Diet

Rinconsaurus, like all sauropods, was an herbivore. Fossilized dung associated with late Cretaceous titanosaurids has revealed phytoliths, silicified plant fragments, that offer clues to a broad, unselective plant diet. Besides the plant remains that might have been expected, such as cycads and conifers, discoveries published in 2005 revealed an unexpectedly wide range of monocotyledons, including palms and grasses (Poaceae), including ancestors of rice and bamboo, which has given rise to speculation that herbivorous dinosaurs and grasses co-evolved. [6]

Titanosaurid teeth are usually spatulate (spoon-shaped) or shaped like pegs. Teeth found in association with Rinconsaurus have been described as "pencil-chisel-like teeth" and had sharply inclined wear facets. [1] Without the ability to grind food, Rinconsaurus, like other sauropods, would have stripped foliage.

See also

Related Research Articles

<i>Argentinosaurus</i> Late Cretaceous giant sauropod dinosaur genus

Argentinosaurus is a genus of giant sauropod dinosaur that lived during the Late Cretaceous period in what is now Argentina. Although it is only known from fragmentary remains, Argentinosaurus is one of the largest known land animals of all time, perhaps the largest, measuring 30–35 m (98–115 ft) long and weighing 65–80 t. It was a member of Titanosauria, the dominant group of sauropods during the Cretaceous. It is widely regarded by many paleontologists as the biggest dinosaur ever, and perhaps lengthwise the longest animal ever, though both claims have no concrete evidence yet.

<span class="mw-page-title-main">Titanosauria</span> Extinct clade of dinosaurs

Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan, estimated at 37 m (121 ft) long with a weight of 69 tonnes, and the comparably-sized Argentinosaurus and Puertasaurus from the same region.

<i>Aeolosaurus</i> Extinct genus of dinosaurs

Aeolosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. Aeolosaurus is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least two species. However, like most titanosaurs, no remains of the skull are known. The holotype of Aeolosaurus rionegrinus consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago. The species A. maximus was transferred over to the new genus Arrudatitan in 2021.

<i>Bonitasaura</i> Extinct genus of dinosaurs

Bonitasaura is a genus of titanosaurian dinosaur hailing from uppermost layers of the Late Cretaceous (Santonian) Bajo de la Carpa Formation, Neuquén Group of the eastern Neuquén Basin, located in Río Negro Province, Northwestern Patagonia, Argentina. The remains, consisting of a partial sub-adult skeleton jumbled in a small area of fluvial sandstone, including a lower jaw with teeth, a partial vertebrae series, and limb bones, were described by Sebastian Apesteguía in 2004.

<i>Andesaurus</i> Extinct genus of dinosaurs

Andesaurus is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Earth, it would have had a small head on the end of a long neck and an equally long tail.

<i>Epachthosaurus</i> Extinct genus of dinosaurs

Epachthosaurus was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America.

<i>Pellegrinisaurus</i> Extinct genus of dinosaurs

Pellegrinisaurus is a genus of titanosaurian sauropod dinosaur that lived in South America during the Late Cretaceous period. The holotype was found in the Allen Formation, Argentina.

<i>Puertasaurus</i> Extinct genus of dinosaurs

Puertasaurus is a genus of sauropod dinosaur that lived in South America during the Late Cretaceous Period. It is known from a single specimen recovered from sedimentary rocks of the Cerro Fortaleza Formation in southwestern Patagonia, Argentina, which probably is Campanian or Maastrichtian in age. The only species is Puertasaurus reuili. Described by the paleontologist Fernando Novas and colleagues in 2005, it was named in honor of Pablo Puerta and Santiago Reuil, who discovered and prepared the specimen. It consists of four well-preserved vertebrae, including one cervical, one dorsal, and two caudal vertebrae. Puertasaurus is a member of Titanosauria, the dominant group of sauropods during the Cretaceous.

<span class="mw-page-title-main">Lithostrotia</span> Extinct clade of dinosaurs

Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch et al. in 2004 as the most recent common ancestor of Malawisaurus and Saltasaurus and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek lithostros, meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch et al. include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous.

Muyelensaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous of Argentina. It was more slender than other titanosaurs. Fossils have been recovered in the Neuquén province of Patagonia and were originally assigned to the Portezuelo Formation but further research showed that these layers belong to the Plottier Formation. The type species is M. pecheni. The name Muyelensaurus first appeared in a 2007 paper by Argentine paleontologists Jorge Calvo of the Universidad Nacional del Comahue and Bernardo González Riga of the Laboratorio de Paleovertebrados, and Brazilian paleontologist Juan Porfiri of the Universidade Federal do Rio de Janeiro.

Malarguesaurus is a genus of titanosauriform sauropod dinosaur from the Late Cretaceous of Mendoza Province, Argentina. Its fossils, consisting of tail vertebrae, chevrons, ribs, and limb bones, were found in the upper Turonian-lower Coniacian Portezuelo Formation of the Neuquén Group. The type species, described by González Riga et al. in 2008, is M. florenciae.

The Bajo Barreal Formation is a geological formation in the Golfo San Jorge Basin of Chubut and Santa Cruz, Argentina whose strata date back to the Middle Cenomanian to Late Turonian. The formation was first described by Teruggi & Rossetto in 1963. The sandstones, claystones, mudstones, conglomerates and tuff were deposited in a fluvial environment. The upper part of formation is laterally equivalent to the Yacimiento El Trébol and Meseta Espinosa Formation and the lower part to the Laguna Palacios, Cañadón Seco and Comodoro Rivadavia Formations. The Bajo Barreal Formation is a reservoir rock in the Golfo San Jorge Basin.

<span class="mw-page-title-main">Lognkosauria</span> Clade of titanosaurian sauropods

Lognkosauria is a clade of giant long-necked sauropod dinosaurs within the clade Titanosauria. It includes some of the largest and heaviest dinosaurs known. They lived in South America and likely Asia during the Late Cretaceous period.

Traukutitan is a genus of titanosaur sauropod dinosaur which lived during the late Cretaceous.

<span class="mw-page-title-main">Aeolosaurini</span> Extinct clade of dinosaurs

Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found Aeolosaurus, Gondwanatitan, Maxakalisaurus, Panamericansaurus and Rinconsaurus to be aeolosaurids.

<i>Overosaurus</i> Extinct genus of dinosaurs

Overosaurus is an extinct genus of sauropod dinosaurs, containing only a single species, Overosaurus paradasorum. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia. Overosaurus paradasorum was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately 10 m (33 ft). It was a ground-dwelling herbivore.

<i>Patagotitan</i> Extinct genus of dinosaurs

Patagotitan is a genus of titanosaurian sauropod dinosaur from the Cerro Barcino Formation in Chubut Province, Patagonia, Argentina. The genus contains a single species known from at least six young adult individuals, Patagotitan mayorum, which was first announced in 2014 and then named in 2017 by José Carballido and colleagues. Preliminary studies and press releases suggested that Patagotitan was the largest known titanosaur and land animal overall, with an estimated length of 37 m (121 ft) and an estimated weight of 69 tonnes. Later research revised the length estimate down to 31 m (102 ft) and weight estimates down to approximately 50–57 tonnes, suggesting that Patagotitan was of a similar size to, if not smaller than, its closest relatives Argentinosaurus and Puertasaurus. Still, Patagotitan is one of the most-known titanosaurs, and so its interrelationships with other titanosaurs have been relatively consistent in phylogenetic analyses. This led to its use in a re-definition of the group Colossosauria by Carballido and colleagues in 2022.

<i>Notocolossus</i> Extinct genus of dinosaurs

Notocolossus is a genus of titanosaurian sauropod dinosaur from late Cretaceous strata of Mendoza Province, Argentina.

<span class="mw-page-title-main">Rinconsauria</span> Extinct clade of dinosaurs

Rinconsauria is an extinct clade of giant titanosaurian sauropods known from the late Cretaceous period of Argentina.

The Lago Colhué Huapí Formation is a Late Cretaceous geologic formation of the Chubut Group in the Golfo San Jorge Basin in Patagonia, Argentina. The formation, named after Lake Colhué Huapí, is overlain by the Salamanca Formation of the Río Chico Group and in some areas by the Laguna Palacios Formation.

References

  1. 1 2 3 4 5 6 Calvo, Jorge O.; B.J.G. Riga (2003). "Rinconsaurus caudamirus gen. et sp nov., a new titanosaurid (Dinosauria, Sauropoda) from the Late Cretaceous of Patagonia, Argentina". Revista Geológica de Chile. 30 (2): 333–353. doi: 10.4067/s0716-02082003000200011 . ISSN   0716-0208.
  2. Pérez Moreno, Agustín; Otero, Alejandro; Carballido, José L.; Salgado, Leonardo; Calvo, Jorge O. (2022-10-07). "The appendicular skeleton of Rinconsaurus caudamirus (Sauropoda: Titanosauria) from the Upper Cretaceous of Patagonia, Argentina". Cretaceous Research. 142: 105389. Bibcode:2023CrRes.14205389P. doi:10.1016/j.cretres.2022.105389. ISSN   0195-6671. S2CID   252799392.
  3. Filippi, Leonardo S.; Juarez Valieri, Ruben D.; Barrios, Francisco (2021-05-31). "The prefrontal of Rinconsaurus caudamirus (Sauropoda, Titanosauria) as a crocodyliform ilium". Cretaceous Research. 125: 104852. Bibcode:2021CrRes.12504852F. doi:10.1016/j.cretres.2021.104852. ISSN   0195-6671.
  4. Pérez Moreno, Agustín; Carballido, José L.; Otero, Alejandro; Salgado, Leonardo; Calvo, Jorge O. (2021-09-22). "The axial skeleton of Rinconsaurus caudamirus (Sauropoda: Titanosauria) from the Late Cretaceous of Patagonia, Argentina". Ameghiniana. 59 (1): 1–46. doi: 10.5710/AMGH.13.09.2021.3427 . ISSN   0002-7014. S2CID   240592543.
  5. Casal, G.; Martínez, R.D.; Luna, M.; Sciutto, J.C; and Lamanna, M.C. (2007). "Aeolosaurus colhuehuapensis sp. nov. (Sauropoda, Titanosauria) de la Formación Bajo Barreal, Cretácico Superior de Argentina [Aeolosaurus colhuehuapensis sp. nov. (Sauropoda, Titanosauria) from the Bajo Barreal Formation, Upper Cretaceous of Argentina]". Revista Brasileira de Paleontologia 10(1):53-62
  6. Prasad, V.; Caroline A. E. Strömberg, Habib Alimohammadian, and Ashok Sahni, CA; Alimohammadian, H; Sahni, A (2005). "Dinosaur Coprolites and the Early Evolution of Grasses and Grazers". Science. 310 (5751): 1177–1180. Bibcode:2005Sci...310.1177P. doi:10.1126/science.1118806. PMID   16293759. S2CID   1816461.{{cite journal}}: CS1 maint: multiple names: authors list (link)