Pilmatueia Temporal range: Early Cretaceous, ![]() | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Superfamily: | † Diplodocoidea |
Family: | † Dicraeosauridae |
Genus: | † Pilmatueia Coria et al., 2019 |
Type species | |
†Pilmatueia faundezi Coria et al., 2019 |
Pilmatueia is a diplodocoid sauropod belonging to the family Dicraeosauridae that lived in Argentina during the Early Cretaceous. Its type and only species is Pilmatueia faundezi. Pilmatueia was probably closely related to other South American dicraeosaurids such as Amargasaurus . Pilmatueia had relatively pneumatic vertebrae compared to other dicraeosaurids, which were otherwise characterized by a reduction in pneumaticity relative to other sauropods. Pilmatueia dates to the Valanginian, an age of the Cretaceous period for which dinosaur faunas are poorly known.
Fossils of Pilmatueia faundezi were discovered in Neuquén Province, Argentina, at a site called Pilmatué. [1] Fossil excavations at Pilmatué began in 2009, [2] and the discovery of dicraeosaurid remains at Pilmatué was first announced in 2012, at a paleontology conference in Buenos Aires. [3] [1] In 2019, [lower-alpha 1] Rodolfo Coria and colleagues named the new genus and species Pilmatueia faundezi. The genus name refers to the Pilmaté locality, and the species epithet recognizes Ramón Faúndez, manager of the Museo Municipal de Las Lajas, who supported the excavation project. The discovery of Pilmatueia in the Valanginian, an age of the Early Cretaceous with poorly known dinosaur faunas, helped fill a gap in the dicraeosaurid fossil record between Jurassic dicraeosaurids and the later Amargasaurus. [1]
Pilmatueia faundezi fossils have been found in the Mulichinco Formation of Argentina. Known material includes the holotype, MLL Pv-005, a posterior dorsal vertebra; [1] the paratype MLL-Pv-002, a posterior cervical vertebra; MLL-Pv-010, a partial skeleton including cervical vertebrae, dorsal vertebrae, ribs, a caudal vertebra, and incomplete scapulae; [4] and other isolated vertebrae. [1] Other dicraeosaurid fossils have been found in the Mulichinco Formation, including a femur [1] and a natural cranial endocast, [5] and it is possible that they also belong to Pilmatueia faundezi. [1] [6]
Pilmatueia, like other dicraeosaurids, had tall, deeply forked neural spines on its cervical and dorsal vertebrae. The cervical neural spines were low compared to those of Amargasaurus and Bajadasaurus. [4] As in most dicraeosaurids, the neural spines were inclined forward, unlike the backswept neural spines seen in Amargasaurus. [4] Unlike most dicraeosaurids, which have reduced vertebral pneumaticity compared to other sauropods, the cervical vertebrae of Pilmatueia contain pneumatic chambers. However, as in other dicraeosaurids, the dorsal centra lack the pneumatic foramina that are characteristic of most sauropods. [7] [8] The cervical centra have ventral keels that are forked anteriorly and posteriorly, unlike the simple midline keels typical of other diplodocoids. [4] A foramen is present on the proximal end of the cervical ribs, which probably is a nutrient foramen and not a pneumatic structure. The anterior dorsal vertebrae show an autapomorphic ridge on the anterior centrodiapophyseal lamina. [1] Even the posterior dorsal vertebrae had forked neural spines, as in its close relative Amargasaurus, and unlike other dicraeosaurids in which forking of the neural spines only extended into the middle dorsal vertebrae. [4] The posterior dorsal vertebrae show an autapomorphic pair of deep fossae near the bases of the neural spines. [1] The scapula has a ridge on the medial surface near the acromion, which is not present in other dicraeosaurids. [4]
Phylogenetic position of Pilmatueia within Dicraeosauridae, following an analysis by Windholtz et al. that recovered their favored hypothesis of South American dicraeosaurid monophyly, [4] with South American dicraeosaurids highlighted in red. |
Pilmatueia is a dicraeosaurid sauropod. Its phylogenetic position within Dicraeosauridae is uncertain, with some analyses finding it to be closely related to Amargasaurus [1] [4] and other analyses finding it to be a more basal dicraeosaurid, outside the clade uniting Dicraeosaurus, Brachytrachelopan, and Amargasaurus. [9] [10] [4] Windholz et al. argued that, while phylogenetic analyses do not currently provide clear resolution for its affinities, on chronological and biogeographic grounds it is most likely to belong to a clade uniting the South American dicraeosaurids. [4]
Pilmatueia lived in what is now Argentina during the Valanginian age of the Cretaceous, and is a member of the Bajadan faunal assemblage. It coexisted with an indeterminate species of diplodocid [6] and the small, early carcharodontosaur Lajasvenator . [2] Podocarp trees were present in the ecosystem. [11]
Argentinosaurus is a genus of giant sauropod dinosaur that lived during the Late Cretaceous period in what is now Argentina. Although it is only known from fragmentary remains, Argentinosaurus is one of the largest known land animals of all time, perhaps the largest, measuring 30–35 metres (98–115 ft) long and weighing 65–80 tonnes. It was a member of Titanosauria, the dominant group of sauropods during the Cretaceous. It is widely regarded by many paleontologists as the biggest dinosaur ever, and perhaps lengthwise the longest animal ever, though both claims have no concrete evidence yet.
Diplodocids, or members of the family Diplodocidae, are a group of sauropod dinosaurs. The family includes some of the longest creatures ever to walk the Earth, including Diplodocus and Supersaurus, some of which may have reached lengths of up to 42 metres (138 ft).
Agustinia is a genus of sauropod dinosaur from the Early Cretaceous of South America. The genus contains a single species, Agustinia ligabuei, known from a single specimen that was recovered from the Lohan Cura Formation of Neuquén Province in Argentina. It lived about 116–108 million years ago, in the Aptian–Albian stages of the Early Cretaceous Period.
Amazonsaurus is a genus of diplodocoid sauropod dinosaur from the Early Cretaceous Period of what is now South America. It would have been a large-bodied quadrupedal herbivore with a long neck and whiplash tail. Although more derived diplodocoids were some of the longest animals ever to exist, Amazonsaurus was probably not more than 12 meters (40 ft) long. Gregory S. Paul estimated in 2010 its weight at 5000 kg.
Amargasaurus is a genus of sauropod dinosaur from the Early Cretaceous epoch of what is now Argentina. The only known skeleton was discovered in 1984 and is virtually complete, including a fragmentary skull, making Amargasaurus one of the best-known sauropods of its epoch. Amargasaurus was first described in 1991 and contains a single known species, Amargasaurus cazaui. It was a large animal, but small for a sauropod, reaching 9 to 13 meters in length. Most distinctively, it sported two parallel rows of tall spines down its neck and back, taller than in any other known sauropod. In life, these spines could have stuck out of the body as solitary structures that supported a keratinous sheath. An alternate hypothesis, now more favored, postulates that they could have formed a scaffold supporting a skin sail. They might have been used for display, combat, or defense.
Dicraeosauridae is a family of diplodocoid sauropods who are the sister group to Diplodocidae. Dicraeosaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as Amargasaurus, Suuwassea, Dicraeosaurus, and Brachytrachelopan. Specimens of this family have been found in North America, Asia, Africa, and South America. In 2023, a dicraeosaurid fossil was discovered in India for the first time. Their temporal range is from the Early or Middle Jurassic to the Early Cretaceous. Few dicraeosaurids survived into the Cretaceous, the youngest of which was Amargasaurus.
Brachytrachelopan is a short-necked sauropod dinosaur from the Late Jurassic of Argentina. The holotype and only known specimen was collected from an erosional exposure of fluvial sandstone within the Cañadón Calcáreo Formation on a hill approximately 25 kilometres (16 mi) north-northeast of Cerro Cóndor, Chubut Province, in west-central Argentina, South America. Though very incomplete, the skeletal elements recovered were found in articulation and include eight cervical, twelve dorsal, and three sacral vertebrae, as well as proximal portions of the posterior cervical ribs and all the dorsal ribs, the distal end of the left femur, the proximal end of the left tibia, and the right ilium. Much of the specimen was probably lost to erosion many years before its discovery. The type species is Brachytrachelopan mesai. The specific name honours Daniel Mesa, a local shepherd who discovered the specimen while searching for lost sheep. The genus name translates as "short-necked Pan", Pan being the god of the shepherds.
Neuquensaurus is a genus of saltasaurid sauropod dinosaur that lived in the Late Cretaceous, about 80 million years ago in Argentina in South America. Its fossils were recovered from outcrops of the Anacleto Formation around Cinco Saltos, near the Neuquén river from which its name is derived.
Puertasaurus is a genus of sauropod dinosaur that lived in South America during the Late Cretaceous Period. It is known from a single specimen recovered from sedimentary rocks of the Cerro Fortaleza Formation in southwestern Patagonia, Argentina, which probably is Campanian or Maastrichtian in age. The only species is Puertasaurus reuili. Described by the paleontologist Fernando Novas and colleagues in 2005, it was named in honor of Pablo Puerta and Santiago Reuil, who discovered and prepared the specimen. It consists of four well-preserved vertebrae, including one cervical, one dorsal, and two caudal vertebrae. Puertasaurus is a member of Titanosauria, the dominant group of sauropods during the Cretaceous.
Amargatitanis is a genus of dicraeosaurid sauropod dinosaur from the Barremian-age La Amarga Formation of Neuquén, Argentina. It is known from a single, incomplete postcranial skeleton consisting of a partial hindlimb, ischium, and two vertebrae. These remains were unearthed by Argentine paleontologist José Bonaparte in 1983 during an expedition by the Museo Argentino de Ciencias Naturales and later described as a new genus and species, Amargatitanis macni by Sebastián Apesteguía. The genus name comes from the words Amarga, where the fossils were collected, and titanis meaning "titan". Its species name is in reference to the MACN, where the remains are stored.
Overosaurus is an extinct genus of sauropod dinosaurs, containing only a single species, Overosaurus paradasorum. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia. Overosaurus paradasorum was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately 10 m (33 ft). It was a ground-dwelling herbivore.
Katepensaurus is an extinct genus of rebbachisaurid sauropod dinosaur known from the Late Cretaceous of south-central Chubut Province of central Patagonia, Argentina. It contains a single species, Katepensaurus goicoecheai.
Leinkupal is a genus of diplodocine sauropod known from the Early Cretaceous of the Bajada Colorada Formation, southeastern Neuquén Basin in the Neuquén Province of Argentina. It contains a single species, Leinkupal laticauda.
Bajadasaurus is a genus of sauropod dinosaur from the Early Cretaceous epoch of northern Patagonia, Argentina. It was first described in 2019 based on a single specimen found in 2010 that includes a largely complete skull and parts of the neck. The only species is Bajadasaurus pronuspinax. The genus is classified as a member of the Dicraeosauridae, a group of relatively small and short-necked sauropods.
The Bajada Colorada Formation is a geologic formation of the southern Neuquén Province in the Neuquén Basin of northern Patagonia, Argentina. The formation belongs to the Mendoza Group and is Late Berriasian to Early Valanginian in age. The formation is renowned for preserving fossil remains of Bajadasaurus pronuspinax, a genus of dicraeosaurid dinosaurs named after the formation.
The Ernesto Bachmann Paleontological Museum (MEB) in Villa El Chocón, Neuquén Province, Argentina, is a municipal museum dedicated to the paleontology, archaeology and history of Villa El Chocón and its surroundings.
Lajasvenator is a genus of carcharodontosaurid dinosaur from the Mulichinco Formation from Neuquén Province in Argentina. The type and only species is Lajasvenator ascheriae. It was probably one of the smallest known allosauroids, being approximately only half the length of Concavenator, about 2.9 m (9.5 ft).
The Mulichinco Formation is a geological formation in Argentina. It is Valanginian in age and is predominantly terrestrial, being deposited at a time of marine regression in the Neuquén Basin, and predominantly consists of siliciclastic rocks.
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