Smitanosaurus

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Smitanosaurus
Temporal range: Late Jurassic, 155.7–145.5  Ma
Smitanosaurus agilis skull and neck.jpg
Back of the skull and neck vertebrae
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Sauropoda
Clade: Eusauropoda
Clade: Neosauropoda
Family: Dicraeosauridae
Genus: Smitanosaurus
Whitlock & Wilson, 2020
Type species
Smitanosaurus agilis
(Marsh, 1889)
Synonyms
  • Camarasaurus agilis(Marsh, 1889)
  • Morosaurus agilisMarsh, 1889

Smitanosaurus (meaning "smith lizard") is a genus of dicraeosaurid sauropod dinosaur from the Late Jurassic of Colorado. The genus contains one species, S. agilis, originally assigned to the defunct genus Morosaurus . [1]

Contents

History

Skull shown from the back; left with proatlas (a) and atlas, and right in oblique view Smitanosaurus skull from the back.jpg
Skull shown from the back; left with proatlas (a) and atlas, and right in oblique view

The species Morosaurus agilis was originally named by Othniel Charles Marsh in 1889 for a partial skull, proatlases, and three cervical vertebrae found in 1883 in the Morrison Formation of Colorado. [2] However, the genus Morosaurus was in 1907 reinterpreted as a junior synonym of Camarasaurus , and most of the species assigned to the former genus were reassigned to the latter. [3] M. agilis, on the other hand, was left defunct without a proper generic assignment. [4] Over time comparisons have been drawn with Haplocanthosaurus , Diplodocus and brachiosaurids, but never with a phylogenetic analysis. Further preparation and analysis by Whitlock & Wilson, 2020, have since reinterpreted it as a dicraeosaurid, warranting the new genus name Smitanosaurus. The new generic name means "smith lizard", from the Old Saxon smitan, referring to J. August Smith, who excavated and sketched the holotype USNM 5384, as well as the Smithsonian Institution, where the remains are stored. [1]

Classification

Whitlock & Wilson's redescription marks the first time S. agilis has been added to a phylogenetic analysis. They recovered it as a member of Dicraeosauridae, making it one of the ever-growing number of North American dicraeosaurids. Their cladogram is shown below: [1]

Dicraeosauridae

Kaatedocus

Smitanosaurus

Bajadasaurus

Lingwulong

MOR 592

Amargatitanis

Suuwassea

Pilmatueia

Dicraeosaurus

Amargasaurus

Brachytrachelopan

Related Research Articles

<i>Apatosaurus</i> Sauropod dinosaur genus from Late Jurassic period

Apatosaurus is a genus of herbivorous sauropod dinosaur that lived in North America during the Late Jurassic period. Othniel Charles Marsh described and named the first-known species, A. ajax, in 1877, and a second species, A. louisae, was discovered and named by William H. Holland in 1916. Apatosaurus lived about 152 to 151 million years ago (mya), during the late Kimmeridgian to early Tithonian age, and are now known from fossils in the Morrison Formation of modern-day Colorado, Oklahoma, New Mexico, Wyoming, and Utah in the United States. Apatosaurus had an average length of 21–22.8 m (69–75 ft), and an average mass of 16.4–22.4 t. A few specimens indicate a maximum length of 11–30% greater than average and a mass of 32.7–72.6 t.

<i>Supersaurus</i> Extinct genus of dinosaurs

Supersaurus is a genus of diplodocid sauropod dinosaur that lived in North America during the Late Jurassic period. The type species, S. vivianae, was first discovered by Vivian Jones of Delta, Colorado, in the middle Morrison Formation of Colorado in 1972. The fossil remains came from the Brushy Basin Member of the formation, dating to about 153 million years ago. A potential second species, S. lourinhanensis, is known from Portugal and has been dated to a similar time.

<i>Camarasaurus</i> Camarasaurid sauropod dinosaur genus from Late Jurassic Period

Camarasaurus was a genus of quadrupedal, herbivorous dinosaurs. It was the most common of the giant sauropods to be found in North America. Its fossil remains have been found in the Morrison Formation of Colorado and Utah, dating to the Late Jurassic epoch, between 155 and 145 million years ago.

Coelurus is a genus of coelurosaurian dinosaur from the Late Jurassic period. The name means "hollow tail", referring to its hollow tail vertebrae. Although its name is linked to one of the main divisions of theropods (Coelurosauria), it has historically been poorly understood, and sometimes confused with its better-known contemporary Ornitholestes. Like many dinosaurs studied in the early years of paleontology, it has had a confusing taxonomic history, with several species being named and later transferred to other genera or abandoned. Only one species is currently recognized as valid: the type species, C. fragilis, described by Othniel Charles Marsh in 1879. It is known from one partial skeleton found in the Morrison Formation of Wyoming, United States. It was a small bipedal carnivore with elongate legs.

<i>Cetiosauriscus</i> Genus of reptiles (fossil)

Cetiosauriscus is a genus of sauropod dinosaur that lived between 166 and 164 million years ago during the Callovian in what is now England. A herbivore, Cetiosauriscus had — by sauropod standards — a moderately long tail, and longer forelimbs, making them as long as its hindlimbs. It has been estimated as about 15 m (49 ft) long and between 4 and 10 t in weight.

Haplocanthosaurus is a genus of intermediate sauropod dinosaur. Two species, H. delfsi and H. priscus, are known from incomplete fossil skeletons. It lived during the late Jurassic period, 155 to 152 million years ago. The type species is H. priscus, and the referred species H. delfsi was discovered by a young college student named Edwin Delfs in Colorado, United States. Haplocanthosaurus specimens have been found in the very lowest layer of the Morrison Formation, along with Hesperosaurus mjosi, Brontosaurus yahnahpin, and Allosaurus jimmadseni.

<i>Elaphrosaurus</i> Ceratosaurian theropod dinosaur genus from the Late Jurassic Period

Elaphrosaurus is a genus of ceratosaurian theropod dinosaur that lived approximately 154 to 150 million years ago during the Late Jurassic Period in what is now Tanzania in Africa. Elaphrosaurus was a medium-sized but lightly built member of the group that could grow up to 6.2 m (20 ft) long. Morphologically, this dinosaur is significant in two ways. Firstly, it has a relatively long body but is very shallow-chested for a theropod of its size. Secondly, it has very short hindlimbs in comparison with its body. Phylogenetic analyses indicate that this genus is likely a ceratosaur. Earlier suggestions that it is a late surviving coelophysoid have been examined but generally dismissed. Elaphrosaurus is currently believed to be a very close relative of Limusaurus, an unusual beaked ceratosaurian which may have been either herbivorous or omnivorous.

<i>Brontosaurus</i> Diplodocid sauropod dinosaur genus from Late Jurassic Period

Brontosaurus is a genus of gigantic quadruped sauropod dinosaurs. Although the type species, B. excelsus, had long been considered a species of the closely related Apatosaurus, researchers proposed in 2015 that Brontosaurus is a genus separate from Apatosaurus and that it contains three species: B. excelsus, B. yahnahpin, and B. parvus.

Suuwassea is a genus of dicraeosaurid sauropod dinosaur found in the Upper Jurassic strata of the Morrison Formation, located in southern Carbon County, Montana, United States. The fossil remains were recovered in a series of expeditions during a period spanning the years 1999 and 2000 and were described by J.D. Harris and Peter Dodson in 2004. They consist of a disarticulated but associated partial skeleton, including partial vertebral series and limb bones.

Diplodocoidea Extinct superfamily of reptiles

Diplodocoidea is a superfamily of sauropod dinosaurs, which included some of the longest animals of all time, including slender giants like Supersaurus, Diplodocus, Apatosaurus, and Amphicoelias. Most had very long necks and long, whip-like tails; however, one family are the only known sauropods to have re-evolved a short neck, presumably an adaptation for feeding low to the ground. This adaptation was taken to the extreme in the highly specialized sauropod Brachytrachelopan. A study of snout shape and dental microwear in diplodocoids showed that the square snouts, large proportion of pits, and fine subparallel scratches in Apatosaurus, Diplodocus, Nigersaurus, and Rebbachisaurus suggest ground-height nonselective browsing; the narrow snouts of Dicraeosaurus, Suuwassea, and Tornieria and the coarse scratches and gouges on the teeth of Dicraeosaurus suggest mid-height selective browsing in those taxa. This taxon is also noteworthy because diplodocoid sauropods had the highest tooth replacement rates of any vertebrates, as exemplified by Nigersaurus, which had new teeth erupting every 30 days.

Dicraeosauridae Extinct family of reptiles

Dicraeosauridae is a family of diplodocoid sauropods who are the sister group to Diplodocidae. Dicraesaurids are a part of the Flagellicaudata, along with Diplodocidae. Dicraeosauridae includes genera such as Amargasaurus, Suuwassea, Dicraeosaurus, and Brachytrachelopan. Specimens of this family have been found in North America, Asia, Africa, and South America. Their temporal range is from the Early or Middle Jurassic to the Early Cretaceous. Few dicraeosaurids survived into the Cretaceous, the youngest of which was Amargasaurus.

<i>Atlantosaurus</i> Extinct genus of dinosaurs

Atlantosaurus is a dubious genus of sauropod dinosaur. It contains a single species, Atlantosaurus montanus, from the upper Morrison Formation of Colorado, United States. Atlantosaurus was the first sauropod to be described during the infamous 19th century Bone Wars, during which scientific methodology suffered in favor of pursuit of academic acclaim.

<i>Tornieria</i> Diplodocid sauropod dinosaur genus from Late Jurassic period

Tornieria is a genus of diplodocid sauropod dinosaur from Late Jurassic of Tanzania. It has a convoluted taxonomic history.

<i>Brachytrachelopan</i> Extinct genus of dinosaurs

Brachytrachelopan is a short-necked sauropod dinosaur from the Late Jurassic of Argentina. The holotype and only known specimen was collected from an erosional exposure of fluvial sandstone within the Cañadón Calcáreo Formation on a hill approximately 25 kilometres (16 mi) north-northeast of Cerro Cóndor, Chubut Province, in west-central Argentina, South America. Though very incomplete, the skeletal elements recovered were found in articulation and include eight cervical, twelve dorsal, and three sacral vertebrae, as well as proximal portions of the posterior cervical ribs and all the dorsal ribs, the distal end of the left femur, the proximal end of the left tibia, and the right ilium. Much of the specimen was probably lost to erosion many years before its discovery. The type species is Brachytrachelopan mesai. The specific name honours Daniel Mesa, a local shepherd who discovered the specimen while searching for lost sheep. The genus name translates as "short-necked Pan", Pan being the god of the shepherds.

<i>Nanosaurus</i> Extinct genus of dinosaurs

Nanosaurus is the name given to a genus of neornithischian dinosaur that lived about 155 to 148 million years ago, during the Late Jurassic-age. Its fossils are known from the Morrison Formation of the south-western United States. The type and only species, Nanosaurus agilis, was described and named by Othniel Charles Marsh in 1877. The taxon has a complicated taxonomic history, largely the work of Marsh and Peter M. Galton, involving the genera Laosaurus, Hallopus, Drinker, Othnielia, and Othnielosaurus, the latter three now being considered to be synonyms of Nanosaurus. It had historically been classified as a hypsilophodont or fabrosaur, types of generalized small bipedal herbivore, but more recent research has abandoned these groupings as paraphyletic and Nanosaurus is today considered a basal member of Neornithischia.

<i>Diplodocus</i> Genus of diplodocid sauropod dinosaurs (fossil)

Diplodocus is a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek διπλός (diplos) "double" and δοκός (dokos) "beam", in reference to the double-beamed chevron bones located in the underside of the tail, which were then considered unique.

<i>Ruyangosaurus</i> Extinct genus of dinosaurs

Ruyangosaurus is a genus of titanosauriform sauropod dinosaur recovered from the Early Cretaceous Haoling Formation of China. The type species is R. giganteus, described in 2009 by Lü Junchang et al.

<i>Leinkupal</i> Extinct genus of reptiles

Leinkupal is a genus of diplodocine sauropod known from the Early Cretaceous of the Bajada Colorada Formation, southeastern Neuquén Basin in the Neuquén Province of Argentina. It contains a single species, Leinkupal laticauda.

<i>Bajadasaurus</i> Genus of dicraeosaurid sauropod from the Early Cretaceous period

Bajadasaurus is a genus of sauropod dinosaur from the Early Cretaceous epoch of northern Patagonia, Argentina. It was first described in 2019 based on a single specimen found in 2010 that includes a largely complete skull and parts of the neck. The only species is Bajadasaurus pronuspinax. The genus is classified as a member of the Dicraeosauridae, a group of relatively small and short-necked sauropods.

References

  1. 1 2 3 John A. Whitlock; Jeffrey A. Wilson Mantilla (2020). "The Late Jurassic sauropod dinosaur 'Morosaurus' agilis Marsh, 1889 reexamined and reinterpreted as a dicraeosaurid". Journal of Vertebrate Paleontology. Online edition: e1780600. doi: 10.1080/02724634.2020.1780600 .
  2. O. C. Marsh. 1889. Notice of new American Dinosauria. The American Journal of Science and Arts, series 3 38:331-336
  3. Gilmore, C.W. 1907. "The type of the Jurassic reptile Morosaurus agilis redescribed, with a note on Camptosaurus ". Proceedings of the United States National Museum32: 151–165
  4. O. P. Hay. 1930. Second Bibliography and Catalogue of the Fossil Vertebrata of North America 390(II):1-1074