Gongxianosaurus Temporal range: Early Jurassic | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Genus: | † Gongxianosaurus |
Species: | †G. shibeiensis |
Binomial name | |
†Gongxianosaurus shibeiensis He et al., 1998 | |
Synonyms | |
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Gongxianosaurus is a genus of basal sauropod dinosaur from the early Jurassic Period (Toarcian stage [1] ). The only species is Gongxianosaurus shibeiensis. Based on four fragmentary to complete specimens found in China (Sichuan Province), it is one of the most completely known early sauropods. The skeleton is known in large part, missing both the hand and the majority of the skull. [2] Gongxianosaurus was firstly named and described in a short note published in 1998; [3] however, a comprehensive description has yet to be published. [4] Gongxianosaurus shibeiensis was named for the place it was found, near the village Shibei in Gong County (珙县; Pinyin: Gǒng Xiàn). [2]
Gongxianosaurus may have reached 14 metres (46 ft) in length. [2] Like other sauropods, it moved quadrupedally (on four legs), as indicated by the elongated fore limbs that reached 70 to 75% of hind limb length. [2] The pedal phalanges were short and robust, as typical for sauropods. The pedal phalangeal formula, counting the number of phalanges for each digit starting from the innermost, was 2-3-4-5-1. All but the outermost digit ended in claws. [2] Unlike in other sauropods, pleurocoels (deep lateral excavations of the vertebrae) were absent; [2] thus, the vertebrae would have been quite massive. The sacrum was made of three fused sacral vertebrae, fewer than in later sauropods. [2] [5] The chevrons were unbifurcated. [2]
An important characteristic of sauropod limbs is their reduced ossification – the tendency to replace bone by cartilage. Gongxianosaurus is the only known sauropod with ossified distal tarsals. Thus, either Gongxianosaurus was one of the basalmost sauropods, or ossified distal tarsals were present in other early sauropods but are simply not preserved due to the fragmentation of the specimens. [5]
Basal Sauropod phylogeny simplified after Apaldetti et al. (2011). [6] |
Because the fossils are not fully described yet, available character information that can be used in cladistic analyses is limited. Thus, only few cladistical analyses have incorporated Gongxianosaurus. A recent analysis by Apaldetti et al. (2011) suggests that Gongxianosaurus was more basal than Vulcanodon , Tazoudasaurus and Isanosaurus , but more derived than the early sauropods Antetonitrus , Lessemsaurus , Blikanasaurus , Camelotia and Melanorosaurus . [6]
Gongxianosaurus fossils were found near the village of Shibei (Sichuan province) in purple mudstones pertaining to the Ziliujing Formation (Dongyueshan Member). [3] These rocks are considered to be Toarcian in age (182.7 to 174.1 mya). [1] Thus, Gongxianosaurus is geologically younger than the "prosauropod" Lufengosaurus but older than the basal sauropod Shunosaurus . [3]
The fossils were found in May 1997 during a geological exploration. Excavation started in the same month and led to the recovery of a wealth of fossils in an area of approximately 200 square meters. While most fossils pertain to a new sauropod genus, remains of theropods have also been found. [3] The sauropod material includes four fragmentary to complete individuals similar in size, together encompassing most of the skeleton, though hand and skull bones were not found except a premaxilla and teeth. In 1998, the sauropod material was briefly described as a new genus and species, Gongxianosaurus shibeiensis, in a preliminary note by palaeontologists led by He Xinlu. [3] A more detailed description was announced, noting that excavation was still in progress while the paper was published. [3] In 2000, a second short description was published by Luo Yaonan and Wang Changsheng, also presenting Gongxianosaurus as a new sauropod and without mentioning the first description that was published two years before. [2] [7] Also, much of the information published by Luo and Wang was already published by He and colleagues. [7]
Luo and Wang suggest that several bones may not pertain to the type species Gongxianosaurus shibeiensis but to a second species of Gongxianosaurus. [2] Those remains were informally placed within the separate genus and species "Yibinosaurus zhoui" in 2001 and 2003 respectively by Ouyang Hui. [8] [9]
The holotype of Gongxianosaurus was kept in situ and a protective exhibition hall was built over it. The exhibition hall subsequently collapsed, and the specimen was most likely destroyed. [10]
Mamenchisaurus is a genus of sauropod dinosaur known for their remarkably long necks which made up nearly half the total body length. Numerous species have been assigned to the genus; however, the validity of these assignments has been questioned. Fossils have been found in the Sichuan Basin and Yunnan Province in China. Several species from the Upper Shaximiao Formation, whose geologic age is uncertain, have been described. However, evidence suggests this formation to be no earlier than the Oxfordian stage of the Late Jurassic. M. sinocanadorum dates to the Oxfordian stage, and M. anyuensis to the Aptian stage of the Early Cretaceous. Most species were medium-large to large sauropods, measuring roughly 15 to 26 meters in length—possibly up to 35 meters (115 ft), based on two undescribed vertebrae.
Vulcanodon is an extinct genus of sauropod dinosaur from the Early Jurassic Uppermost Forest Sandstone of southern Africa. The only known species is V. karibaensis. Discovered in 1969 in Zimbabwe, it was regarded as the earliest-known sauropod for decades, and is still one of the most primitive sauropods that has been discovered. As a quadrupedal, ground-dwelling herbivore, Vulcanodon already showed the typical sauropod body plan with column-like legs and a long neck and tail. It was smaller than most other sauropods, measuring approximately eleven metres (36 ft) in length. Vulcanodon is known from a fragmentary skeleton including much of the pelvic girdle, hindlimbs, forearms, and tail, but lacking the trunk and neck vertebrae as well as the skull.
Cetiosaurus meaning 'whale lizard', from the Greek keteios/κήτειος meaning 'sea monster' and sauros/σαυρος meaning 'lizard', is a genus of herbivorous sauropod dinosaur from the Middle Jurassic Period, living about 168 million years ago in what is now Britain.
Tazoudasaurus is a genus of gravisaurian, probably a vulcanodontid sauropod dinosaurs hailing from the late Early Jurassic (Toarcian), that was recovered in the "Toundoute Continental Series" located in the High Atlas Mountains of Morocco in North Africa. Along with Patagosaurus, Volkheimeria, Bagualia and Perijasaurus represents one of the few sauropods named from this stage on Gondwana, as well the only one from Africa.
Yunnanosaurus is an extinct genus of sauropodomorph dinosaur that lived approximately 199 to 183 million years ago in what is now the Yunnan Province, in China, for which it was named. Yunnanosaurus was a large sized, moderately-built, ground-dwelling, quadrupedal herbivore, that could also walk bipedally, and ranged in size from 7 meters (23 feet) long and 2 m (6.5 ft) high to 4 m (13 ft) high in the largest species.
Antetonitrus is a genus of sauropodiform dinosaur found in the Early Jurassic Elliot Formation of South Africa. The only species is Antetonitrus ingenipes. Sometimes considered a basal sauropod, it is crucial for the understanding of the origin and early evolution of this group. It was a quadrupedal herbivore, like its later relatives, but shows primitive adaptations to use the forelimbs for grasping, instead of purely for weight support.
Omeisaurus is a genus of sauropod dinosaur from the Middle Jurassic Period of what is now China. Its name comes from Mount Emei, where it was discovered in the lower Shaximiao Formation of Sichuan Province.
Volkheimeria is an extinct genus of sauropod dinosaurs that lived in what is now Argentina during the Early Jurassic, 178–179 million years ago. Its type and only species is Volkheimeria chubutensis.
Chinshakiangosaurus is a genus of dinosaur and probably one of the most basal sauropods known. The only species, Chinshakiangosaurus chunghoensis, is known from a fragmentary skeleton found in Lower Jurassic rocks in China. Chinshakiangosaurus is one of the few basal sauropods with preserved skull bones and therefore important for the understanding of the early evolution of this group. It shows that early sauropods may have possessed fleshy cheeks.
Dracopelta is a monospecific genus of ankylosaur dinosaur from Portugal that lived during the Late Jurassic in what is now the Lourinhã Formation. The type and only species is Dracopelta zbyszewskii, which is represented by a partial skeleton including unpublished material.
Emausaurus is a genus of thyreophoran or armored dinosaur from the Early Jurassic. Its fossils have been found in Mecklenburg-Vorpommern, northern Germany. Emausaurus is the only known Toarcian thyreophoran, as well as the only dinosaur from the zone of the same age with a formal name.
Lapparentosaurus is a genus of sauropod dinosaur from the Middle Jurassic. Its fossils were found in Madagascar. The type species is L. madagascariensis.
Gobititan is a genus of herbivorous sauropod dinosaur from the Aptian faunal stage of the Early Cretaceous. The name of this genus is derived from the Gobi desert region and the Titans of Greek mythology, which is a reference to its large body size. The specific name shenzhouensis, is derived from "Shenzhou", an ancient name for China.
Yuanmousaurus was a sauropod dinosaur from the Middle Jurassic period of China. It is known from incomplete remains, recovered in 2000 from the Zhanghe Formation in Yuanmou County in Yunnan Province. Yuanmousaurus was a relatively large sauropod and may have reached about 17 meters (56 ft) in length. It was a basal member of the Sauropoda, but its exact systematic position is unclear. A recent study placed Yuanmousaurus within the family Mamenchisauridae. It may be a dubious genus. The only and type species was Yuanmousaurus jiangyiensis.
The Ziliujing Formation is a geological formation in China, It is Early Jurassic in age. It is part of the stratigraphy of the Sichuan Basin. The dinosaur Gongxianosaurus and indeterminate theropod material are known from the Dongyuemiao Member of the formation, as well as dinosaur footprints, Zizhongosaurus and indeterminate prosauropods from the Da'anzhai Member. The basal sauropod Sanpasaurus is known from the Maanshan Member. An unnamed stegosaur and the pliosauroid plesiosaur Sinopliosaurus are also known from this formation but they were found an indeterminate member. An unnamed teleosaurid known from a complete skull has also been found in the formation, pending a formal description. The deposition environment during the Da'anzhai Member in the lower Toarcian is thought to have been that of a giant freshwater lake encompassing the whole of the Sichuan basin, around 3 times larger than Lake Superior, coeval with the Toarcian Oceanic Anoxic Event around 183 Ma.
Dongbeititan is a genus of sauropod dinosaur from the Early Cretaceous-age Yixian Formation of Beipiao, Liaoning, China. It is based on holotype DNHM D2867, a partial postcranial skeleton including bones from the limbs, shoulder and pelvic girdles, and vertebrae, which was described in 2007. Its describers suggested it was as a basal titanosauriform, not as derived as Gobititan or Jiutaisaurus, but more derived than Euhelopus, Fusuisaurus, and Huanghetitan. The type species is D. dongi, and it is the first named sauropod from the Yixian Formation, which is part of the well-known Jehol Group. The genus name refers to the region Dongbei and to Greek titan, "giant". The specific name honours the Chinese paleontologist Dong Zhiming. Like other sauropods, Dongbeititan would have been a large quadrupedal herbivore.
Shidaisaurus is a genus of metriacanthosaurid dinosaur. Its fossil was found in early Middle Jurassic-age rocks of the Chuanjie Formation in Yunnan, China. It is known from a partial skeleton, holotype DML-LCA 9701-IV, found at the bottom of an assemblage of nine dinosaur individuals, lacking most of the tail vertebrae, ribs, pectoral girdle, and limb bones. Shidaisaurus was described in 2009 by Wu and colleagues. The type species is Shidaisaurus jinae. Generic name and specific name in combination refer to the Jin-Shidai Company that oversaw excavation and inspection of the Jurassic World Park near the site.
Tonganosaurus is a genus of mamenchisaurid sauropod dinosaur, similar to Omeisaurus. It is known from one specimen consisting of twenty vertebrae, a front limb and pectoral girdle, and a complete hind limb with partial hip. It was discovered in the Yimen Formation, China. The horizon of the specimen and the age of the Yimen Formation is controversial. The formation has been divided into three levels, and Tonganosaurus appears to be of late Early Jurassic (Pliensbachian) age. Tonganosaurus is the oldest known member of the mamenchisaurids, being almost 15 million years older than the next-oldest members of the group. It was first named by Li Kui, Yang Chun-Yan, Liu Jian and Wang Zheng-Xin in 2010 and the type species is Tonganosaurus hei.
Qijianglong is a genus of herbivorous mamenchisaurid sauropod dinosaur from the Early Cretaceous of China.