Turiasaurs | |
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Amanzia skeletal reconstruction, showing known material in blue | |
Moabosaurus, with an incorrect Camarasaurus like head | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Eusauropoda |
Clade: | † Turiasauria Royo-Torres et al., 2006 |
Genera [2] | |
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Synonyms | |
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Turiasauria is an unranked clade of basal sauropod dinosaurs known from Middle Jurassic to Early Cretaceous deposits in Europe, North America, and Africa.
Turiasauria was originally erected by Royo-Torres et al. (2006) to include Turiasaurus , Galvesaurus and Losillasaurus , all of which hail from the Villar del Arzobispo Formation (Tithonian-Berriasian) of Spain. Turiasuria was defined by the authors as "all Eusauropoda closer to Turiasaurus riodevensis than to Saltasaurus loricatus". [7] Cladistic analysis (Royo-Torres et al., 2006; 1927) of 309 characters and 33 taxa suggests that the turiasaurians lie outside the Neosauropoda and form a monophyletic group. The clade is diagnosed by the presence of vertical neural spines, posterior centroparapohyseal laminae on the dorsal vertebrae, the absence of pre- and postspinal laminae on the dorsal vertebrae, the absence of a scapular acromial crest, the presence of a prominent humeral deltopectoral crest, medial deflection of the proximal end of the humerus, and a distinct vertical ridge on the caudal side of the distal half of the ulna.
Turiasaurs were initially considered confined to Europe, with Turiasaurus from Spain and Zby from Portugal, [8] and the tooth taxa Cardiodon, Neosodon, and Oplosaurus were referred to the clade, but additional members were found in North America and Africa. Heart-shaped teeth are considered a synapomorphy of the turiasauria. Recently a heart shaped tooth found from the Jaisalmer Formation confirmed the presence of this clade in India during the Middle Jurassic (Bathonian). [9] Together with the Narindasaurus from the Bathonian of Madagascar, these are the oldest records of the group. It is therefore suggested that Turiasauria might have originated in Gondwana during the Middle Jurassic. [9] A tooth discovered in the Lower Pliensbachian (Lower Jurassic) Hasle Formation of Bornholm, NHMD 1185136, was also referred to the family, being, if truly a member, 17 My older than any previously known turiasaur. [1]
North American Mierasaurus and Moabosaurus from the Early Cretaceous are also considered to be turiasaurs. [4]
Remains of a very large species of turiasaur, not yet formally identified, have recently been unearthed from the earliest Cretaceous (Berriasian) aged Angeac-Charente bonebed in western France. [10] [11]
Indeterminate turiasaur material, consisting of a single vertebra, has been described from an unknown locality in the Early Cretaceous Wealden Group of England. [12]
Turiasaurus demonstrates that the evolution of enormous body size was not restricted to neosauropod clades such as the Diplodocidae and Titanosauria, but developed independently at least once in a lineage of more basal sauropods, the turiasaurians.
A 2009 thesis published by José Barco proposed that neither Galvesaurus nor Losillasaurus were turiasaurians. [13] Later, a master thesis by Francisco Gascó (2009) and Royo-Torres et al. (2009) reaffirmed the validity of Turiasauria. [14] [15]
Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located. Fossil evidence suggests that macronarian dinosaurs lived from the Middle Jurassic (Bathonian) through the Late Cretaceous (Maastrichtian). Macronarians have been found globally, including discoveries in Argentina, the United States, Portugal, China, and Tanzania. Like other sauropods, they are known to have inhabited primarily terrestrial areas, and little evidence exists to suggest that they spent much time in coastal environments. Macronarians are diagnosed through their distinct characters on their skulls, as well as appendicular and vertebral characters. Macronaria is composed of several subclades and families notably including Camarasauridae and Titanosauriformes, among several others. Titanosauriforms are particularly well known for being some of the largest terrestrial animals to ever exist.
Cardiodon was a herbivorous genus of sauropod dinosaur, based on a tooth from the late Bathonian-age Middle Jurassic Forest Marble Formation of Wiltshire, England. Historically, it is very obscure and usually referred to Cetiosaurus, but recent analyses suggest that it is a distinct genus, and possibly related to Turiasaurus. Cardiodon was the first sauropod genus named.
Atlasaurus is a genus of sauropod dinosaurs from Middle Jurassic beds in North Africa.
Neosodon was a genus of sauropod dinosaur from the Late Tithonian-age Upper Jurassic Sables et Gres a Trigonia gibbosa of Pas-de-Calais department, France. It has never been formally given a species name, but is often seen as N. praecursor, which actually comes from a different animal. Often in the past, it had been assigned to the wastebasket taxon Pelorosaurus, but restudy has suggested that it could be related to Turiasaurus, a roughly contemporaneous giant Spanish sauropod. It is only known from six teeth.
Lapparentosaurus is a genus of sauropod dinosaur from the Middle Jurassic. Its fossils were found in Madagascar. The type species is L. madagascariensis.
Losillasaurus is a genus of sauropod dinosaur from the Late Jurassic and possibly Early Cretaceous (Kimmeridgian-?Berriasian) in the southeast of Spain. The type species of the turiasaurian Losillasaurus giganteus was discovered in the Villar del Arzobispo Formation in Valencia and formally described by Casanovas, Santafé and Sanz in 2001. The holotype material is from a subadult and includes part of a skull; complete cervical, dorsal, sacral, and caudal vertebrae as well as several fragments; skeletal elements from the limbs including a humerus, ulna, radius, and metacarpal; sternal plates; and from the pelvis: the ilium, ischium, and pubis. The genus is characterized by the dimension and shape of the neural spine of the proximal caudal vertebrae. The humerus is 143 centimetres (56 in) long, which despite being from a subadult specimen is within 20% of the size of Paralititan. The size estimation proposed by Francisco Gascó in his master thesis is 15–18 m (49–59 ft) and 12-15 tons.
Galvesaurus, or Galveosaurus, is a genus of brachiosaurid sauropod dinosaur from the Late Jurassic period. Fossils of the only known species, G. herreroi, were found in Galve, Spain, hence its generic name, "Galve lizard". The specific name G. herreroi honours the discoverer, José María Herrero. Some researchers suggest that the taxon might represent a junior synonym of Lusotitan.
Moabosaurus is a genus of turiasaurian sauropod dinosaur from the Early Cretaceous Cedar Mountain Formation of Utah, United States.
Turiasaurus is a genus of sauropod dinosaurs. It is known from a single fossil specimen representing the species Turiasaurus riodevensis, found in the Kimmeridgian Villar del Arzobispo Formation of Teruel, Spain.
Camarasauridae is a family of sauropod dinosaurs. Among sauropods, camarasaurids are small to medium-sized, with relatively short necks. They are visually identifiable by a short skull with large nares, and broad, spatulate teeth filling a thick jaw. Based on cervical vertebrae and cervical rib biomechanics, camarasaurids most likely moved their necks in a vertical, rather than horizontal, sweeping motion, in contrast to most diplodocids.
Galve is a municipality located in the province of Teruel, Aragon, Spain. According to the 2006 census (INE), the municipality has a population of 145 inhabitants. There is an important paleontological site.
The Isalo III Formation is a geological formation in Madagascar, off the eastern coast of Africa. It dates back to the Middle Jurassic. The use of the term "Isalo III" is somewhat controversial as the two prior units Isalo I and II are Triassic cross-bedded sandstone units that form a continuous depositional sequence, while the "Isalo III" sandstones are not part of the same depositional sequence, and were deposited considerably later. and are perhaps better treated as part of several separate formations. It is traditionally divided into two subunits the lower, Bajocian aged Isalo IIIa unit also known as the Beronono Formation and the upper, Bathonian aged Isalo IIIb unit also known as the Sakaraha Formation or Sakahara Formation. The Sakaraha Formation consists of sandstones, marls and carbonates and represents a coastal plain environment, and is laterally equivalent to the predominantly carbonate Bemaraha Formation, which represents a coastal barrier lagoon complex. The formation is found in the northwest and in the southeast of the country and has provided a variety of fossils.
Ruyangosaurus is a genus of titanosauriform sauropod dinosaur recovered from the Early Cretaceous Haoling Formation of China. The type species is R. giganteus, described in 2009 by Lü Junchang et al.
Eusauropoda is a derived clade of sauropod dinosaurs. Eusauropods represent the node-based group that includes all descendant sauropods starting with the basal eusauropods of Shunosaurus, and possibly Barapasaurus, and Amygdalodon, but excluding Vulcanodon and Rhoetosaurus. The Eusauropoda was coined in 1995 by Paul Upchurch to create a monophyletic new taxonomic group that would include all sauropods, except for the vulcanodontids.
The Villar del Arzobispo Formation is a Late Jurassic to possibly Early Cretaceous geologic formation in eastern Spain. It is equivalent in age to the Lourinhã Formation of Portugal. It was originally thought to date from the Late Tithonian-Middle Berriasian, but more recent work suggests a Kimmeridigan-Late Tithonian, possibly dating to the Early Berriasian in some areas. The Villar del Arzobispo Formation's age in the area of Riodeva in Spain has been dated based on stratigraphic correlations as middle-upper Tithonian, approximately 145-141 million years old. In the area of Galve, the formation potentially dates into the earliest Cretaceous.
Laurasiformes is an extinct clade of sauropod dinosaurs from the late Early Cretaceous of Europe, North and South America. It was defined in 2009 by the Spanish paleontologist Rafael Royo-Torres as a clade containing sauropods more closely related to Tastavinsaurus than to Saltasaurus. Genera purported to form part of this clade include Aragosaurus, Galvesaurus, Phuwiangosaurus, Venenosaurus, Cedarosaurus, Tehuelchesaurus, Sonorasaurus and Tastavinsaurus.
Zby is an extinct genus of turiasaurian sauropod dinosaur known from the Late Jurassic of the Lourinhã Formation, central west Portugal. It contains a single species, Zby atlanticus. It is named after Georges Zbyszewski, who studied the geology and paleontology of Portugal.
Mierasaurus is an extinct genus of sauropod dinosaur from the Early Cretaceous of Utah, United States. The taxon was first described and named in 2017 by Rafael Royo-Torres and colleagues, from a mostly complete skeleton including a disarticulated partial skull and mandible, teeth, multiple vertebrae from along the length of the body, both scapulae, radius and ulna bones, a left manus, a complete pelvis, both femora and the entire left hindlimb. Additionally, they referred a lower jaw and femur from juvenile individuals, which were found nearby, to the genus. Collectively, Mierasaurus is among the most completely known North American sauropods. The genus name honours Bernardo de Miera y Pacheco, the first European scientist to enter what is now Utah. The type species for Mierasaurus is Mierasaurus bobyoungi, named after Robert Glen Young, a paleontologist who researched the Early Cretaceous of Utah.
Narindasaurus is a genus of turiasaurian sauropod dinosaur from the Middle Jurassic Isalo III Formation of Madagascar. The type species, N. thevenini was formally described by Royo-Torres et al. in 2020. The holotype, which consists of one specimen, is currently stored at the Muséum national d’Histoire naturelle and has been since 1906 or 1907.
Oblitosaurus is a genus of ankylopollexian ornithopod dinosaur from the Late Jurassic Villar del Arzobispo Formation of Spain. The type species is Oblitosaurus bunnueli.
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