Kaijutitan | |
---|---|
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Macronaria |
Clade: | † Titanosauria |
Genus: | † Kaijutitan Filippi, Salgado & Garrido, 2019 |
Type species | |
†Kaijutitan maui Filippi, Salgado & Garrido, 2019 |
Kaijutitan (meaning "Kaiju titan" after the type of Japanese movie monsters) is a genus of basal titanosaur dinosaur from the Sierra Barrosa Formation from Neuquén Province in Argentina. The type and only species is Kaijutitan maui. [1]
Kaijutitan was discovered by a team of researchers from the Museo Municipal Argentino Urquiza and the Museo Provincial de Ciencias Naturales “Prof. Dr. Juan Olsacher" from a layer from the Sierra Barrosa Formation, in Cañadón Mistringa, about 9 km southwest of the city of Rincón de los Sauces, in Neuquen, Argentina.
Kaijutitan is known from the holotype MAU-Pv-CM-522, stored at the Museo Municipal Argentino Urquiza, Paleontología de Vertebrados, Cañadón Mistringa, which is a partial skeleton preserving elements from several parts of the skeleton.
It can be distinguished from other titanosaurians through the possession of unique traits: the width between the basal tuberosities is almost four times the width of the foramen magnum; the foramen for the internal carotid artery is located on the back of the basipterygoid , almost at the middle of the distance between the basipterygoid and the basal tuberosities; the anterior cervical vertebrae have bifid neural spines; a medial tuber located behind both metapophyses on both cervical vertebrae; a spinopostzygapophyseal lamina bifurcated in the spinal sector, generating elongated deep pneumatic cavities on anterior cervical vertebrae; a posteroventral keel on the anterior cervical vertebrae, generated from the convergence of two ridges that originate from the bottom edge of the parapophysis; an accessory lamina that runs from postzygadiapophyseal lamina up to the spinoprezygapophyseal lamina in cervical vertebra; absence of proximal pneumatopores in dorsal ribs ; the prespinal lamina is triangular, which is a product of a dorsal expansion in anterior caudal vertebra; the absence of a ventromedial process in the bottom of the scapula ; the tibial proximal condyle is narrow, with its long axis anteroposteriorly oriented; the tibial cnemial crest projects forward; and foramina at the base of the ascending process of the astragalus . [1]
To test the systematic position of Kaijutitan, a phylogenetic analysis was conducted to resolve its affinities. A simplified cladogram, showing the results of the analysis, is presented below. [1]
Argentinosaurus is a genus of giant sauropod dinosaur that lived during the Late Cretaceous period in what is now Argentina. Although it is only known from fragmentary remains, Argentinosaurus is one of the largest known land animals of all time, perhaps the largest, measuring 30–35 metres (98–115 ft) long and weighing 65–80 tonnes. It was a member of Titanosauria, the dominant group of sauropods during the Cretaceous. It is widely regarded by many paleontologists as the biggest dinosaur ever, and perhaps lengthwise the longest animal ever, though both claims have no concrete evidence yet.
Aeolosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. Aeolosaurus is well known for a titanosaur, as it is represented by the remains of several individuals belonging to at least two species. However, like most titanosaurs, no remains of the skull are known. The holotype of Aeolosaurus rionegrinus consists of a series of seven tail vertebrae, as well as parts of both forelimbs and the right hindlimb. It was discovered in the Angostura Colorada Formation in Argentina, which dates from the Campanian stage of the Late Cretaceous, about 83 to 74 million years ago. The species A. maximus was transferred over to the new genus Arrudatitan in 2021.
Andesaurus is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Earth, it would have had a small head on the end of a long neck and an equally long tail.
Brachytrachelopan is a short-necked sauropod dinosaur from the Late Jurassic of Argentina. The holotype and only known specimen was collected from an erosional exposure of fluvial sandstone within the Cañadón Calcáreo Formation on a hill approximately 25 kilometres (16 mi) north-northeast of Cerro Cóndor, Chubut Province, in west-central Argentina, South America. Though very incomplete, the skeletal elements recovered were found in articulation and include eight cervical, twelve dorsal, and three sacral vertebrae, as well as proximal portions of the posterior cervical ribs and all the dorsal ribs, the distal end of the left femur, the proximal end of the left tibia, and the right ilium. Much of the specimen was probably lost to erosion many years before its discovery. The type species is Brachytrachelopan mesai. The specific name honours Daniel Mesa, a local shepherd who discovered the specimen while searching for lost sheep. The genus name translates as "short-necked Pan", Pan being the god of the shepherds.
Huabeisaurus was a genus of dinosaur from the Late Cretaceous. It was a sauropod which lived in what is present-day northern China. The type species, Huabeisaurus allocotus, was first described by Pang Qiqing and Cheng Zhengwu in 2000. Huabeisaurus is known from numerous remains found in the 1990s, which include teeth, partial limbs and vertebrae. Due to its relative completeness, Huabeisaurus represents a significant taxon for understanding sauropod evolution in Asia. Huabeisaurus comes from Kangdailiang and Houyu, Zhaojiagou Town, Tianzhen County, Shanxi province, China. The holotype was found in the unnamed upper member of the Huiquanpu Formation, which is Late Cretaceous (?Cenomanian–?Campanian) in age based on ostracods, charophytes, and fission-track dating.
Epachthosaurus was a genus of titanosaurian sauropod dinosaur from the Late Cretaceous. It was a basal lithostrotian titanosaur. Its fossils have been found in Central and Northern Patagonia in South America.
Neuquensaurus is a genus of saltasaurid sauropod dinosaur that lived in the Late Cretaceous, about 80 million years ago in Argentina in South America. Its fossils were recovered from outcrops of the Anacleto Formation around Cinco Saltos, near the Neuquén river from which its name is derived.
Lirainosaurus is a genus of titanosaur sauropod which lived in what is now Spain. The type species, Lirainosaurus astibiae, was described by Sanz, Powell, Le Loeuff, Martinez, and Pereda-Suberbiola in 1999. It was a relatively small sauropod, measuring 4 metres (13 ft) long, possibly up to 6 metres (20 ft) long for the largest individuals, and weighed about 2–4 metric tons.
Puertasaurus is a genus of sauropod dinosaur that lived in South America during the Late Cretaceous Period. It is known from a single specimen recovered from sedimentary rocks of the Cerro Fortaleza Formation in southwestern Patagonia, Argentina, which probably is Campanian or Maastrichtian in age. The only species is Puertasaurus reuili. Described by the paleontologist Fernando Novas and colleagues in 2005, it was named in honor of Pablo Puerta and Santiago Reuil, who discovered and prepared the specimen. It consists of four well-preserved vertebrae, including one cervical, one dorsal, and two caudal vertebrae. Puertasaurus is a member of Titanosauria, the dominant group of sauropods during the Cretaceous.
Amargatitanis is a genus of dicraeosaurid sauropod dinosaur from the Barremian-age La Amarga Formation of Neuquén, Argentina. It is known from a single, incomplete postcranial skeleton consisting of a partial hindlimb, ischium, and two vertebrae. These remains were unearthed by Argentine paleontologist José Bonaparte in 1983 during an expedition by the Museo Argentino de Ciencias Naturales and later described as a new genus and species, Amargatitanis macni by Sebastián Apesteguía. The genus name comes from the words Amarga, where the fossils were collected, and titanis meaning "titan". Its species name is in reference to the MACN, where the remains are stored.
Diamantinasaurus is a genus of titanosaurian sauropod from Australia that lived during the early Late Cretaceous, about 94 million years ago. The type species of the genus is D. matildae, first described and named in 2009 by Scott Hocknull and colleagues based on fossil finds in the Winton Formation. Meaning "Diamantina lizard", the name is derived from the location of the nearby Diamantina River and the Greek word sauros, "lizard". The specific epithet is from the Australian song Waltzing Matilda, also the locality of the holotype and paratype. The known skeleton includes most of the forelimb, shoulder girdle, pelvis, hindlimb and ribs of the holotype, and one shoulder bone, a radius and some vertebrae of the paratype.
Bonapartenykus is a monospecific genus of alvarezsauroid dinosaur from Argentina that lived during the Late Cretaceous (Campanian-Maastrichtian) in what is now the upper Allen Formation of the Río Negro Province. The type and only species, Bonapartenykus ultimus, is known from a nearly articulated but partial skeleton that was found in close association to two incomplete eggs and several clusters of eggshells belonging to the oogenus Arriagadoolithus. Bonapartenykus was named in 2012 by Federico L. Agnolin, Jaime E. Powell, Fernando E. Novas and Martin Kundrát. Bonapartenykus has an estimated length of 2.5 m (8.2 ft) and weight of 72 kg (159 lb), making it the largest member of the clade Alvarezsauroidea.
Elaltitan is an extinct genus of large lithostrotian titanosaur sauropod dinosaur known from the Late Cretaceous of Chubut Province, southern Argentina. It contains a single species, Elaltitan lilloi.
Overosaurus is an extinct genus of sauropod dinosaurs, containing only a single species, Overosaurus paradasorum. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia. Overosaurus paradasorum was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately 10 m (33 ft). It was a ground-dwelling herbivore.
Katepensaurus is an extinct genus of rebbachisaurid sauropod dinosaur known from the Late Cretaceous of south-central Chubut Province of central Patagonia, Argentina. It contains a single species, Katepensaurus goicoecheai.
Quetecsaurus is a genus of titanosaurian sauropod dinosaur known from the Late Cretaceous of the southern Mendoza Province, western Argentina. It contains a single species, Quetecsaurus rusconii.
Pilmatueia is a diplodocoid sauropod belonging to the family Dicraeosauridae that lived in Argentina during the Early Cretaceous. Its type and only species is Pilmatueia faundezi. Pilmatueia was probably closely related to other South American dicraeosaurids such as Amargasaurus. Pilmatueia had relatively pneumatic vertebrae compared to other dicraeosaurids, which were otherwise characterized by a reduction in pneumaticity relative to other sauropods. Pilmatueia dates to the Valanginian, an age of the Cretaceous period for which dinosaur faunas are poorly known.
Nullotitan is a genus of lithostrotian titanosaur from the Chorrillo Formation from Santa Cruz Province in Argentina. The type and only species is Nullotitan glaciaris. It was a contemporary of the ornithopod Isasicursor which was described in the same paper.
Punatitan is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Ciénaga del Río Huaco Formation of La Rioja, Argentina. It contains one species, Punatitan coughlini.
Barrosasuchus is a genus of peirosaurid notosuchian from the Santonian of Argentina and part of the extensive peirosaurid record of Late Cretaceous Patagonia. It contains one species, Barrosasuchus neuquenianus. B. neuquenianus is known from an almost complete skull and the majority of the articulated postcranial skeleton, making it the best preserved Patagonian peirosaurid.