Rhomaleopakhus

Last updated

Rhomaleopakhus
Temporal range: Late Jurassic 145  Ma
O
S
D
C
P
T
J
K
Pg
N
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Sauropoda
Family: Mamenchisauridae
Genus: Rhomaleopakhus
Upchurch et al., 2021
Species:
R. turpanensis
Binomial name
Rhomaleopakhus turpanensis
Upchurch et al., 2021

Rhomaleopakhus (meaning "strong forearm") is a genus of mamenchisaurid sauropod, dinosaur from the Late Jurassic Kalaza Formation of China. The type and only species is Rhomaleopakhus turpanensis. [1]

Contents

History

The holotype IVPP-V11121-1, was found by a Chinese-Japanese Chunichi Shinibun expedition near Qiketai in Shanshan, Xinjiang province in 1993, along with the holotype of Hudiesaurus , [2] and is a partially complete forelimb consisting of a humerus, ulna, radius, one carpal, and a partially complete manus that was originally assigned to the coeval mamenchisaurid Hudiesaurus . [2] Paul Upchurch in 2004 rejected the identity because of a lack of overlapping material. [3] However, in a 2021 reassessment of the latter genus, the forelimb was separated from its hypodigm and named the holotype of a new genus and species, Rhomaleopakhus turpanensis; the specific name refers to the Turpan Basin where the specimen was collected. [1]

Description

Upchurch et al. note that the robustness of the forelimb, after which the taxon was named, convergently evolved in what are called the "Core Mamenchisaurus -like taxa" (CMTs), [4] titanosaurs, and ceratopsids. They believe this correlates with a more flexed orientation of the forearm, an enhanced role of the forearm in feeding, and a more anterior shift in the center of mass. It is possible that CMTs and titanosaurs specialized in a feeding strategy that involved efficient locomotion between sparsely-located food sources. [1]

Classification

Upchurch et al. used Moore et al. (2020)'s [4] phylogenetic analysis to determine the relationships of Rhomaleopakhus. They found it to be a mamenchisaurid sister to Chuanjiesaurus, in a clade that also includes Analong. Their cladogram is shown below: [1]

Mamenchisauridae

Tienshanosaurus

Omeisaurus junghsiensis

Wamweracaudia

Analong

Chuanjiesaurus

Rhomaleopakhus

Hudiesaurus

"Mamenchisaurus" hochuanensis (referred)

"Mamenchisaurus" youngi

Xinjiangtitan

"Mamenchisaurus" hochuanensis (holotype)

Shishugou cervicodorsals

Klamelisaurus

Phu Kradung taxon

Related Research Articles

<i>Mamenchisaurus</i> Sauropod dinosaur genus from Late Jurassic-Early Cretaceous Period

Mamenchisaurus is a genus of sauropod dinosaur known for their remarkably long necks which made up nearly half the total body length. Numerous species have been assigned to the genus; however, the validity of these assignments has been questioned. Fossils have been found in the Sichuan Basin and Yunnan Province in China. Several species from the Upper Shaximiao Formation, whose geologic age is uncertain, have been described. However, evidence suggests this formation to be no earlier than the Oxfordian stage of the Late Jurassic. M. sinocanadorum dates to the Oxfordian stage, and M. anyuensis to the Aptian stage of the Early Cretaceous. Most species were medium-large to large sauropods, measuring roughly 15 to 26 meters in length—possibly up to 35 meters (115 ft), based on two undescribed vertebrae.

<span class="mw-page-title-main">Titanosauria</span> Extinct clade of dinosaurs

Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan, estimated at 37 m (121 ft) long with a weight of 69 tonnes, and the comparably-sized Argentinosaurus and Puertasaurus from the same region.

Agustinia is a genus of sauropod dinosaur from the Early Cretaceous of South America. The genus contains a single species, A. ligabuei, known from a single specimen that was recovered from the Lohan Cura Formation of Neuquén Province in Argentina. It lived about 116–108 million years ago, in the Aptian–Albian stages of the Early Cretaceous Period.

<span class="mw-page-title-main">Macronaria</span> Extinct clade of dinosaurs

Macronaria is a clade of sauropod dinosaurs. Macronarians are named after the large diameter of the nasal opening of their skull, known as the external naris, which exceeded the size of the orbit, the skull opening where the eye is located. Fossil evidence suggests that macronarian dinosaurs lived from the Middle Jurassic (Bathonian) through the Late Cretaceous (Maastrichtian). Macronarians have been found globally, including discoveries in Argentina, the United States, Portugal, China, and Tanzania. Like other sauropods, they are known to have inhabited primarily terrestrial areas, and little evidence exists to suggest that they spent much time in coastal environments. Macronarians are diagnosed through their distinct characters on their skulls, as well as appendicular and vertebral characters. Macronaria is composed of several subclades and families notably including Camarasauridae and Titanosauriformes, among several others. Titanosauriforms are particularly well known for being some of the largest terrestrial animals to ever exist.

<i>Euhelopus</i> Extinct genus of dinosaurs

Euhelopus is a genus of sauropod dinosaur that lived between 145 and 133 million years ago during the Berriasian and Valanginian ages of the Early Cretaceous in what is now Shandong Province in China. It was a large quadrupedal herbivore. Like sauropods such as brachiosaurs and titanosaurs, Euhelopus had longer forelegs than hind legs. This discovery was paleontologically significant because it represented the first dinosaur scientifically investigated from China: seen in 1913, rediscovered in 1922, and excavated in 1923 and studied by T'an during the same year. Unlike most sauropod specimens, it has a relatively complete skull.

Chinshakiangosaurus is a genus of dinosaur and probably one of the most basal sauropods known. The only species, Chinshakiangosaurus chunghoensis, is known from a fragmentary skeleton found in Lower Jurassic rocks in China. Chinshakiangosaurus is one of the few basal sauropods with preserved skull bones and therefore important for the understanding of the early evolution of this group. It shows that early sauropods may have possessed fleshy cheeks.

<i>Hudiesaurus</i> Extinct genus of dinosaurs

Hudiesaurus is a herbivorous sauropod genus of dinosaur from China. Its fossil remains were found in 1993 by a Chinese-Japanese expedition near Qiketai in Shanshan, Xinjiang province. The genus contains a single species, Hudiesaurus sinojapanorum, was named and described by Dong Zhiming in 1997. The generic name is derived from Mandarin hudie, meaning "butterfly," and refers to a flat butterfly-shaped process on the front base of the vertebral spine. The specific name refers to the members of the Sino-Japan Silk Road Dinosaur Expedition but can also be read as "central part" in Chinese, a pun on the Japanese Chunichi Shimbun press group, which financed the research.

<i>Klamelisaurus</i> Extinct genus of dinosaurs

Klamelisaurus is a genus of herbivorous sauropod dinosaur from the Middle Jurassic Shishugou Formation of China. The type species is Klamelisaurus gobiensis, which was named by Zhao Xijin in 1993, based on a partial skeleton discovered in 1982 near the abandoned town of Jiangjunmiao. Zhao described Klamelisaurus as the only member of a new subfamily, Klamelisaurinae, among the now-defunct primitive sauropod order Bothrosauropodoidea. Since Zhao's description, Klamelisaurus received limited attention from researchers until Andrew Moore and colleagues redescribed it in 2020.

<span class="mw-page-title-main">Camarasauridae</span> Extinct family of dinosaurs

Camarasauridae is a family of sauropod dinosaurs. Among sauropods, camarasaurids are small to medium-sized, with relatively short necks. They are visually identifiable by a short skull with large nares, and broad, spatulate teeth filling a thick jaw. Based on cervical vertebrae and cervical rib biomechanics, camarasaurids most likely moved their necks in a vertical, rather than horizontal, sweeping motion, in contrast to most diplodocids.

Malarguesaurus is a genus of titanosauriform sauropod dinosaur from the Late Cretaceous of Mendoza Province, Argentina. Its fossils, consisting of tail vertebrae, chevrons, ribs, and limb bones, were found in the upper Turonian-lower Coniacian Portezuelo Formation of the Neuquén Group. The type species, described by González Riga et al. in 2008, is M. florenciae.

The Kalaza Formation is a geological formation in Xinjiang, China whose strata date back to the Late Jurassic. There is some confusion with the stratigraphy of this unit, as the term is used for sediments of equivalent age in both the Junggar Basin and the Turpan Basin. Dinosaur remains are among the fossils that have been recovered from formation.

<i>Daxiatitan</i> Extinct genus of dinosaurs

Daxiatitan is a genus of sauropod dinosaur known from the Lower Cretaceous of Gansu, China. Its type and only species is Daxiatitan binglingi. It is known from a single partial skeleton consisting of most of the neck and back vertebrae, two tail vertebrae, a shoulder blade, and a thigh bone. At the time of its discovery in 2008, Daxiatitan was regarded as potentially the largest known dinosaur from China.

<span class="mw-page-title-main">Mamenchisauridae</span> Extinct family of dinosaurs

Mamenchisauridae is a family of sauropod dinosaurs belonging to Eusauropoda known from the Jurassic and Early Cretaceous of Asia and Africa. Some members of the group reached gigantic sizes, amongst the largest of all sauropods.

<i>Xianshanosaurus</i> Extinct genus of dinosaurs

Xianshanosaurus is a genus of sauropod dinosaur from the Early Cretaceous (Aptian-Albian) of the Ruyang Basin in Henan Province, China. Its type and only species is Xianshanosaurus shijiagouensis. It was described in 2009 by a team of paleontologists led by Lü Junchang. Xianshanosaurus may be a titanosaur, and Daxiatitan may be its closest relative, but its evolutionary relationships remain controversial.

<i>Xinjiangtitan</i> Extinct genus of reptiles

Xinjiangtitan is an extinct genus of mamenchisaurid sauropod that lived during the Middle Jurassic of what is now Xinjiang, northwestern China. Its type and only species is Xinjiangtitan shanshanesis, known from a single incomplete skeleton recovered from the Qiketai Formation. The holotype preserves one of the most complete vertebral columns of any sauropod found in Asia, and has the longest complete neck known for any animal.

Wamweracaudia is a large herbivorous sauropod dinosaur from the Late Jurassic Tendaguru Formation of Tanzania, Africa, 155-145 million years ago.

Fushanosaurus is a genus of sauropod dinosaur from the Shishugou Formation from Xinjiang Province in China. The type and only species is Fushanosaurus qitaiensis. It is solely known from the holotype specimen FH000101, a complete right femur.

Ruixinia is an extinct genus of somphospondylan titanosauriform dinosaur from the Early Cretaceous (Barremian) Yixian Formation of China. The genus contains a single species, Ruixinia zhangi. The Ruixinia holotype is a partial articulated skeleton with the most complete series of caudal vertebrae known from any Asian titanosauriform.

References

  1. 1 2 3 4 Upchurch P, Mannion PD, Xu X, Barrett PM (2021). "Re-assessment of the Late Jurassic eusauropod dinosaur Hudiesaurus sinojapanorum Dong, 1997, from the Turpan Basin, China, and the evolution of hyper-robust antebrachia in sauropods". Journal of Vertebrate Paleontology. 41 (4): e1994414. doi: 10.1080/02724634.2021.1994414 . S2CID   245164168.
  2. 1 2 Dong, Z. (1997). "A gigantic sauropod (Hudiesaurus sinojapanorum gen. et sp. nov.) from the Turpan Basin, China." Pp. 102-110 in Dong, Z. (ed.), Sino-Japanese Silk Road Dinosaur Expedition. China Ocean Press, Beijing.
  3. Upchurch, P.; Barrett, P.M.; Dodson, P.; 2004 "Sauropoda". In: Weishampel, D.B. and Dodson, P. and Osmolska, H., (eds.) The Dinosauria. p 259 - 322. University of California Press: Berkeley and Los Angeles
  4. 1 2 Moore, A.J.; Upchurch, P.; Barrett, P.M.; Clark, J.M.; Xing, X. (2020). "Osteology of Klamelisaurus gobiensis (Dinosauria, Eusauropoda) and the evolutionary history of Middle–Late Jurassic Chinese sauropods". Journal of Systematic Palaeontology. 18 (16): 1299–1393. doi:10.1080/14772019.2020.1759706. S2CID   219749618.