Lusotitan Temporal range: Late Jurassic, | |
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Holotype humerus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Macronaria |
Family: | † Brachiosauridae |
Genus: | † Lusotitan Antunes & Mateus, 2003 |
Species: | †L. atalaiensis |
Binomial name | |
†Lusotitan atalaiensis (Lapparent & Zbyszewski, 1957 [originally Brachiosaurus]) | |
Synonyms | |
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Lusotitan is a genus of herbivorous brachiosaurid sauropod dinosaur from the Late Jurassic of Portugal and possibly Spain.
In 1947 Manuel de Matos, a member of the Geological Survey of Portugal, discovered large sauropod fossils in the Portuguese Lourinhã Formation that date back to the Tithonian stage of the Late Jurassic period. [1] In 1957 Albert-Félix de Lapparent and Georges Zbyszewski named the remains as a new species of Brachiosaurus : Brachiosaurus atalaiensis. [2] The specific name referred to the site Atalaia. In 2003 Octávio Mateus and Miguel Telles Antunes named it as a separate genus: Lusotitan. The type species is Lusotitan atalaiensis. The generic name is derived from Luso, the Latin name for an inhabitant of Lusitania, and from the Greek word "Titan", a mythological giant. [1]
The finds consisted of a partial skeleton lacking the skull and individual vertebrae uncovered in several locations. De Lapparent did not assign a holotype. In 2003 Mateus chose the skeleton as the lectotype. Its bones have the inventory numbers MIGM 4798, 4801–10, 4938, 4944, 4950, 4952, 4958, 4964–6, 4981–2, 4985, 8807, and 8793-5. These remains include 28 vertebrae and elements of the appendicular skeleton.
The lectotype was re-described by Mannion and colleagues in 2013. [3] In 2017, Mocho, Royo-Torres and Ortega suggested that Galvesaurus or Galveosaurus from the Late Jurassic of Spain might represent a junior synonym of this taxon. [4] However, a 2019 description of new material of Galvesaurus by Perez-Pueyo et al. identified phylogenetically informative characters to distinguish it from Lusotitan which was recovered as its sister taxon. [5]
Lusotitan was a large sauropod, reaching 21 m (69 ft) in length and 30 tonnes (33 short tons) in body mass. [6] It had long forearms with the humerus and femur measuring 2.05 m (6.7 ft) and 2 m (6.6 ft) in length, respectively. [7]
The Lourinhã Formation of western Portugal was likely to be formed during the Kimmeridgian or Tithonian ages of the Late Jurassic period. The area is a coastal region with a strong marine influence. Its flora and fauna are similar to the Morrison Formation in the United States, and the Tendaguru Formation in Tanzania. Lusotitan is the largest dinosaur that has been discovered in the area. Lusotitan lived alongside species of the predatory theropods Allosaurus (A. europaeus), Ceratosaurus , Lourinhanosaurus , and Torvosaurus , the ankylosaurian Dracopelta , the sauropods Bothriospondylus , Lourinhasaurus , and Zby , and the stegosaurs Dacentrurus and Miragaia . [7]
Torvosaurus is a genus of large megalosaurine theropod dinosaur that lived approximately 165 to 148 million years ago during the Callovian to Tithonian ages of the late Middle and Late Jurassic period in what is now Colorado, Portugal, Germany, and possibly England, Spain, Tanzania, and Uruguay. It contains two currently recognized species, Torvosaurus tanneri and Torvosaurus gurneyi, plus a third unnamed species from Germany.
Giraffatitan is a genus of sauropod dinosaur that lived during the late Jurassic Period in what is now Lindi Region, Tanzania. Only one species is known, G. brancai, named in honor of German paleontologist Wilhelm von Branca, who was a driving force behind the expedition that discovered it in the Tendaguru Formation. Giraffatitan brancai was originally described by German paleontologist Werner Janensch as a species of the North American sauropod Brachiosaurus from the Morrison Formation, as Brachiosaurus brancai. Recent research shows that the differences between the type species of Brachiosaurus and the Tendaguru material are so large that the African material should be placed in a separate genus.
Bothriospondylus is a dubious genus of neosauropod sauropod dinosaur. It lived during the Late Jurassic in England, and the type and only species is B. suffossus.
Dracopelta is a monospecific genus of ankylosaur dinosaur from Portugal that lived during the Late Jurassic in what is now the Lourinhã Formation. The type and only species is Dracopelta zbyszewskii, which is represented by a partial skeleton including unpublished material.
Dinheirosaurus is a genus of diplodocid sauropod dinosaur that is known from fossils uncovered in modern-day Portugal. It may represent a species of Supersaurus. The only species is Dinheirosaurus lourinhanensis, first described by José Bonaparte and Octávio Mateus in 1999 for vertebrae and some other material from the Lourinhã Formation. Although the precise age of the formation is not known, it can be dated around the early Tithonian of the Late Jurassic.
Lourinhasaurus is an extinct genus of herbivorous sauropod dinosaur genus dating from Late Jurassic strata of Estremadura, Portugal.
Lourinhanosaurus was a genus of carnivorous theropod dinosaur that lived during the Late Jurassic Period (Kimmeridgian/Tithonian) in Portugal. It is one of many large predators discovered at the Lourinhã Formation and probably competed with coeval Torvosaurus gurneyi, Allosaurus europaeus, and Ceratosaurus.
Europasaurus is a basal macronarian sauropod, a form of quadrupedal herbivorous dinosaur. It lived during the Late Jurassic of northern Germany, and has been identified as an example of insular dwarfism resulting from the isolation of a sauropod population on an island within the Lower Saxony basin.
Galvesaurus, or Galveosaurus, is a genus of brachiosaurid sauropod dinosaur from the Late Jurassic period. Fossils of the only known species, G. herreroi, were found in Galve, Spain, hence its generic name, "Galve lizard". The specific name G. herreroi honours the discoverer, José María Herrero. Some researchers suggest that the taxon might represent a junior synonym of Lusotitan.
Turiasauria is an unranked clade of basal sauropod dinosaurs known from Middle Jurassic to Early Cretaceous deposits in Europe, North America, and Africa.
Octávio Mateus is a Portuguese dinosaur paleontologist and biologist Professor of Paleontology at the Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa. He graduated in Universidade de Évora and received his PhD at Universidade Nova de Lisboa in 2005. He collaborates with Museu da Lourinhã, known for their dinosaur collection.
Brohisaurus is a genus of titanosauriform sauropod dinosaur from the Late Jurassic (Kimmeridgian-aged) Sembar Formation of Pakistan. The type species is B. kirthari.
The Lourinhã Formation is a geological formation in western Portugal, outstanding for its abundant fossilized fauna and flora, including dinosaur bones, dinosaur eggs and nests, and fossil tracks, to name but some examples.
At its upper limit, the formation includes some Early Cretaceous (Berriasian) strata, but it mostly consists of Late Jurassic (Kimmeridgian/Tithonian) rocks. The stratigraphy of the formation is generally complex and controversial, with the classification and grouping of its lithostratigraphic units varying between different stratigraphers.
The fossil biota present there is very similar to that of the Morrison Formation in the United States and, to a lesser extent, the Tendaguru Formation in Tanzania. There are also similarities to the Villar del Arzobispo Formation in Spain, and the neardy Alcobaça Formation.
The formation is named after the municipality of Lourinhã, the geographic area it mostly occupies.
Brachiosaurus is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154 to 150 million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in the Colorado River valley in western Colorado, United States. Riggs named the dinosaur Brachiosaurus altithorax; the generic name is Greek for "arm lizard", in reference to its proportionately long arms, and the specific name means "deep chest". Brachiosaurus is estimated to have been between 18 and 22 meters long; body mass estimates of the subadult holotype specimen range from 28.3 to 46.9 metric tons. It had a disproportionately long neck, small skull, and large overall size, all of which are typical for sauropods. Atypically, Brachiosaurus had longer forelimbs than hindlimbs, which resulted in a steeply inclined trunk, and a proportionally shorter tail.
Miragaia is a long-necked stegosaurid dinosaur. Its fossils have been found in Upper Jurassic rocks in Portugal and possibly also Wyoming, United States. Miragaia has the longest neck known for any stegosaurian, which included at least seventeen vertebrae. Some researchers consider this taxon to be a junior synonym of Dacentrurus.
The Villar del Arzobispo Formation is a Late Jurassic to possibly Early Cretaceous geologic formation in eastern Spain. It is equivalent in age to the Lourinhã Formation of Portugal. It was originally thought to date from the Late Tithonian-Middle Berriasian, but more recent work suggests a Kimmeridigan-Late Tithonian, possibly dating to the Early Berriasian in some areas. The Villar del Arzobispo Formation's age in the area of Riodeva in Spain has been dated based on stratigraphic correlations as middle-upper Tithonian, approximately 145-141 million years old. In the area of Galve, the formation potentially dates into the earliest Cretaceous.
Zby is an extinct genus of turiasaurian sauropod dinosaur known from the Late Jurassic of the Lourinhã Formation, central west Portugal. It contains a single species, Zby atlanticus. It is named after Georges Zbyszewski, who studied the geology and paleontology of Portugal.
Oceanotitan is a genus of sauropod dinosaur known from the Upper Jurassic Lourinha Formation of Portugal. It is represented by a single specimen consisting of several tail vertebrae and appendicular bones. It contains one species, Oceanotitan dantasi. Oceanotitan is classified as possibly one of the earliest members of the Somphospondyli, a group of sauropods that includes the titanosaurs.
Oblitosaurus is a genus of ankylopollexian ornithopod dinosaur from the Late Jurassic Villar del Arzobispo Formation of Spain. The type species is Oblitosaurus bunnueli.
Ardetosaurus is an extinct genus of diplodocid sauropod dinosaurs from the Late Jurassic (Kimmeridgian) Morrison Formation of northern Wyoming, United States. The genus contains a single species, Ardetosaurus viator. It was first described in 2024 on the basis of a partial articulated skeleton, including vertebrae from the neck, back, and tail, hip bones, and part of the left hindlimb. The genus is a member of the Diplodocinae, a subfamily of large long-necked dinosaurs with whiplike tails. Ardetosaurus represents one of many distinct sauropod taxa that coexisted in this formation.