Garrigatitan Temporal range: Late Cretaceous, | |
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Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | † Sauropodomorpha |
Clade: | † Sauropoda |
Clade: | † Macronaria |
Clade: | † Titanosauria |
Clade: | † Lithostrotia |
Subfamily: | † Lirainosaurinae |
Genus: | † Garrigatitan Díaz et al., 2021 |
Species: | †G. meridionalis |
Binomial name | |
†Garrigatitan meridionalis Díaz et al., 2021 | |
Garrigatitan (meaning "garrigue giant") is a genus of titanosaurian dinosaur from the late Cretaceous Period of the Grès à Reptiles Formation in France. The genus contains a single species, Garrigatitan meridionalis. [1]
Between 2009 and 2012, excavations were carried out at Velaux-La Bastide Neuve by the Palaios Association and the University of Poitiers. During the excavations, the holotype of Garrigatitan was discovered along with the remains of Atsinganosaurus, another titanosaurian. [2]
In 2021, the type species Garrigatitan meridionalis was named and described by Verónica Díez Díaz, Géraldine Garcia, Xabier Pereda Suberbiola, Benjamin Jentgen-Ceschino, Koen Stein, Pascal Godefroit and Xavier Valentin. The holotype, MMS / VBN.09.17, was found in a layer of sandstone of the Begudian, the second level of the second series, dating back to the late Campanian. It consists of a sacrum belonging to an immature individual. [1]
Additional fossil material has been assigned to the species including a cervical vertebra, two humeri, a left ilium, and a right ischial bone. Other specimens recovered include a neural spine, a right humerus, part of the right leg, and a left femur. The assigned specimens come from the third level of the second series. The fossils were found within an area of 375 square meters and a thickness of 1.2 metres (3.9 ft). They were not associated and presumably represent different individuals. They are all part of the Moulin Seigneurial de Velaux collection. [1]
The generic name, "Garrigatitan," is a combination of the Occitan "garriga," meaning "dry thicket", referring to a type of Mediterranean vegetation characterized by drought-resistant shrubs, and the Greek "titan", after the Greek mythological family of giants. The specific epithet, "meridonalis," means "southern" in Latin, in reference to the discovery location in southern France. [1]
Some Garrigatitan specimens regarded as subadults or adults belonged to individuals measuring 4–6 metres (13–20 ft) long and weighing 2–2.5 metric tons (2.2–2.8 short tons). A tentatively referred specimen belonged to an adult individual measuring about 12–16 metres (39–52 ft) long. [1]
Cladistic analysis of Garrigatitan shows that it belonged to the subfamily Lirainosaurinae. [1]
Lirainosaurinae |
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Ampelosaurus is a titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now France. Its type species is A. atacis, named by Le Loeuff in 1995. Its remains were found in a level dating from 71.5 million years ago representing the early Maastrichtian.
Pararhabdodon is a genus of tsintaosaurin hadrosaurid dinosaur, from the Maastrichtian-age Upper Cretaceous Tremp Group of Spain. The first remains were discovered from the Sant Romà d’Abella fossil locality and assigned to the genus Rhabdodon, and later named as the distinct species Pararhabdodon isonensis in 1993. Known material includes assorted postcranial remains, mostly vertebrae, as well as maxillae from the skull. Specimens from other sites, including remains from France, a maxilla previously considered the distinct taxon Koutalisaurus kohlerorum, an additional maxilla from another locality, the material assigned to the genera Blasisaurus and Arenysaurus, and the extensive Basturs Poble bonebed have been considered at different times to belong to the species, but all of these assignments have more recently been questioned. It was one of the last non-avian dinosaurs known from the fossil record that went extinct during the Cretaceous-Paleogene extinction event.
Lirainosaurus is a genus of titanosaur sauropod which lived in what is now Spain. The type species, Lirainosaurus astibiae, was described by Sanz, Powell, Le Loeuff, Martinez, and Pereda-Suberbiola in 1999. It was a relatively small sauropod, measuring 4 metres (13 ft) long, possibly up to 6 metres (20 ft) long for the largest individuals, and weighed about 2–4 metric tons.
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Argiles et Grès à Reptiles Formation also known as the Argiles Rutilantes Formation is an early Maastrichtian French geologic formation in the département of Var preserving the remains of several types of dinosaurs and other extinct organisms.
Opisthocoelicaudiinae is a subfamily of titanosaurian dinosaurs from the Late Cretaceous. It was named by John McIntosh in 1990. Opisthocoelicaudiines are known from Mongolia, Argentina, and the United States. Two genera were assigned to Opisthocoelicaudiinae by Gonzalez et al. (2009): Alamosaurus and Opisthocoelicaudia, a conclusion also found by Díez Díaz et al. (2018). The hands of opisthocoelicaudiines lacked wrist bones and phalanges.
Atsinganosaurus is a genus of titanosaurian sauropod dinosaur which existed in what is now France during the Late Cretaceous. Well-preserved remains of Atsinganosaurus were collected from the Grès à Reptiles Formation of the Aix-en-Provence Basin. The type and only species is A. velauciensis.
Paludititan is a genus of titanosaurian sauropod dinosaur which lived in the area of present Romania during the Late Cretaceous. It existed in the island ecosystem known as Hațeg Island.
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Aralosaurini is a proposed tribe of hadrosaurid dinosaurs belonging to the subfamily Lambeosaurinae. The members of this group lived in Asia and Europe during the end of the Late Cretaceous about 83.6 to 66.0 million years ago. The clade may not be monophyletic, with Canardia and Aralosaurus potentially instead being unrelated primitive members of the subfamily.
Lohuecotitan is an extinct genus of titanosaurian sauropod dinosaur which lived during the Late Cretaceous in Spain. The only species known in the genus is Lohuecotitan pandafilandi, described and named in 2016.
Chenanisaurus is a genus of predatory abelisaurid dinosaur, with a single known species C. barbaricus. It comes from the upper Maastrichtian phosphates of the Ouled Abdoun Basin in Morocco, North Africa. The animal is known from a holotype, consisting of a partial jaw bone, and several isolated teeth found in the same beds. Chenanisaurus is one of the largest members of the Abelisauridae, and one of the last, being a contemporary of the North American Tyrannosaurus. It would have been among the dinosaur species wiped out by the Chicxulub asteroid impact and the Cretaceous-Paleogene mass extinction that followed.
Matheronodon is a genus of rhabdodontid ornithopod dinosaur from the late Cretaceous Period of the Grès à Reptiles Formation in France. The genus contains a single species, M. provincialis, which is known from a single maxilla and associated teeth. It was named by Pascal Godefroit and colleagues in 2017. The teeth of Matheronodon are large but few in number. The teeth are also in an unusual arrangement, emerging alternatingly from one of a pair of fused tooth sockets in its mouth. In life, the teeth would have functioned like a pair of scissors, allowing Matheronodon to feed on the tough leaves of monocot plants.
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