Paralititan

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Paralititan
Temporal range: Late Cretaceous, 99.6–93.5  Ma
Paralititan stromeri by Hatem Moushir 9.JPG
Humeri at the Egyptian Geological Museum
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Sauropoda
Clade: Macronaria
Clade: Titanosauria
Clade: Lithostrotia
Genus: Paralititan
Smith et al., 2001
Type species
Paralititan stromeri
Smith et al., 2001

Paralititan (meaning "tidal giant" [1] ) was a giant titanosaurian sauropod dinosaur genus discovered in coastal deposits in the Upper Cretaceous Bahariya Formation of Egypt. It lived between 99.6 and 93.5 million years ago. [2]

Contents

Discovery

First caudal vertebra and other fossils Paralititan stromeri by Hatem Moushir 3.JPG
First caudal vertebra and other fossils

Joshua Smith in 1999 in the Bahariya Oasis rediscovered the Gebel el Dist site where Richard Markgraf in 1912, 1913 and 1914 had excavated fossils for Ernst Stromer. In 2000, an American expedition was mounted to revisit the site. However, apparently Markgraf had already removed all more complete skeletons, leaving only limited remains behind. At a new site, the nearby Gebel Fagga, the expedition succeeded in locating a partial sauropod skeleton. [3] It was identified by Lacovara as a species new to science. It was named and described by Joshua B. Smith, Matthew C. Lamanna, Kenneth J. Lacovara, Peter Dodson, Jennifer R. Smith, Jason Charles Poole, Robert Giegengack and Yousri Attia in 2001 as the type species Paralititan stromeri. The generic name means "Stromer's tidal (Greek para + halos "near sea") titan" or "Stromer's tidal giant", in reference to the "paralic" tidal flats the animal lived on. [4] The specific name honors Ernst Stromer von Reichenbach, a German paleontologist and geologist who first established the presence of dinosaur fossils in this area in 1911. Paralititan represents the first tetrapod reported from the Bahariya Formation since Romer's publication of 1935. [1]

The holotype specimen of Paralititan, CGM 81119, was found in a layer of the Bahariya Formation, dating from the Cenomanian. It consists of a partial skeleton lacking the skull. It is incomplete, apart from bone fragments containing two fused posterior sacral vertebrae, two anterior caudal vertebrae, both incomplete scapulae, two humeri and a metacarpal. The Paralititan type specimen shows evidence of having been scavenged by a carnivorous dinosaur as it was disarticulated within an oval of eight metres length with the various bones being clustered. A Carcharodontosaurus tooth was discovered in between the clusters. The holotype is part of the collection of the Cairo Geological Museum. [1]

The large anterior dorsal vertebra 1912V11164, in 1932 by Stromer referred to an undetermined "Giant Sauropod", was in 2001 tentatively referred to Paralititan. [1]

Description

Hypothetical scale diagram comparing Paralititan to some humans, with some of the known material in white. Paralititan-Scale-Diagram-SVG-Steveoc86.svg
Hypothetical scale diagram comparing Paralititan to some humans, with some of the known material in white.

Joshua Smith, who informally led the research team that found the dinosaur fossils, told an interviewer, "It was a truly enormous dinosaur by any reckoning." [5]

Life restoration Paralititan stromeri.jpg
Life restoration

Little of Paralititan is known, so its exact size is difficult to estimate. However, the limited material, especially the long humeri, suggested that it is one of the most massive dinosaurs ever discovered, with an estimated weight of 59 t (65 short tons). [6] The complete right humerus measured 1.69 meters (5.54 ft) long which at the time of discovery was the longest known in a Cretaceous sauropod; this was surpassed in 2016 with the discovery of Notocolossus which had a 1.76 m (5 ft 9 in) humerus. [7] Using Saltasaurus as a guide, Carpenter estimated its length at around 26 m (85 ft) in 2006. [8]

Scott Hartman estimates an animal that is massive, but still smaller than the biggest titanosaurs such as Puertasaurus , Alamosaurus , and Argentinosaurus . [9] In 2010, Gregory S. Paul estimated its length at 20+ meters (66+ ft), and its weight at 20 tonnes (24.2 short tons). [10] In 2012 Holtz gave a length of 32 meters (105 ft) and an estimated weight of 65.3–72.5 tonnes (72–80 short tons). [2] In 2016, using equations that estimate body mass based on the circumference of the humerus and femur of quadrupedal animals, it was given an estimated weight of ~50 t (55 short tons). [7] In 2019 Gregory S. Paul estimated Paralititan between 30–55 tonnes (33–60.6 short tons). [11] In 2020 Molina-Perez and Larramendi estimated the size of the animal at 27 meters (88.6 ft) and 30 tonnes (33 short tons). [12]

From the formation another sauropod had already been known, Aegyptosaurus . Paralititan differs from Aegyptosaurus in its larger size, the latter genus weighing only fifteen tons, possibly in not having pleurocoels in its front tail vertebrae, and in possessing a relatively longer deltopectoral crest on its humerus. [1]

Palaeoenvironment

Restoration of Paralititan with contemporaneous animals of the Bahariya Formation Bahariya Formation McAfee.jpg
Restoration of Paralititan with contemporaneous animals of the Bahariya Formation

The autochthonous, scavenged skeleton was preserved in tidal flat deposits containing in the form of fossil leaves and root systems, a mangrove vegetation of seed ferns, Weichselia reticulata. The mangrove ecosystem it inhabited was situated along the southern shore of the Tethys Sea. Paralititan is the first dinosaur demonstrated to have inhabited a mangrove habitat. It lived at approximately the same time and place as giant predators Carcharodontosaurus , Spinosaurus , and the sauropod Aegyptosaurus . [1]

Related Research Articles

<i>Argentinosaurus</i> Late Cretaceous giant sauropod dinosaur genus

Argentinosaurus is a genus of giant sauropod dinosaur that lived during the Late Cretaceous period in what is now Argentina. Although it is only known from fragmentary remains, Argentinosaurus is one of the largest known land animals of all time, perhaps the largest, measuring 30–35 metres (98–115 ft) long and weighing 65–80 tonnes. It was a member of Titanosauria, the dominant group of sauropods during the Cretaceous. It is widely regarded by many paleontologists as the biggest dinosaur ever, and perhaps lengthwise the longest animal ever, though both claims have no concrete evidence yet.

<i>Spinosaurus</i> Genus of spinosaurid dinosaur

Spinosaurus is a genus of spinosaurid dinosaur that lived in what now is North Africa during the Cenomanian to upper Turonian stages of the Late Cretaceous period, about 99 to 93.5 million years ago. The genus was known first from Egyptian remains discovered in 1912 and described by German palaeontologist Ernst Stromer in 1915. The original remains were destroyed in World War II, but additional material came to light in the early 21st century. It is unclear whether one or two species are represented in the fossils reported in the scientific literature. The best known species is S. aegyptiacus from Egypt, although a potential second species, S. maroccanus, has been recovered from Morocco. The contemporary spinosaurid genus Sigilmassasaurus has also been synonymized by some authors with S. aegyptiacus, though other researchers propose it to be a distinct taxon. Another possible junior synonym is Oxalaia from the Alcântara Formation in Brazil.

<i>Sauroposeidon</i> Sauropod dinosaur genus from the Early Cretaceous period

Sauroposeidon is a genus of sauropod dinosaur known from several incomplete specimens including a bone bed and fossilized trackways that have been found in the U.S. states of Oklahoma, Wyoming, and Texas.

<i>Carcharodontosaurus</i> Genus of carcharodontosaurid dinosaur from the Cretaceous period

Carcharodontosaurus is a genus of carnivorous theropod dinosaur that lived in North Africa from about 99 to 94 million years ago during the Cenomanian stage of the Late Cretaceous. Two teeth of the genus, now lost, were first described from Algeria by French paleontologists Charles Depéret and Justin Savornin as Megalosaurus saharicus. A partial skeleton was collected by crews of German paleontologist Ernst Stromer during a 1914 expedition to Egypt. Stromer did not report the Egyptian find until 1931, in which he dubbed the novel genus Carcharodontosaurus, making the type species C. saharicus. Unfortunately, this skeleton was destroyed during the Second World War. In 1995 a nearly complete skull of C. saharicus, the first well-preserved specimen to be found in almost a century, was discovered in the Kem Kem Beds of Morocco; it was designated the neotype in 1996. Fossils unearthed from the Echkar Formation of northern Niger were described and named as another species, C. iguidensis, in 2007.

<span class="mw-page-title-main">Ernst Stromer</span> German paleontologist

Ernst Freiherr Stromer von Reichenbach was a German paleontologist best remembered for his expedition to Egypt, during which the discovery of the first known remains of Spinosaurus was made.

<i>Alamosaurus</i> Extinct genus of dinosaurs

Alamosaurus is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs containing a single known species, Alamosaurus sanjuanensis, from the Maastrichtian age of the Late Cretaceous period in what is now southwestern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to Argentinosaurus and Puertasaurus, which would make it the absolute largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age. Specimens of a juvenile Alamosaurus sanjuanensis have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species. Alamosaurus is the only known sauropod to have inhabited North America after their nearly 30-million year absence from the North American fossil record and probably represents an immigrant from South America.

<i>Aegyptosaurus</i> Titanosaurid sauropod dinosaur genus from late Cretaceous Period

Aegyptosaurus is a genus of sauropod dinosaur that lived in what is now Africa, around 95 million years ago, during the Late Cretaceous Period.

<i>Bahariasaurus</i> Extinct genus of dinosaurs

Bahariasaurus is an enigmatic genus of large theropod dinosaur. Bahariasaurus is known to have included at least 1 species, Bahariasaurus ingens, which was found in North African rock layers dating to the Cenomanian and Turonian ages of the Late Cretaceous. The only fossils confidently assigned to Bahariasaurus were found in the Bahariya Formation of the Bahariya oasis in Egypt by Ernst Stromer but were destroyed during a World War II bombing raid with the same raid taking out the holotype of Spinosaurus and Aegyptosaurus among other animals found in the Bahariya Formation. While there have been more fossils assigned to the genus such as some from the Farak Formation of Niger, these remains are referred to with much less certainty. Bahariasaurus is, by most estimations, one of the largest theropods, approaching the height and length of other large bodied theropods such as Tyrannosaurus rex and the contemporaneous Carcharodontosaurus. The aforementioned estimations tend to put it at around 11–12.2 metres in length and 4-4.8 tonnes in overall weight.

<i>Giraffatitan</i> Sauropod dinosaur genus from the late Jurassic Period in Lindi Region, Tanzania

Giraffatitan is a genus of sauropod dinosaur that lived during the late Jurassic Period in what is now Lindi Region, Tanzania. It was originally named as an African species of Brachiosaurus, but this has since been moved to its own genus. Giraffatitan was for many decades known as the largest dinosaur but recent discoveries of several larger dinosaurs prove otherwise; giant titanosaurians appear to have surpassed Giraffatitan in terms of sheer mass. Also, the sauropod dinosaur Sauroposeidon is estimated to be taller and possibly heavier than Giraffatitan.

<i>Isisaurus</i> Extinct genus of dinosaurs

Isisaurus is a genus of titanosaurian dinosaur from the Late Cretaceous Lameta Formation of India and Pab Formation of Pakistan. The genus contains a single species, Isisaurus colberti.

<i>Futalognkosaurus</i> Extinct genus of dinosaurs

Futalognkosaurus is a genus of titanosaurian dinosaur. The herbivorous Futalognkosaurus lived approximately 87 million years ago in the Portezuelo Formation, in what is now Argentina, of the Coniacian stage of the late Cretaceous Period. The fish and fossilized leaf debris on the site, together with other dinosaur remains, suggest a warm tropical climate in Patagonia during this period.

<i>Rinconsaurus</i> Extinct genus of dinosaurs

Rinconsaurus is a genus of titanosaur sauropod dinosaur from the Late Cretaceous in what is now Argentina. The type species, Rinconsaurus caudamirus, was described by Calvo and Riga in 2003, and is based on three partial skeletons.

<i>Puertasaurus</i> Extinct genus of dinosaurs

Puertasaurus is a genus of sauropod dinosaur that lived in South America during the Late Cretaceous Period. It is known from a single specimen recovered from sedimentary rocks of the Cerro Fortaleza Formation in southwestern Patagonia, Argentina, which probably is Campanian or Maastrichtian in age. The only species is Puertasaurus reuili. Described by the paleontologist Fernando Novas and colleagues in 2005, it was named in honor of Pablo Puerta and Santiago Reuil, who discovered and prepared the specimen. It consists of four well-preserved vertebrae, including one cervical, one dorsal, and two caudal vertebrae. Puertasaurus is a member of Titanosauria, the dominant group of sauropods during the Cretaceous.

Egypt has many fossil-bearing geologic formations, in which many dinosaurs have been discovered.

Stromerosuchus is a dubious genus of Late Cretaceous crocodyliform. Fragmentary remains have been found from the Cenomanian-age Bahariya Formation of Egypt. The genus was named in 1936 by Oskar Kuhn. It is named in honor of Ernst Stromer, the German paleontologist who found the fossils in the Bahariya Oasis in 1911 and described them in 1922. After their discovery, the fossils, along with many others found from Bahariya, were in the possession of the Egyptian Geological Survey. In 1922, the fossils were sent back to Stromer, but they were badly crushed in shipment from Egypt. Because the known remains are so poor, the genus is now regarded as a nomen dubium. Some material has been referred to the genera Aegyptosuchus and Stomatosuchus, both named by Stromer from the Bahariya material.

<i>Patagotitan</i> Extinct genus of dinosaurs

Patagotitan is a genus of titanosaurian sauropod dinosaur from the Cerro Barcino Formation in Chubut Province, Patagonia, Argentina. The genus contains a single species known from at least six young adult individuals, Patagotitan mayorum, which was first announced in 2014 and then named in 2017 by José Carballido and colleagues. Preliminary studies and press releases suggested that Patagotitan was the largest known titanosaur and land animal overall, with an estimated length of 37 m (121 ft) and an estimated weight of 69 tonnes. Later research revised the length estimate down to 31 m (102 ft) and weight estimates down to approximately 50–57 tonnes, suggesting that Patagotitan was of a similar size to, if not smaller than, its closest relatives Argentinosaurus and Puertasaurus. Still, Patagotitan is one of the most-known titanosaurs, and so its interrelationships with other titanosaurs have been relatively consistent in phylogenetic analyses. This led to its use in a re-definition of the group Colossosauria by Carballido and colleagues in 2022.

<span class="mw-page-title-main">Kenneth Lacovara</span> American paleontologist

Kenneth John Lacovara is an American paleontologist and geologist at Rowan University and fellow of the Explorers Club, known for the discovery of the titanosaurian dinosaur Dreadnoughtus and his involvement in the discovery and naming of the giant sauropod dinosaur Paralititan, as well as his work applying 3D printing technology to paleontology. Lacovara is the founder of the Edelman Fossil Park of Rowan University and the author of the general-audience book, Why Dinosaurs Matter (2017), for which he received a Nautilus Book Award. Additionally, he serves as Paleontology Fellow of the Academy of Natural Sciences. He is a recipient of the Explorers Club Medal, the highest honor bestowed by The Explorers Club.

<i>Dreadnoughtus</i> Extinct genus of dinosaur

Dreadnoughtus is a genus of titanosaurian sauropod dinosaur containing a single species, Dreadnoughtus schrani. D. schrani is known from two partial skeletons discovered in Upper Cretaceous rocks of the Cerro Fortaleza Formation in Santa Cruz Province, Argentina. It is one of the largest terrestrial vertebrates known, with the immature type specimen measuring 26 metres (85 ft) in total body length and weighing 48–49 metric tons. D. schrani is known from more complete skeletons than any other gigantic titanosaurian.

<i>Notocolossus</i> Extinct genus of dinosaurs

Notocolossus is a genus of titanosaurian sauropod dinosaur from late Cretaceous strata of Mendoza Province, Argentina.

Mnyamawamtuka is a genus of lithostrotian titanosaur sauropod dinosaur from the Cretaceous Galula Formation in Tanzania. The type and only species is M. moyowamkia.

References

  1. 1 2 3 4 5 6 Smith JB, Lamanna MC, Lacovara KJ, Dodson P, Smith JR, Poole JC, Giegengack R, Attia Y (June 2001). "A giant sauropod dinosaur from an Upper Cretaceous mangrove deposit in Egypt" (PDF). Science. 292 (5522): 1704–6. Bibcode:2001Sci...292.1704S. doi:10.1126/science.1060561. PMID   11387472. S2CID   33454060.
  2. 1 2 Holtz, Thomas R. Jr. (2012) Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages, Winter 2011 Appendix.
  3. Nothdurft, William; Joshua Smith; Matt Lamana; Ken Lacovara; Jason Poole & Jen Smith, 2002, The Lost Dinosaurs of Egypt: The Astonishing and Unlikely True Story of One of the Twentieth Century's Greatest Paleontological Discoveries, Random House, 272 pp
  4. "Paralititan Stromeri". Paleobiology Database. Retrieved December 31, 2012.
  5. Roach, John (May 31, 2001). "'Tidal Giant' Roamed Coastal Swamps of Ancient Africa". National Geographic News. Washington, D.C.: National Geographic Society. Archived from the original on June 5, 2001. Retrieved December 31, 2012.
  6. Burness, G.P. and Flannery, T. (2001). "Dinosaurs, Dragonslayer, and Dwarfs: The Evolution of Maximal Body Size." Proceedings of the National Academy of Sciences, 98(25): 14518-14523.
  7. 1 2 González Riga BJ, Lamanna MC, Ortiz David LD, Calvo JO, Coria JP (January 2016). "A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot". Scientific Reports. 6 (1): 19165. Bibcode:2016NatSR...619165G. doi:10.1038/srep19165. PMC   4725985 . PMID   26777391.
  8. Carpenter K (2006). Foster JR, Lucas SG (eds.). "Biggest of the Big: a Critical Re-evaluation of the Mega-sauropod Amphicoelias fragillimus Paleontology and Geology of the Upper Jurassic Morrison Formation" (PDF). New Mexico Museum of Natural History and Science Bulletin. 36: 131–138. Archived from the original (PDF) on 2016-03-08.
  9. Hartman, Scott. "The biggest of the big". Skeletal Drawings.
  10. Paul, G.S., 2010, The Princeton Field Guide to Dinosaurs, Princeton University Press p. 209
  11. Paul, Gregory S. (2019). "Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals" (PDF). Annals of the Carnegie Museum. 85 (4): 335–358. doi:10.2992/007.085.0403. S2CID   210840060.
  12. Molina-Perez & Larramendi (2020). Dinosaur Facts and Figures: The Sauropods and Other Sauropodomorphs. New Jersey: Princeton University Press. p. 267.