Titanosaurs were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as Patagotitan—estimated at 37 m (121 ft) long with a weight of 69 tonnes —and the comparably-sized Argentinosaurus and Puertasaurus from the same region.
Alamosaurus is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs containing a single known species, Alamosaurus sanjuanensis, from the Maastrichtian age of the Late Cretaceous period in what is now southwestern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to Argentinosaurus and Puertasaurus, which would make it the absolute largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age. Specimens of a juvenile Alamosaurus sanjuanensis have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species. Alamosaurus is the only known sauropod to have inhabited North America after their nearly 30-million year absence from the North American fossil record and probably represents an immigrant from South America.
Euhelopus is a genus of sauropod dinosaur that lived between 145 and 133 million years ago during the Berriasian and Valanginian stages of the Early Cretaceous in what is now Shandong Province in China. It was a large quadrupedal herbivore. Like sauropods such as brachiosaurs and titanosaurs, Euhelopus had longer forelegs than hind legs. This discovery was paleontologically significant because it represented the first dinosaur scientifically investigated from China: seen in 1913, rediscovered in 1922, and excavated in 1923 and studied by T'an during the same year. Unlike most sauropod specimens, it has a relatively complete skull.
Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch et al. in 2004 as the most recent common ancestor of Malawisaurus and Saltasaurus and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek lithostros, meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch et al. include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous.
Pakasuchus is a genus of notosuchian crocodyliform distinguished by its unusual mammal-like appearance, including mammal-like teeth that would have given the animal the ability to chew. It also had long, slender legs and a doglike nose. Pakasuchus lived approximately 105 million years ago, in the mid-Cretaceous. Fossils have been found in the Galula Formation of Rukwa Rift Basin of southwestern Tanzania, and were described in 2010 in the journal Nature. The type species is P. kapilimai. Pakasuchus means "cat crocodile" in reference to its catlike skull.
Panamericansaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of Argentina. The holotype is very similar to known fossil material of Aeolosaurus, which Panamericansaurus is thought to be closely related to.
Angolatitan is a genus of titanosauriform sauropod dinosaur from the Upper Cretaceous. It is also the first non-avian dinosaur discovered in Angola. The genus contains a single species, Angolatitan adamastor, known from a partial right forelimb. Angolatitan was a relict form of its time; it was a Late Cretaceous basal titanosauriform, when more derived titanosaurs were far more common.
Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found Aeolosaurus, Gondwanatitan, Maxakalisaurus, Panamericansaurus and Rinconsaurus to be aeolosaurids.
Overosaurus is an extinct genus of sauropod dinosaurs, containing only a single species, Overosaurus paradasorum. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia. Overosaurus paradasorum was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately 10 m (33 ft). It was a ground-dwelling herbivore.
Rukwasuchus is an extinct genus of peirosaurid mesoeucrocodylian known from the middle Cretaceous Galula Formation of southwestern Tanzania. It contains a single species, Rukwasuchus yajabalijekundu.
Rukwatitan is a genus of titanosaur sauropod dinosaur from the Galula Formation in Tanzania. It lived around 100 million years ago, during the middle Cretaceous. The species, which shared features with another southern African species, Malawisaurus dixeyi, measured 30 feet (9.1 m) from the head to the tip of the tail, and had forelimbs that were estimated around 6.5 feet (2.0 m) long. Its fossils were found embedded in a cliff face near Lake Rukwa in the Rukwa Valley, from which it gets its name.
Lohuecotitan is an extinct genus of titanosaurian sauropod dinosaur which lived during the Late Cretaceous in Spain. The only species known in the genus is Lohuecotitan pandafilandi, described and named in 2016.
Sibirotitan is a genus of somphospondyl sauropod from the Ilek Formation of Russia. The type and only species is S. astrosacralis.
Mansourasaurus is a genus of herbivorous lithostrotian sauropod dinosaur from the Quseir Formation of Egypt. The type and only species is Mansourasaurus shahinae.
The Galula Formation is a geological formation located south of Lake Rukwa in Tanzania, part of the Red Sandstone Group of the Rukwa Rift Basin. Along with the unconformably overlying Oligocene Nsungwe Formation. It is divided into two members, the lower Mtuka Member and the upper Namba Member.
Mnyamawamtuka is a genus of lithostrotian titanosaur sauropod dinosaur from the Cretaceous Galula Formation in Tanzania. The type and only species is M. moyowamkia.
The Nsungwe Formation is a formation in the Rukwa Rift Basin of the East African Rift System, it is Oligocene in age based on U-Pb dating of a tuff horizon within the formation. It is part of the Red Sandstone Group along with the uncomfortably underlying Mid-Cretaceous Galula Formation It is divided into two members, the lower Utengule Member, and the upper Songwe member. It is notable for being one of the most important Paleogene fossil deposits in Sub-Saharan Africa.
Bravasaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Ciénaga del Río Huaco Formation of La Rioja, Argentina. It contains one species, Bravasaurus arreirosorum.
Punatitan is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Ciénaga del Río Huaco Formation of La Rioja, Argentina. It contains one species, Punatitan coughlini.
Abditosaurus is an extinct genus of titanosaurian sauropod dinosaur from the Late Cretaceous (Maastrichtian) Tremp Group of Catalonia, Spain. The type and only species is Abditosaurus kuehnei. Phylogenetic analyses recover it within a clade of South American and African saltasaurines, distinct from other insular dwarf sauropods from the European archipelago. Abditosaurus inhabited the Ibero-Armorican Island, a prehistoric island made up of what is now Spain, Portugal, and southern France, and would have been the largest titanosaur species in its environment.
